Introduction

Colombia is a major biodiversity hotspot, it has one of the richest orchid floras in the world and it represents a major diversity center for the Pleurothallidinae. It is conveniently situated in the northwestern corner of South America, being the only country in that subcontinent to have both Atlantic and Pacific coasts, a terrestrial connection with Central America, and borders with Ecuador, Brazil, Peru and Venezuela. A third of its 1,141,748 km² of land is mountainous, a preferred ecosystem for pleurothallids, with the Andes splitting into three main branches known as the Eastern, Central and Western cordilleras. Isolated mountain ranges in both the Atlantic (Sierra Nevada de Santa Marta) and Pacific (Serranía del Baudó) regions, along with fragments of the Guiana Shield in the Amazon (Serrania del Chiribiquete, Serrania de la Macarena), also contribute to this outstanding plant diversity. Colombia’s National Park System consists of 56 protected areas, covering 113,910 km² of land (9.98% of the country’s land area).

Orchidaceae is by far the most species-rich of all plant families of the Colombian flora (Bernal et al. 2016). Estimates range from 3591-4270 species of orchids in the country (Bernal et al. 2016, Betancur et al. 2015), that is up to three times as many species as the second most specious family: the Asteraceae. The cited studies find that close to 35% of all Colombian orchids are members of the Pleurothallidinae subtribe, which is the largest subtribe in the Orchidaceae (Karremans 2016, Pérez-Escobar et al. 2017, Pridgeon 2005). This exclusively neotropical group has one of the highest evolutionary diversification rates in the orchid family (Pérez-Escobar et al. 2017), and grows at a rate of little less than 100 new species recognized per year (Karremans & Davin 2017). About half of all the Orchidaceae species described yearly in the world belong to this subtribe. This means that, this is also the fastest growing group, in species numbers, of all orchids. Of the top ten most diverse plant genera in Colombia four are orchid genera, and three of them belong to the Pleurothallidinae. Orchidaceae is also the family with most species endemic to Colombia, two thirds of those are pleurothallids (Bernal et al. 2016, Betancur et al. 2015).

Organizing the wealth information regarding pleurothallid diversity in a country such as Colombia represents a real challenge. An important effort of systematically compiling the information from previous works, herbaria and libraries was made by Viveros & Higgins (2007), who recognized 1112 species in their Checklist of Pleurothallidinae from Colombia. Subsequent works led by the Universidad Nacional de Colombia and the Colombian Government offered exhaustive compilations of the orchids of Colombia (Betancur et al. 2015), followed by the even more ambitious catalogue of plants and lichens of Colombia (Bernal et al. 2016). Betancur et al.’s Plan para el estudio y la conservación de las orquídeas en Colombia offered a notorious increase in the number of Pleurothallidinae recorded previously, listing 1529 names. However, not all of them were supported by voucher specimens. Bernal et al.’s Catálogo de Plantas y Líquenes de Colombia, the most comprehensive checklist ever compiled of the plants in the country, resulted in a more conservative estimate of 1286 species for the subtribe.

The striking differences in species numbers between these catalogues published within a decade reflects not only how actively Colombian Pleurothallidinae have been worked on in recent years, but also how difficult it is to recover, revise and reconcile the enormous body of information on these orchids that comes from botanical databases, herbaria, monographic works and other publications around the world. An unpublished compilation of all names attributable to Pleurothallidinae by the first author today includes 5639 accepted species and almost 13 thousand described taxon names (Karremans 2016, Karremans & Vieira-Uribe 2020); close to a third of those names are referable to Colombian pleurothallids. However, the preparation of a carefully crafted catalogue of the Pleurothallidinae of Colombia can only be achieved through a collaborative effort with a network of multiple local experts, and it has to be continuously revised and updated, as there is always room for improvement.

Here, we have extracted and reviewed all the pleurothallid taxa attributable to Colombia and recognize 1862 accepted species occurring in the country (Fig. 1), with ca. 60% of them (1103 species) being endemic. Colombia, according to our assessment, hosts more or less a third of all the species currently recognized in the subtribe, and is by all means one of the most important pleurothallid diversity centers in the world. The current account represents a significant increase of several hundred species from the 1286-1529 range of just a few years ago (Betancur et al. 2015, Bernal et al. 2016) in the number of Pleurothallidinae recorded for Colombia. These numbers are even more notorious considering we are explicitly excluding 142 names that had been listed previously for the country (see Excluded Taxa below). This reflects not only how active the work on Colombian pleurothallids still is, but also the necessity of continued efforts to document and understand the country’s extremely rich orchid flora. We hope that this catalogue serves as a tool for students, researchers, conservation and policy makers, and can be seen for what it is: a compilation, to the best of our knowledge, of the current understanding of the most species rich orchid group in one of the most orchidrich countries in the world. And, therefore, imperfect and ever evolving.

Materials and methods

Taxa included in the catalogue have been either listed in previous works or are here cited for the first time based on materials determined by one of us. We relied on the pleurothallid monographs published by Luer (1986, 1987, 1988, 1989, 1990, 1991a, 1992, 1993, 1994, 1995, 1996a, 1996b, 1996c, 1997, 1998a, 1998b, 1999, 2000a, 2000b, 2000c, 2001, 2002a, 2002b, 2003a, 2003b, 2004, 2005, 2006, 2007, 2009a, 2009b, 2010, 2011, 2016a, 2016b, 2017a, 2017b, 2018a,2018b, 2018c) and Luer & Thoerle (2012a), in addition to previous checklists of Colombian Pleurothallidinae (Bernal et al. 2016, Betancur et al. 2015, Viveros & Higgins 2007), and online databases such as Epidendra, GBIF, IPNI, Tropicos and the WCSP (Govaerts et al. 2022).

An attempt has been made to curate previous lists of names. We include only those taxa for which a Colombian voucher was available, occasionally a photographic voucher is indicated when no physical voucher has been located. Names for which no voucher has been located are not included for the time being. A single, representative voucher is indicated for each taxon. Whenever possible, we have made an effort to revise the determination of vouchers cited in other works. This has resulted in the exclusion of certain taxa previously cited for the country. An extensive list of excluded taxa is provided at the end, these include species listed previously for which no voucher could be located, vouchers that had been misidentified, or names for which no reference could be found. The exclusion of a taxon is not evidence that it does not occur in the country, but simply that there is no voucher to support its current inclusion. Some excluded taxa, as well as other new ones, will eventually be confirmed to occur in the country. Further curation is certainly required in large and taxonomically challenging genera, especially Lepanthes Sw., Masdevallia Ruiz & Pav., Pleurothallis R.Br. and Stelis Sw. The citation of vouchers is an important step in this direction, and curation is significantly aided by the availability of illustrations and photographs based on local specimens.

Vouchers were inspected at AMES, BM, BR, CAUP, COL, CUVC, F, FAUC, FMB, G, GH, GOET, HPUJ, HUA, JAUM, JBB, JBL, K, LE, MEDEL, MO, MOL, MPU, NY, P, QPLS, SEL, TOLI, UTMC, VALLE, W, WU, US. Type specimens are given for each species discussed, lectotypes and neotypes are proposed when holotypes are lost or they were not clearly established. Unfortunately, we have not been able to locate original materials of a few taxa which remain poorly understood. These obscure species are accepted for the time being pending further evaluation. For each accepted taxon we offer a list of homotypic synonyms. Heterotypic synonyms of taxa based on Colombian specimens are listed at the end. Accepted species include country level distributions. Taxa that are endemic to Colombia are distinguished by an asterisk. A single representative Colombian voucher is given for taxa with foreign types.

Generic circumscription follows Pridgeon (2005) and Karremans (2016), with modifications proposed in Acianthera Scheidw. (Karremans & Rincón-González 2015, Karremans et al. 2016a), Andinia Luer (Wilson et al. 2017a), Andreettaea Luer (Doucette et al. 2022), Madisonia Luer (Smidt et al. 2021), Ophidion Luer (Karremans & Vieira-Uribe 2020), Specklinia Lindl. (Karremans et al. 2016b), Stelis (Karremans et al. 2013) and the genera in the Anathallis Barb.Rodr. Lepanthes Trichosalpinx Luer complex (Bogarín et al. 2019, Karremans 2014).

Results and discussion

We estimate that Colombia currently holds 1862 accepted species of Pleurothallidinae, of which 1103 species (ca. 60%) are endemic to the country. We expect pleurothallids to represent close to half the Colombian orchid flora. These taxa are unevenly distributed across 40 genera represented in the country (Fig. 2, Table 1), out of 46 in the whole subtribe. The numbers per genus range from a single species in Draconanthes (Luer) Luer, Gravendeelia Karremans & Bogarín, Frondaria Luer, Restrepiella Garay & Dunst. and Stellamaris Mel.Fernández & Bogarín, to over 500 taxa in Stelis. The most species rich genus in the country is Stelis with 521 species, followed by Lepanthes with 377 species, Pleurothallis with 248 species and Masdevallia with 171 species; all other genera are represented by 80 or less species. This pattern of species diversity is consistent with that we find across the subtribe, with Stelis, Lepanthes, Pleurothallis and Masdevallia respectively being the four most species-rich genera in the pleurothallids (Karremans & Vieira-Uribe 2020). Colombia is an extremely important center of radiation for species in these genera and Pleurothallidinae in general.

Just about 40% of pleurothallids occurring in Colombia are found elsewhere in the Neotropics. The greatest floristic affinity is found with Ecuador, with which Colombia shares 31.2% of its Pleurothallidinae. Strikingly, Ecuador hosts over 76% of the pleurothallid species found in Colombia which are not endemic to the country (Table 2). It is followed at a distance by other Andean countries, especially Venezuela (12.9% overall shared, 31.7% of non-endemics shared) and Peru (11.0% overall shared, 26.9% of non-endemics shared). Between 6.5-7.7% of the Pleurothallidinae occurring in Colombia are also found in either Bolivia, Costa Rica or Panama, while less than 5% affinity is found with species-rich but geographically distant, countries such as Brazil, Guatemala and Mexico.

Endemism rates and affinity patters differ among pleurothallid genera. Some genera, including Dresslerella Luer, Echinosepala Pridgeon & M.W.Chase, and Pendusalpinx Karremans & Mel.Fernández, seem to have no Colombian endemics, while endemism is very high in others such as Andinia, Dracula Luer, Lepanthes, Masdevallia, and Stelis. It is striking that of the whopping 377 species of Lepanthes known to occur in Colombia only 5 species (1.3%) are shared with Costa Rica and Panama (Bogarín et al. 2014, Pupulin 2002), two highly biodiverse countries that are the closest to Colombia in Central America. A similar situation occurs with Masdevallia, where only 6 species (3.5%) are shared with those two countries. Both well below the average number of shared taxa for the whole subtribe.

Overall, about a third of all currently known pleurothallids occur in Colombia, with marked differences among the genera (Table 1). Only 2.9-7.3% of the species of Brazilian-centered genera such as Anathallis and Pabstiella Brieger & Senghas are found to occur in the Colombia. Also, poorly represented are the northern genus Echinosepala, and the southern genus Octomeria R.Br., as well as Acianthera, Dresslerella Luer and Teagueia (Luer) Luer. Contrarily, a significant number of species belonging to the specious genera Andinia, Dracula, Ophidion, Pleurothallis, Pleurothallopsis Porto & Brade, Restrepia Kunth and Zootrophion Luer, are found to occur in Colombia, as well as all the species of the smaller genera Diodonopsis Pridgeon & M.W.Chase, Gravendeelia, Pseudolepanthes (Luer) Archila and Stellamaris. Only six pleurothallid genera are not currently represented in the Colombian flora: Atopoglossum Luer, Chamelophyton Garay, Luerella Braas, Opilionanthe Karremans & Bogarín, Phloeophila Hoehne & Schltr., and Pupulinia Karremans & Bogarín. All but Atopoglossum occur in countries sharing a border with Colombia and could be eventually recorded in the country.

A significant portion of the Pleurothallidinae of Colombia have been described in modern times. Hundreds of taxa were proposed in the last few decades by Carl Luer, leading pleurothallid authority. Colombian nationals including Rodrigo Escobar, father Pedro Ortiz Valdivieso, Oscar Duque, and Guillermo Misas Urreta, among others, have also made significant impact. However, several historically important botanical expeditions, early collectors, and botanists, have greatly influenced the research on the Orchidaceae of Colombia. The country’s rich orchid history includes many celebrated names such as Bonpland, Funck, Goudot, Hartweg, von Humboldt, von Jacquin, Karsten, Lehmann, Linden, Mutis, Purdie, Roezl, Schlim, Triana, Wagener, Wallis, von Warscewicz, and countless others. A thorough historical account is given by Ossenbach & Jenny (2021), our main focus here is on the typification of the materials collected. The interpretation in modern times of original materials of certain early taxa can represent a true challenge. Many classical botanists provided little information in the original descriptions and they often lacked a very precise accompanying illustration. In such cases the study of the original materials becomes essential in understanding a taxon. However, holotypes may not always be available, especially those by authors such as Friedrich Kraenzlin and Rudolf Schlechter whose main collections were destroyed in Berlin. We provide lectotypifications for several dozen taxa for which the holotype has been lost or it was not clearly selected. We follow Pupulin et al. (2022) in recognizing Schlechter’s floral analyses reproduced by Mansfeld as original material and designate them as lectotypes when no other original material is available. Even though rudimentary, those illustrations are important guides to determining the identities of those taxa. Unfortunately, we were unable to locate original materials, including illustrations, for some species. Little-known botanical figures such as Hermann Hopf and Werner Hopp, sent their personal collections directly to Rudolf Schlechter (Ossenbach & Jenny 2021), and it is likely that all their materials were lost in Berlin (Ossenbach pers. comm.). Those taxa, even though often accepted, are likely to remain cryptic.

One would expect such issues not to occur in modern times, but unfortunately this is not always the case when it comes to the description of orchids. Certain authors continue to propose new species in complex groups, such as the Pleurothallidinae, based only on a single (sometimes aberrant) specimen, often without proper descriptions, and detailed accompanying drawings or photographs. Many of these become modern cryptic taxa that cannot be adequately placed among their closest relatives, and become accepted just because they are poorly documented and understood. Often, the proposing authors themselves have no firsthand knowledge of the species in nature and/ or have not physically inspected the type specimen themselves, and therefore lack sufficient information to properly place their alleged novel taxon. In the more extreme cases this unfamiliarity leads to severe misdiagnoses. Examples of the latter are Acianthera ortizii Uribe Vélez, Sauleda & Szlach. and Crocodeilanthe asymmetrica Kolan. & Szlach. Both of which, despite possible differences in the interpretation of generic circumscriptions, have prior names in genus Pleurothallis where they ought to have been placed. Another rather curious example is the description of Crocodeilanthe velaticauloides Kolan. & Szlach., the holotype of which is an isotype of a prior taxon: Stelis pennelliana (Luer) Pridgeon & M.W.Chase. The latter, which is clearly a synonym, is not even mentioned in the diagnosis (Kolanowska & Szlachetko 2020b) despite being conspecific and indistinguishable. This raises suspicion as to how faithful and reliable such descriptions really are, especially given that S. pennelliana is used in that very same publication as a basis of comparison in the diagnosis of a few other newly proposed taxa. In addition, it is worrying that the type specimens of some of these modern taxa (see notes below) could not be retrieved in the Colombian herbaria where they were allegedly deposited.

Discovering and describing new taxa has become much more accesible with the advent of universal online access through the internet to databases, literature, and social media. Online publication, which was introduced as a means to increase the access and speed of formally describing taxa, has unfortunately promoted informality and illegality in the process. Some resources offer overnight publication, without the hassle of having an editorial board or a peer review process. This may appear convenient, but in practice it mostly stimulates poorly executed research, and the publication of taxa that often lack locality information (sometimes to avoid legal prosecution), proper illustrations, and stantardized data analyses. But poor-quality taxonomic research is also published in more formal journals, and can only be discouraged by more strict requirements by the code of botanical nomenclature, proper journal review and stern governmental policies. Good quality drawings and photographs are not required for the formal description of new species but should be considered a bare minimum for any serious taxonomic proposal at this day and age. The description of a taxon that none of the authors has physically inspected should not be permitted either, nor should describing a new taxon formally using only a schematic illustration of floral segments. Such descriptions only inflate publication numbers and lists of species names, but serve very little in the understanding of biodiversity. Similarly, detrimental is the use of drawings or photographs based on specimens from other countries to illustrate the flora of Colombia, such as has been done with pleurothallids, and other orchids, in the Orchids of the Darien Gap (Kolanowska 2014). This practice was commonplace in the past when species were believed to be highly variable and widespread, but today, given the levels of local endemism and strict biogeographical patterns, it is simply bad science.

It is high time that institutions in and researchers from Latin American countries assume a leading role in the study of their own flora, taking a front seat in floristic and systematic projects. This can only be done by combining stern governmental policies towards scientific research and a strong group of local researchers and institutions that collaborate in generating, curating and publishing high quality research. Much work is still needed to fully grasp the diversity of pleurothallids, and orchids in general, in Colombia. We would like encourage researchers working on these, and other plants, to do so, adhering to critical ethical and scientific standards, so that the information produced can be useful to understanding and conserving biodiversity. There are enough novelties in Pleurothallidinae to keep several generations of students busy.

Catalogue of the Pleurothallidinae of Colombia

Index to Generic Names (with species counts)

Acianthera Scheidw. (53)

Anathallis Barb.Rodr. (9)

Andinia (Luer) Luer (37)

Andreettaea Luer (22)

Barbosella Schltr. (5)

Brachionidium Lindl. (17)

Diodonopsis Pridgeon & M.W.Chase (4)

Draconanthes (Luer) Luer (1)

Dracula Luer (80)

Dresslerella Luer (2)

Dryadella Luer (15)

Echinosepala Pridgeon & M.W.Chase (3)

Frondaria Luer (1)

Gravendeelia Bogarín & Karremans (1)

Karma Karremans (16)

Lankesteriana Karremans (6)

Lepanthes Sw. (377)

Lepanthopsis (Cogn.) Ames (14)

Madisonia Luer (2)

Masdevallia Ruiz & Pav. (171)

Myoxanthus Poepp. & Endl. (19)

Octomeria R.Br. (22)

Ophidion Luer (12)

Pabstiella Brieger & Senghas (4)

Pendusalpinx Karremans & Mel.Fernández (3)

Platystele Schltr. (51)

Pleurothallis R.Br. (248)

Pleurothallopsis Porto & Brade (12)

Porroglossum Schltr. (13)

Pseudolepanthes (Luer) Archila (10)

Restrepia Kunth (31)

Restrepiella Garay & Dunst. (1)

Scaphosepalum Pfitzer (17)

Specklinia Lindl. (29)

Stelis Sw. (521)

Stellamaris Mel.Fernández & Bogarín (1)

Teagueia (Luer) Luer (3)

Trichosalpinx Luer (7)

Trisetella Luer (7)

Zootrophion Luer (15)

Index to other generic names

Aberrantia Luer = see Acianthera

Acinopetala Luer = see Masdevallia

Acostaea Schltr. = see Specklinia

Acronia C.Presl = see Pleurothallis

Alaticaulia Luer = see Masdevallia

Ancipitia (Luer) Luer = see Pleurothallis

Anthereon Pridgeon & M.W.Chase = see Pabstiella

Apatostelis Garay = see Stelis

Apoda-prorepentia (Luer) Luer = see Acianthera

Arthrosia (Luer) Luer = see Acianthera

Aspegrenia Poepp. & Endl. = see Octomeria

Brachycladium (Luer) Luer = see Andinia

Brenesia Schltr. = see Echinosepala

Buccella Luer = see Masdevallia

Byrsella Luer = see Masdevallia

Colombiana Ospina = see Pleurothallis

Condylago Luer = see Stelis

Crocodeilanthe Rchb.f. & Warsz. = see Pleurothallis, Stelis

Cryptophoranthus Barb.Rodr. = see Ophidion , Zootrophion

Dialissa Lindl. = see Stelis

Didactylus Luer = see Acianthera

Dubois-reymondia H.Karst. = see Myoxanthus

Duboisia R.Br. = see Myoxanthus

Echinella Pridgeon & M.W.Chase = see Echinosepala

Effusiella Luer = see Stelis

Elongatia (Luer) Luer = see Pleurothallis

Expedicula Luer = see Lepanthopsis

Fissia (Luer) Luer = see Masdevallia

Gyalanthos Szlach. & Marg. = see Pabstiella

Kraenzlinella Kuntze = see Acianthera, Anathallis

Lindleyalis Luer = see Pleurothallis

Lothiania Kraenzl. = see Porroglossum

Lueranthos Szlach. & Marg. = see Andinia

Luzama Luer = see Masdevallia

Megema Luer = see Masdevallia

Mixis Luer = see Pleurothallis

Muscarella Luer = see Andreettaea

Neooreophilus Archila = see Andinia

Niphantha Luer = see Stelis

Oreophilus W.E.Higgins & Archila = see Andinia

Panmorphia Luer = see Anathallis, Lankesteriana

Penducella Luer & Thoerle = see Andinia

Petalodon Luer = see Masdevallia

Physosiphon Lindl. = see Lepanthopsis, Stelis

Physothallis Garay = see Acianthera

Pteroon Luer = see Masdevallia

Reichantha Luer = see Masdevallia

Restrepiopsis Luer = see Pleurothallopsis

Rhynchopera Link, Klotzsch & Otto = see Pleurothallis

Rodrigoa Braas = see Masdevallia

Ronaldella Luer = see Pabstiella

Sarcinula Luer = see Specklinia

Spectaculum Luer = see Masdevallia

Spilotantha Luer = see Masdevallia

Streptoura Luer = see Masdevallia

Sylphia Luer = see Andreettaea, Specklinia

Talpinaria H.Karst. = see Pleurothallis

Tigivesta Luer = see Pleurothallis

Triaristella (Rchb.f.) Brieger ex Luer = see Trisetella

Triaristellina Rauschert = see Trisetella

Tubella (Luer) Archila = see Karma

Unciferia Luer = see Stelis

Verapazia Archila = see Andreettaea

Vestigium Luer = see Pleurothallis

Xenosia Luer = see Andinia

Zahleria Luer = see Masdevallia

Zosterophyllanthos Szlach. & Marg. = see Pleurothallis

Country abbreviations (listed alphabetically)

ARG Argentina

BEL Belize

BOL Bolivia

BRA Brazil

COL Colombia

COS Costa Rica

CUB Cuba

DOM Dominican Republic

ECU Ecuador

ELS El Salvador

FRE French Guiana

GUA Guatemala

GUY Guyana

HAI Haiti

HON Honduras

JAM Jamaica

L.A. Lesser Antilles

MEX Mexico

NIC Nicaragua

PAN Panama

PER Peru

PUE Puerto Rico

SUR Suriname

URU Uruguay

USA United States of America

VEN Venezuela

Acianthera Scheidw

Acianthera aberrans (Luer) Pupulin & Bogarín, Lankesteriana 8(2): 53-55. 2008. Aberrantia aberrans (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 310. 2005. Aberrantia aberrans (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004, nom. inval. Pleurothallis aberrans Luer, Selbyana 2(4): 382-383. 1978.

Illustration: Luer 2004, f. 108; Bogarín et al. 2008, f. 1, 15A; Karremans & Vieira-Uribe 2020, p. 23, p. 24; Misas Urreta 2005, p. 509.

Distribution: COL COS ECU PAN.

Type: Panama, Luer & Dressler 1628 (holotype, SEL).

Voucher: Colombia, Misas Urreta 255 (HPUJ).

Acianthera agathophylla (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 241. 2001. Humboldtia agathophylla (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis agathophylla Rchb.f., Xenia Orchid. 3: 25. 1881.

Illustration: Luer 2004, f. 3; Misas Urreta 2005, p. 515; Karremans & Vieira-Uribe 2020, p. 49.

Distribution: BOL BRA COL PER.

Type: Bolivia, Mandon 1132 (holotype, W-R45939; isotypes, AMES-00074025, BR-0000006586072, BM-000082439, G-00168967, K-000584177, MPU-018064, NY-00009220, P-00489101, P-00489102, P-00489103, P-00489104, S05-6340, W-R-0075167; tracing, W-2017-02129).

Voucher: Colombia, Misas Urreta 106 (HPUJ).

Acianthera angustifolia (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 118. 2007. Acianthera angustifolia (Lindl.) Carnevali & G.A.Romero, Nuevo Cat. Fl. Vasc. Venezuela 753. 2008, nom. superfl. Pleurothallis angustifolia Lindl., Edwards’s Bot. Reg. 21: sub t. 1797. 1835. Humboldtia angustifolia (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 2004, f. 4; Stenzel 2004, f. 41; Stenzel 2007, f. 29; Karremans & Vieira-Uribe 2020, p. 40.

Distribution: BEL COL CUB DOM GUA HAI HON JAM PUE L.A. MEX VEN.

Type: Mexico, Deppe s.n. (holotype, K-L000079872; isotypes, BM-000082661).

Voucher: Colombia, Silverstone-Sopkin 6292 (MO; Luer ill. 16340)

Acianthera aphthosa (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 242. 2001. Pleurothallis aphthosa Lindl., Edwards’s Bot. Reg. 24(Misc.): 42-43. 1838. Specklinia aphthosa (Lindl.) F.Barros, Hoehnea 10: 109. 1983 [1984].

Illustration: Cogniaux 1896, t. 107 (III), t. 108 (II); Luer 2004, f. 6; Karremans & Vieira-Uribe 2020, p. 18.

Distribution: BOL BRA COL ECU GUA MEX PER.

Type: Unknown country, Bateman s.n. (holotype, K-L-000674223).

Voucher: Colombia, McCullough 1862 (SEL; Luer ill. 228-S).

Acianthera beyrodtiana (Kraenzl.) Karremans, Harvard Pap. Bot. 21(2): 185. 2016.

Pleurothallis beyrodtiana Kraenzl., Repert. Spec. Nov. Regni Veg. 6: 65. 1908. Pleurothallis cogniauxiana Kraenzl., Orchis 2: 28. 1907, nom. illeg. Non Pleurothallis cogniauxiana Schltr. [1907].

Illustration: Luer 2004, f. 21, f. 22.

Distribution: COL ECU.

Type: Colombia, Beyrodt s.n. (holotype, HBG501882).

Acianthera carcinopsis Luer & Sijm, Harvard Pap. Bot. 17(2): 334. 2012.

Illustration: Luer & Thoerle 2012b, f. 3; Karremans & Vieira-Uribe 2020, p. 26.

Distribution: *COL.

Type: Colombia, Sijm 20110830 (holotype, MO).

Acianthera chamelopoda (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 253. 2004. Pleurothallis chamelopodaLuer, Orquideología 20(2): 201-203. 1996.

Illustration: Luer 1996f, 202; Luer 2004, f. 16.

Distribution: *COL.

Type: Colombia, Lehmann 346 (holotype, K-L000079872; isotype, W).

Acianthera chionopa (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 242. 2001. Pleurothallis chionopa Luer, Phytologia 46(6): 361. 1980.

Illustration: Luer 2004, f. 17; Karremans & Vieira-Uribe 2020, p. 60.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 4494 (holotype, SEL).

Voucher: Colombia, Killip & Smith 16751 (AMES, NY, US).

Acianthera ciliata (Knowles & Westc.) F.Barros & L.R.S. Guim., Neodiversity 5: 28. 2010. Humboldtia ciliata (Knowles & Westc.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis ciliata Knowles & Westc., Fl. Cab. 1: 39-40, t. 19. 1837.

Illustration: Luer 2004, f. 56, f. 57; Karremans & Vieira-Uribe 2020, p. 35.

Distribution: BEL BOL BRA COL COS ECU GUY L.A. PER SUR VEN.

Type: Guyana, Henchman ex Wilmore s.n [holotype, not preserved; lectotype (Christenson 1996), Flor. Cab. 1: 39, t. 19].

Voucher: Colombia, Lehmann 312 (K, L).

Acianthera cremasta (Luer & J. Portilla) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 253. 2004. Pleurothallis cremasta Luer & J. Portilla, Selbyana 22(2): 119-121. 2001.

Illustration: Luer 2004, f. 28; Karremans & Vieira-Uribe 2020, p. 72. Distribution: COL ECU.

Type: Ecuador, Hirtz 7132 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & VieiraUribe 2020, p. 72.

Acianthera decipiens (Ames & C.Schweinf.) Pridgeon & M.W.Chase, Lindleyana 16(4): 243. 2001. Pleurothallis decipiens Ames & C.Schweinf., Sched. Orch. 8: 26-27. 1925.

Illustration: Luer 2004, f. 29; Karremans & Vieira-Uribe 2020, p. 38.

Distribution: COL COS ECU PAN VEN.

Type: Costa Rica, Valerio 68 (holotype, US; isotypes AMES).

Voucher: Colombia, Cuatrecasas 18770 (AMES).

Acianthera decurrens (Poepp. & Endl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 243. 2001. Humboldtia decurrens (Poepp. & Endl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis decurrens Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 48, pl. 83. 1836.

Illustration: Luer 2004, f. 30; Damián et al. 2018, f. 1D; Karremans & Vieira-Uribe 2020, p. 36.

Distribution: BOL COL ECU PER.

Type: Peru, Poepping 1604 [t.83] [lectotype, W-R0047866 (Karremans et al. 2023); isolectotype, W-R-0047867].

Voucher: Colombia, Betancur et al. 7018 (COL, MO).

Acianthera discophylla (Luer & Carnevali) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 253. 2004. Pleurothallis discophylla Luer & Carnevali, Novon 3(2): 158. 1993.

Illustration: Luer 2004, f. 31, f. 32, f. 33; Damián et al. 2018, f. 1E; Karremans & Vieira-Uribe 2020, p. 40.

Distribution: BOL BRA COL ECU PER VEN.

Type: Ecuador, Hirtz & Ortega 13927 (holotype, MO).

Voucher: Colombia, Barona 15 (COAH).

Acianthera dubbeldamii Luer & Sijm, Harvard Pap. Bot. 17(2): 336. 2012.

Illustration: Luer & Thoerle 2012b, f. 4; Karremans & Vieira-Uribe 2020, p. 35.

Distribution: *COL.

Type: Colombia, Sijm 20110829 (holotype, MO).

Acianthera erinacea (Rchb.f.) A.Doucette, Phytotaxa 275(3): 271. 2016. Humboldtia erinacea (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Kraenzlinella erinacea (Rchb.f.) Solano, Icon. Orchid. (Mexico) 5-6: xi. 2002 [2003]. Kraenzlinella erinacea (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004, nom. superfl. Pleurothallis erinacea Rchb.f., Bonplandia (Hannover) 3: 72. 1855.

Illustration: Luer 1994, pl. 3 (p. 97); Karremans & Vieira-Uribe 2020, p. 57.

Distribution: BEL BOL COL COS ECU GUA HON MEX NIC PAN PER SUR VEN.

Type: Colombia, Wagener s.n. (holotype, W-R53828; isotype, K-L).

Acianthera floribunda (Lindl.) F.Barros, Bradea 8: 294. 2002. Arthrosia floribunda (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 248. 2006. Pleurothallis floribunda (Lindl.) Lindl., Edwards’s Bot. Reg. Misc. 73. 1842. Specklinia floribunda Lindl., Gen. Sp. Orchid. Pl. 9. 1830.

Illustration: Damián et al. 2018, f. 1H.

Distribution: BRA, COL, ECU, L.A., PER.

Type: Martinique, Sieber 206 (holotype, K-L; isotypes, M, P, MO).

Voucher: Colombia, Viveros 135 (HUQ).

Acianthera fockei (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 243. 2001. Humboldtia fockei (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis fockei Lindl., Fol. Orchid. ~Pleurothallis~ 23. 1859. Pleurothallis tricarinata H.Focke, Bot. Zeitung (Berlin) 11: 339. 1853, nom. illeg. Non Pleurothallis tricarinata Poepp. & Endl. [1836]. Illustration: Luer 2004, f. 42, f. 98; Karremans & Vieira-Uribe 2020, p. 59.

Distribution: COL FRE GUY SUR VEN.

Type: Suriname, Wullschlaegel 1591 (holotype, GOET-013949; type illustr., K-L, W-R-0008958; tracing, W-R-0011538).

Voucher: Colombia, Clavijo-Romero 813 (COL).

Acianthera geminicaulina (Ames) Pridgeon & M.W.Chase, Lindleyana 16(4): 243. 2001. Pleurothallis geminicaulina Ames, Sched. Orch. 6: 59-60. 1923.

Illustration: Luer 1975a, f. 8 (p. 73); Luer 2004, f. 45; Karremans & Vieira-Uribe 2020, p. 15, p. 34; Pérez-Escobar et al. 2022, f. 4a-d; Misas Urreta 2005, p. 539.

Distribution: COL COS ECU PER.

Type: Costa Rica, Lankester 454 (holotype, AMES).

Voucher: Colombia, Espina 2752 (CHOCO, MO).

Acianthera gracilis (Barb.Rodr.) F.Barros & L.R.S. Guim., Neodiversity 5(1): 28. 2010. Arthrosia gracilis (Barb.Rodr.) Campacci, Bol. CAOB 69-70: 26. 2008. Pleurothallis gracilis Barb.Rodr., Gen. Sp. Orchid. 1: 6. 1877.

Distribution: BRA COL.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype, tab. 175, fig. B, in Iconogr. Orchid. Bresil, vol. 3, at the Library of Rio de Janeiro Botanical Garden; reproduced in color in Sprunger et al. (1996, vol. 1: 223) (Barros & Guimarães 2010)].

Voucher: Colombia, Ospina 117 (JAUM).

Acianthera hagsateriO.Pérez, L.Oses & Bogarín, Lankesteriana 22(1): 18. 2022.

Illustration: Pérez-Escobar et al. 2022, f. 1, f. 2, f. 4e.

Distribution: *COL.

Type: Colombia, Pérez & Parra 1488 (holotype, CUVC).

Acianthera harpophylla (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 253. 2004.

Humboldtia harpophylla (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis harpophylla Rchb.f., Bonplandia (Hannover) 3: 72. 1855.

Illustration: Luer 2004, f. 47.

Distribution: *COL.

Type: Colombia, Wagener s.n. [L549] (holotype, W-R-0021870; isotype; K-L).

Acianthera henrici (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 253. 2004. Pleurothallis henrici Schltr., Repert. Spec. Nov. Regni Veg. Beih. 8: 58. 1921.

Illustration: Luer 2004, f. 49; Zambrano & Solano 2021, f. 4b.

Distribution: COL ECU.

Type: Ecuador, Eggers 15628 [holotype, B, destroyed; lectotype, K-000584158 (Luer 2004)].

Voucher: Colombia, Haught 4823 (COL).

Acianthera hirtipes (Schltr.) Rojas-Alv. & Karremans, Phytotaxa 507: 252. 2021. Myoxanthus hirtipes (Schltr.) Luer, Selbyana 7(1): 43. 1982. Pleurothallis hirtipes Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 104. 1920.

Illustration: Mansfeld 1929, t. 39 (No. 136).

Distribution: *COL.

Type: Colombia, Madero s.n. (holotype, B, destroyed). Colombia, Madero s.n. [neotype, illustration in Mansfeld 1929, t. 39, No. 136 (Rojas-Alvarado et al. 2021)].

Acianthera hirtzii (Luer) Karremans & RincónGonzález, Phytotaxa 238(2): 178. 2015. Apoda-prorepentia hirtziiLuer, Harvard Pap. Bot. 18(2): 174. 2013.

Illustration: Luer & Thoerle 2013, f. 4; Karremans & Vieira-Uribe 2020, p. 30.

Distribution: COL ECU.

Type: Ecuador, Hirtz 10239 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & VieiraUribe 2020, p. 30.

Acianthera hygrophila (Barb.Rodr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 244. 2001. Arthrosia hygrophila (Barb.Rodr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 249. 2006. Pleurothallis hygrophila Barb.Rodr., Gen. Sp. Orchid. 1: 7. 1877. Specklinia hygrophila (Barb.Rodr.) F.Barros, Hoehnea 10: 110. 1984. Specklinia hygrophila (Barb.Rodr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004, nom. superfl. Stelis hygrophila (Barb.Rodr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001.

Illustration: Rossado et al. 2014, f. 1; Karremans & Vieira-Uribe 2020, p. 31.

Distribution: ARG BRA COL URU.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype (selected here), tab. 140, fig. B, in Iconogr. Orchid. Bresil, vol. 3, at the Library of Rio de Janeiro Botanical Garden; reproduced in color in Sprunger et al. (1996, vol. 1: 198)]. Brazil. Rio de Janeiro: Serra dos Órgãos, de Moura 14 [neotype, RB (Gonçalves & Waechter 2004), here rejected and selected as epitype; photo, RB0542551].

Voucher: Colombia, Wasum 12647 (MO).

Note: A neotype for this taxon was selected by Gonçalves & Waechter (2004) arguing that the extant type illustration reproduced in Sprunger et al. (1996) is insufficient to precisely apply the name. Nevertheless, this is still the only extant material among original materials and it does provide significant information regarding the floral features. The neotype is therefore here rejected and selected as epitype, which serves the purpose of an interpretative type.

Acianthera kateora (Garay) Karremans & RincónGonzález, Phytotaxa 238(2): 178. 2015. Apodaprorepentia kateora (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004. Physothallis kateora Garay, Orquideología 9: 133. 1973. Pleurothallis kateora (Garay) Luer, Phytologia 49(3): 209. 1981.

Illustration: Luer 2007, f. 5; Karremans & VieiraUribe 2020, p. 29.

Distribution: *COL.

Type: Colombia, Escobar 1078 (holotype, AMES).

Acianthera lojae (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Pleurothallis lojae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 8: 60-61. 1921.

Illustration: Luer 2004, f. 59, f. 60, f. 61; Karremans & Vieira-Uribe 2020, p. 37.

Distribution: BOL COL ECU PER.

Type: Ecuador, Lehmann 8202 [holotype, B, destroyed; lectotype (selected here), K-000584144].

Voucher: Colombia, Luer et al. 11365 (MO).

Acianthera madisonii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 244. 2001. Pleurothallis madisonii Luer, Selbyana 5(2): 171-172. 1979.

Illustration: Luer 2004, f. 62; Ortiz et al. 2010, p. 57, 58; Karremans & Vieira-Uribe 2020, p. 36.

Distribution: COL ECU.

Type: Ecuador, Madison 3922 (holotype, SEL).

Voucher: Colombia, Pérez et al. 512 (holotype, VALLE).

Acianthera melanoglossa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Pleurothallis melanoglossaLuer & R.Escobar, Orquideología 20: 74. 1996.

Illustration: Luer & Escobar 1996, p. 73; Luer 2004, f. 64.

Distribution: *COL.

Type: Colombia, Luer et al. 16795 (holotype, MO).

Acianthera miqueliana (H.Focke) Pridgeon & M.W.Chase, Lindleyana 16(4): 244. 2001. Humboldtia miqueliana (H.Focke) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis miqueliana (H.Focke) Lindl., Fol. Orchid. ~Pleurothallis~ 17, no. 93. 1859. Specklinia miqueliana H.Focke, Tijdschr. WisNatuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Stelis miqueliana (H.Focke) Lindl., Fol. Orchid. ~Pleurothallis~ 17. 1859.

Illustration: Cogniaux 1896, f. 2 (p. 116); Luer 2004, f. 67; Damián et al. 2018, f. 2A.

Distribution: BRA COL ECU FRE GUY PER SUR VEN.

Type: Suriname, Focke s.n. (holotype, W; isotype, BR).

Voucher: Colombia, Betancur 13433 (COAH).

Acianthera moronae (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Pleurothallis moronaeLuer & Hirtz, Lindleyana 11(3): 171-172. 1996.

Illustration: Luer 2004, f. 69; Luer 1996c, f. 39; Karremans & Vieira-Uribe 2020, p. 37; Rincón-González et al. 2020a, f. 1.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 13998 (holotype, MO).

Voucher: Colombia, Rincón et al. 1297 (TOLIspirit).

Acianthera nellyae (P.Ortiz) Karremans, Harvard Pap. Bot. 21(2): 185. 2016. Pleurothallis nellyaeP.Ortiz, Orquideología 20: 325-326. 1997.

Illustration: Ortiz 1997, p. 317 (center left), p. 322 (below right).

Distribution: *COL.

Type: Colombia, Ortiz 1081 (holotype, HPUJ).

Acianthera omissa (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 245. 2001. Pleurothallis omissaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 175, f. 30a. 1999.

Illustration: Luer 1999a, f. 30a; Luer 2004, f. 73; Karremans & Vieira-Uribe 2020, p. 59.

Distribution: COL ECU VEN.

Type: Venezuela, Dunsterville 794 (holotype, MO).

Voucher: Colombia, Pipoly 12937 (COL).

Acianthera oscitans (Ames) Pridgeon & M.W.Chase, Lindleyana 16(4): 245. 2001. Didactylus oscitans (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 310. 2005. Didactylus oscitans (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004, nom. inval. Pleurothallis oscitans Ames, Bot. Mus. Leafl. 2(2): 25-27. 1934.

Illustration: Luer 2004, f. 110; Karremans & Vieira-Uribe 2020, p. 55, p. 56.

Distribution: COL COS ECU HON PAN.

Type: Honduras, Edwards 515 (holotype, AMES).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 55.

Acianthera pantasmi (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 245. 2001. Humboldtia pantasmi (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pantasmi Rchb.f., Bonplandia (Hannover) 3: 224. 1855.

Illustration: Luer 2004, f. 74; Luer 1975e, f. 91; Karremans & Vieira-Uribe 2020, p. 22.

Distribution: COL COS ECU HON NIC.

Type: Nicaragua, Oersted s.n. [L553] (holotype, W-R-0020817! isotypes, K-L-00079865).

Voucher: Colombia, Ortiz-Valdivieso 914 (HPUJ).

Acianthera pantasmoides (C.Schweinf.) Pridgeon & M.W.Chase, Lindleyana 16(4): 245. 2001. Pleurothallis pantasmoides C.Schweinf., Bot. Mus. Leafl. 15(3): 98. 1951.

Illustration: Luer 2004, f. 75; Luer & Thoerle 2012b, f. 2; Damián et al. 2018, f. 2B; Karremans & Vieira-Uribe 2020, p. 22.

Distribution: COL ECU PER.

Type: Peru, Vargas 6499 (holotype, AMES).

Voucher: Colombia, Pinto-Escobar 393 (COL).

Acianthera paziiLuer, Selbyana 30(1): 2. 2009. Pleurothallis pazii (Luer) Pfahl, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 2. 2013. 2013.

Illustration: Luer 2009b, f. 2.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin & Paz 8522 (holotype, CUVC; isotype, MO).

Acianthera phyllostachys (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 245. 2001. Pleurothallis phyllostachys Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 76. 1921.

Illustration: Cogniaux 1896, f. 2 (p. 119); Luer 2004, f. 78; Karremans & Vieira-Uribe 2020, p. 42.

Distribution: BOL BRA COL ECU PER.

Type: Peru, Weberbauer 6313 [holotype, B, destroyed; lectotype, AMES (Luer 2004)].

Voucher: Colombia, Lehmann 7100 (K, W).

Acianthera polystachya (Ruiz & Pav.) Pupulin, Anales Jard. Bot. Madrid 69: 27. 2012. Humboldtia polystachya Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 234. 1798. Stelis polystachya (Ruiz & Pav.) Willd., Sp. Pl. 4: 139. 1805.

Illustration: Luer 1975a, f. 3 (p. 63); Luer 2004, f. 13, f. 14; Damián et al. 2018, f. 2C. Fig. 3A; Karremans & Vieira-Uribe 2020, p. 41, p. 78.

Distribution: BOL COL ECU PER VEN.

Type: Peru, Ruiz & Pavón s.n. (holotype, MA).

Voucher: Colombia, Luer 10279 (MO).

Acianthera portilloi (Luer & R.Escobar) Karremans & Rincón-González, Phytotaxa 238(2): 178. 2015. Apoda-prorepentia portilloi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004. Pleurothallis portilloiLuer & R.Escobar, Orquideología 15(2-3): 144. 1982.

Illustration: Luer & Escobar 1982, p. 145; Luer 2007, f. 7.

Distribution: *COL.

Type: Colombia, Luer et al. 6688 (holotype, SEL; isotype, JAUM).

Acianthera prognatha (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16: 245. 2001.

Pleurothallis prognathaLuer & R.Escobar, Orquideología 16(1): 30. 1983.

Illustration: Luer & Escobar 1983, p. 29; Luer 2004, f. 79; Karremans & Vieira-Uribe 2020, p. 21.

Distribution: COL VEN.

Type: Colombia, Luer et al. 7918 (holotype, SEL; isotypes, COL, JAUM).

Acianthera pubescens (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 245-246. 2001. Humboldtia pubescens (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pubescens Lindl., Companion Bot. Mag. 2: 355. 1836.

Illustration: Cogniaux 1896, f. 1, f. 2 (p. 107); Luer 2004, f. 81; Zambrano & Solano 2021, f. 4d, f. 6, f. 7, f. 8, f. 9; Karremans & Vieira-Uribe 2020, p. 53, p. 54.

Distribution: BOL BRA COL COS ECU DOM GUA HON MEX NIC PAN PER PUE VEN.

Type: Mexico, Barker s.n. (holotype, K).

Voucher: Colombia, Luer et al. 10156 (MO).

Acianthera rodrigoi (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Pleurothallis rodrigoiLuer, Revista Soc. Boliv. Bot. 4(1): 19, f. 10. 2003.

Illustration: Luer 2003a, f. 10; Luer 2004, f. 85.

Distribution: *COL.

Type: Colombia, Escobar 5170 (holotype, MO).

Acianthera scalpricaulis (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 242. 2001. Pleurothallis scalpricaulis Luer, Selbyana 1(4): 424, f. 103. 1976.

Illustration: Luer 2004, f. 89; Damián et al. 2018, f. 2F; Karremans & Vieira-Uribe 2020, p. 18.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 619 (holotype, SEL).

Voucher: Colombia, Sierra-Ariza 108 (TOLI).

Acianthera serratifolia Rincón-González & Karremans, Phytotaxa 238(2): 178. 2015.

Illustration: Karremans & Rincón-González 2015, f. 2-4; Karremans & Vieira-Uribe 2020, p. 28.

Distribution: *COL.

Type: Colombia, Rincón-González 12962 (holotype, TOLI).

Acianthera sicaria (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 246. 2001. Humboldtia sicaria (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis sicaria Lindl., Edwards’s Bot. Reg. 27: Misc. 91. 1841.

Illustration: Luer 1975a, f. 17 (p. 91); Luer 2004, f. 90, f. 91, f. 92; Damián et al. 2018, f. 2G; Karremans & Vieira-Uribe 2020, p. 19, p. 20; Misas Urreta 2005, p. 568, 569.

Distribution: BOL COL COS ECU HON L.A. PAN PER VEN.

Type: Trinidad, Lodigges s.n. (holotype, K-L- 000079864).

Voucher: Colombia, Luer et al. 4569 (K, SEL).

Acianthera testifolia (Sw.) Solano, Acta Bot. Mex. 97: 50. 2011. Apoda-prorepentia testifolia (Sw.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004. Cymbidium testifolium (Sw.) Sw., Nova Acta Regiae Soc. Sci. Upsal. 6: 71. 1799. Epidendrum testifolium Sw., Prodr. Fl. Ind. Orient. 1: 122. 1788. Humboldtia testifolia (Sw.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis testifolia (Sw.) Lindl., Ann. Mag. Nat. Hist., ser. 3 1(5): 328. 1858.

Illustration: Stenzel 2004, f. 37; Luer 2007, f. 8; Stenzel 2007, f. 27; Karremans & Vieira-Uribe 2020, p. 29.

Distribution: COL COS CUB DOM GUA HAI HON JAM L.A. MEX NIC SUR VEN.

Type: Jamaica, Swartz s.n. (holotype, S-R1973; isotypes, BM-000082277, LD-1743206, M-0226572, SBT-11224, W-R-0016036).

Voucher: Colombia, Chaparro de Barrera 496 (COL).

Acianthera tunguraguae (F.Lehm. & Kraenzl.) A. Doucette, Phytotaxa 275(3): 272. 2016. Kraenzlinella tunguraguae (F.Lehm. & Kraenzl.) Engl. & Prantl, Nat. Pflanzenfam. Nachtr. 3: 86. 1906. Otopetalum tunguraguae F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26(3-4): 457. 1899.

Illustration: Luer 1975a, f. 12 (p. 81); Luer 1994, pl. 7 (p. 105); Damián et al. 2018, f. 2K; Karremans & Vieira-Uribe 2020, p. 57.

Distribution: COL ECU PER.

Type: Ecuador, Lehmann 8088 (holotype, K-000584257; isotypes, K-000584258, AMES00074566).

Acianthera verecunda (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 246. 2001. Pleurothallis verecunda Schltr., Repert. Spec. Nov. Regni Veg. Beih. 17: 24. 1922.

Illustration: Luer 2004, f. 100; Karremans & Vieira-Uribe 2020, p. 37.

Distribution: COL COS ECU PAN.

Type: Panama, Powell 91 [holotype, B, destroyed; lectotype, AMES-23795 (Christenson 1991; superfl. selected by Luer 2004); isolectotypes, K, MO955945].

Voucher: Colombia, Luer 18437 (MO).

Acianthera villahermosaeSierra-Ariza, Rinc.-González & Karremans, Phytotaxa 474(1): 93-97. 2020.

Illustration: Rincón-González et al. 2020b, f. 1-3, 4a; Karremans & Vieira-Uribe 2020, p. 29 (bottom left). Fig. 3B.

Distribution: *COL.

Type: Colombia, Ariza & Bohórquez 110 (holotype, TOLI-spirit).

Acianthera wageneriana (Klotzsch) Pridgeon & M.W.Chase, Lindleyana 16(4): 247. 2001.

Pleurothallis wageneriana Klotzsch, Allg. Gartenzeitung 20: 274. 1852.

Illustration: Cogniaux 1896, f. 1 (p. 105); Luer 2004, f. 103, f. 104; Karremans & Vieira-Uribe 2020, p. 21.

Distribution: BRA COL ECU PER VEN.

Type: Venezuela, Wagener 44 [lectotype (selected here), W-R-0025671; isolectotype, W-R-0017712; type illustr. W-R-0025672].

Voucher: Colombia, Escobar 8345 (MO).

Acianthera wyvern (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 247. 2001.

Pleurothallis wyvern Luer & R.Escobar, Orquideología 14(2): 184. 1981.

Illustration: Luer & Escobar 1981a, p. 185; Luer 2004, f. 105.

Distribution: COL ECU.

Type: Colombia, Luer et al. 3726 (holotype, SEL).

Acianthera yauaperyensis (Barb.Rodr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 247. 2001. Pleurothallis yauaperyensis Barb.Rodr., Vellosia 116. 1891.

Illustration: Cogniaux 1896, f. 3 (p. 116); Luer 2004, f. 106; Karremans & Vieira-Uribe 2020, p. 39.

Distribution: BRA COL GUY L.A. VEN.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype (selected here), tab. 126, fig. C, in Iconogr. Orchid. Bresil, vol. 2, at the Library of Rio de Janeiro Botanical Garden; reproduced in color in Sprunger et al. (1996, vol. 1: 182)].

Voucher: Colombia, Idrobo 2134 (COL).

Anathallis Barb.Rodr.

Anathallis anfracta (Luer) Karremans, Harvard Pap. Bot. 21(2): 186. 2016. Kraenzlinella anfracta (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004. Pleurothallis anfractaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 92, pl. 1. 1994.

Illustration: Ortiz 1997, p. 317 (center); Luer 1994, pl. 1 (p. 93); Karremans & Vieira-Uribe 2020, p. 83.

Distribution: *COL.

Type: Colombia, Lehmann 2388 (holotype, G).

Anathallis angustilabia (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 247. 2001. Pleurothallis angustilabia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 71. 1921. Specklinia angustilabia (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004.

Illustration: Luer 1999a, f. 3 (p. 126); Karremans & Vieira-Uribe 2020, p. 85.

Distribution: BOL COL ECU PER VEN.

Type: Peru, Weberbauer 6540 [holotype, B, destroyed; lectotype, AMES; isolectotype, F (Luer 1999a)].

Voucher: Colombia, Lehmann 9076 (AMES, K).

Anathallis brevipes (H.Focke) Pridgeon & M.W.Chase, Lindleyana 16(4): 247-248. 2001. Humboldtia brevipes (H.Focke) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Panmorphia brevipes (H.Focke) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 148, f. 103. 2006. Pleurothallis brevipes H.Focke, Tijdschr. WisNatuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 2: 198-199. 1849. Specklinia brevipes (H.Focke) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004. Trichosalpinx brevipes (H.Focke) Archila, Revista Guatemal. 17(1): 67. 2014 [2015].

Illustration: Luer 2006, f. 103; Karremans & Vieira-Uribe 2020, p. 98.

Distribution: BOL BRA COL ECU PER SUR.

Type: Suriname, Wullschlaegel 1589 [Focke illustr. 69] [lectotype (selected here), W-R-0043349; isolectotype, GOET-013893; type illustr., W-R0020789].

Voucher: Colombia, Luer 1961 (MO).

Anathallis citrina (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 248. 2001. Pleurothallis citrina Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 103. 1920. Specklinia citrina (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004.

Illustration: Luer 1999a, f. 8 (p. 127); Santos et al. 2019, f. 1d, f. 15; Karremans & Vieira-Uribe 2020, p. 87.

Distribution: BOL BRA COL PER.

Type: Colombia, Lehmann 6926 [lectotype (selected here), K-000584166; isolectotypes, K-000584167; LE-00001411].

Anathallis deborana (Carnevali & I.Ramírez) Carnevali & I.Ramírez, Harvard Pap. Bot. 21(1): 24. 2016. Pleurothallis deborana Carnevali & I.Ramírez, Ann. Missouri Bot. Gard. 77: 553. 1990. Specklinia deborana (Carnevali & I.Ramírez) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004.

Illustration: Romero-González et al. 2016, f. 2, f. 3

Distribution: COL VEN.

Type: Venezuela, Carnevali & Ramírez 2737 (holotype, VEN).

Anathallis herpethophyton (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 258. 2009. Panmorphia herpethophyton (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 158-159. 2006. Pleurothallis herpethophyton Schltr., Repert. Spec. Nov. Regni Veg. 27: 52. 1929. Specklinia herpethophyton (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004. Trichosalpinx herpethophyton (Schltr.) Archila, Revista Guatemal. 17(1): 69. 2014 [2015].

Illustration: Luer 2006, f. 119; Karremans & Vieira-Uribe 2020, p. 93. Fig. 3C.

Distribution: BOL COL ECU.

Type: Bolivia, Buchtien 6345 [holotype, B, destroyed; lectotype, US (Luer 2006)].

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 93.

Anathallis obovata (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Humboldtia obovata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis obovata (Lindl.) Lindl., Edwards’s Bot. Reg. 28: Misc. 75. 1842. Specklinia obovata Lindl., Edwards’s Bot. Reg. 25: Misc. 86. 1839.

Illustration: Cogniaux 1896, f. 2 (p. 110), f. 3 (p. 111), f. 1, f. 2 (p. 112); Luer 1999a, f. 19 (p. 130); Stenzel 2004, f. 26; Santos et al. 2019, f. 1l, f. 20; Karremans & Vieira-Uribe 2020, p. 85, p. 86.

Distribution: BOL BRA COL COS CUB DOM ECU ELS GUA GUY HAI HON NIC PER PUE VEN.

Type: Brazil, unknown collector (holotype, K-000674215; isotype, W).

Voucher: Colombia, Killip & Smith 16401 (AMES, NY).

Anathallis polygonoides (Griseb.) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Humboldtia polygonoides (Griseb.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Panmorphia polygonoides (Griseb.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 171, f. 139. 2006. Pleurothallis polygonoides Griseb., Fl. Brit. W. I. 609. 1864. Specklinia polygonoides (Griseb.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Trichosalpinx polygonoides (Griseb.) Archila, Revista Guatemal. 17(1): 72. 2014 [2015].

Illustration: Luer 2006, f. 139; Karremans & Vieira-Uribe 2020, p. 93.

Distribution: COL ECU FRE L.A. PER SUR VEN.

Type: Trinidad, Fendler 793 (holotype, K-000584077).

Voucher: Colombia, Prieto-Cruz 1072 (FMB).

Anathallis rabei (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 259. 2009.

Panmorphia rabei (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 171-172. 2006. Pleurothallis rabeiFoldats, Acta Bot. Venez. 3: 387. 1968. Specklinia rabei (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Trichosalpinx rabei (Foldats) Archila, Revista Guatemal. 17(1): 72. 2014 [2015].

Illustration: Foldats 1968, f. 27; Luer 1975b, f. 24; Luer 2006, f. 140a, f. 140b; Karremans & Vieira-Uribe 2020, p. 81.

Distribution: COL ECU PER VEN.

Type: Venezuela, Steyermark & Rabe 97263 (holotype, VEN).

Voucher: Colombia, Lehmann s.n. HK-140 (K).

Andinia (Luer) Luer

Andinia auriculipetala S.Vieira-Uribe & N. Gut., Spec. Orchid. 4(1): LCDP 37. 2020.

Illustration: Vieira-Uribe & Gutiérrez 2020, f. 37; Karremans & Vieira-Uribe 2020, p. 116. Fig. 3D

Distribution: *COL.

Type: Colombia, Calderón Hernández 190 (holotype, JBB).

Andinia cardiocheila (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 123. 2017. Brachycladium cardiocheilum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes cardiocheilaLuer & R.Escobar, Orquideología 19(2): 86-87. 1994. Lepanthes cardiochilaLuer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 6, pl. 1. 1994, sphalm. [?]. Neooreophilus cardiochilus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 75. 2009 [2011]. Oreophilus cardiochilus (Luer & R.Escobar) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella cardiocheila (Luer & R.Escobar) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer & Escobar 1994, f. 85 (p. 84); Luer 1994, pl. 1; Karremans & Vieira-Uribe 2020, p. 125.

Distribution: *COL.

Type: Colombia, Luer 16773 (holotype, MO).

Andinia catella (Luer & R.Escobar) Karremans & S. Vieira-Uribe, Phytotaxa 295(2): 124. 2017.

Brachycladium catellum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes catellaLuer & R.Escobar, Orquideología 16(1): 10. 1983. Neooreophilus catellus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 75. 2009 [2011]. Oreophilus catellus (Luer & R.Escobar) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella catella (Luer & R.Escobar) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer & Escobar 1983, p. 11; Luer 1994, pl. 2; Karremans & Vieira-Uribe 2020, p. 115.

Distribution: *COL.

Type: Colombia, Luer et al. 7554 (holotype, SEL).

Andinia chaoae (S.Vieira-Uribe & L. Jost) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Neooreophilus chaoaeS.Vieira-Uribe & L.Jost, Lankesteriana 15(3): 213. 2015.

Illustration: Vieira-Uribe & Jost 2015, f. 1, f. 2, f. 3b; Vieira-Uribe & Karremans 2017d, f. 1; Karremans & Vieira-Uribe 2020, p. 114.

Distribution: *COL.

Type: Colombia, Vieira 0020 (holotype, JAUM).

Andinia ciliaris (Luer & Hirtz) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017.

Brachycladium ciliare (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes ciliarisLuer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 8, f. 2. 1994. Neooreophilus ciliaris (Luer & Hirtz) Archila, Revista Guatemal. 12(2): 76. 2009 [2011]. Oreophilus ciliaris (Luer & Hirtz) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella ciliaris (Luer & Hirtz) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer 1994, pl. 3; Karremans & Vieira-Uribe 2020, p. 112. Fig. 3E.

Distribution: COL ECU.

Type: Ecuador, Hirtz 2887 (holotype, MO).

Voucher: Colombia, Gutiérrez-Morales 13 (JBB).

Andinia composita (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium compositum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes composita Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 11, pl. 4. 1994. Neooreophilus compositus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 77. 2009 [2011]. Oreophilus compositus (Luer & R.Escobar) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella composita (Luer & R.Escobar) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer & Escobar 1994, f. 87 (p. 84); Luer 1994, pl. 4; Karremans & Vieira-Uribe 2020, p. 123.

Distribution: *COL.

Type: Colombia, De Wilde 397 (holotype, MO).

Andinia dactyla (Garay) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium dactylum (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes dactyla Garay, Orquideología 6(1): 13. 1971. Neooreophilus dactylus (Garay) Archila, Revista Guatemal. 12(2): 78. 2009 [2011]. Oreophilus dactylus (Garay) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella dactyla (Garay) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer 1994, pl. 5; Karremans & Vieira-Uribe 2020, p. 117, p. 118.

Distribution: *COL.

Type: Colombia, Escobar et al. 645 (holotype, AMES).

Andinia destituta (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium destitutum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes destitutaLuer & R.Escobar, Orquideología 17(3): 179-182. 1988. Neooreophilus destitutus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 78. 2009 [2011]. Oreophilus destitutus (Luer & R.Escobar) Archila, Selbyana 29(2): 203. 2008 [2009]. Penducella destituta (Luer & R.Escobar) Luer, Orchid Digest 74(2): 69. 2010.

Illustration: Luer & Escobar 1988, p. 181; Luer 1994, pl. 6; Karremans & Vieira-Uribe 2020, p. 126.

Distribution: *COL.

Type: Colombia, Luer et al. 10342 (holotype, MO).

Andinia geminipetala (Luer & J. Portilla) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium geminipetalum (Luer & J. Portilla) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes geminipetalaLuer & J. Portilla, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 89, f. 314. 2002. Neooreophilus geminipetalus (Luer & J. Portilla) Archila, Revista Guatemal. 12(2): 79. 2009 [2011]. Oreophilus geminipetala (Luer & J. Portilla) Archila, Selbyana 29(2): 204. 2008 [2009]. Penducella geminipetala (Luer & J. Portilla) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Ortiz 2009b, f. 5 (p. 126); Luer 2002a, f. 314; Karremans & Vieira-Uribe 2020, p. 118.

Distribution: COL ECU.

Type: Ecuador, Luer 19656 (holotype, MO).

Voucher: Colombia, Ramirez et al. 7171 (CAUP).

Andinia hernandoi Est.Domínguez & S. Vieira-Uribe, Pleurothallids Neotropical Jewels I: 120. 2020.

Illustration: Karremans & Vieira-Uribe 2020, p. 119, p. 120. Fig. 3F.

Distribution: *COL.

Type: Colombia, Domínguez et al. 1673 (holotype, MEDEL).

Andinia hippocrepica (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium hippocrepicum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes hippocrepicaLuer & R.Escobar, Orquideología 18(1): 54, 88. 1991. Neooreophilus hippocrepicus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 80. 2009 [2011]. Oreophilus hippocrepicus (Luer & R.Escobar) Archila, Selbyana 29(2): 204. 2008 [2009]. Penducella hippocrepica (Luer & R.Escobar) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer & Escobar 1991a, p. 55; Luer 1994, pl. 8; Karremans & Vieira-Uribe 2020, p. 113.

Distribution: COL ECU.

Type: Colombia, Luer et al. 11343 (holotype, MO; isotype, JAUM).

Andinia hystricosa (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 251. 2001. Pleurothallis hystricosaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 54, 56, pl. 2. 1994.

Illustration: Luer 1994, pl. 2 (p. 57).

Distribution: *COL.

Type: Colombia, Little Jr. 8519 (holotype, US).

Andinia irrasa (Luer & R.Escobar) Karremans & S. Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium irrasum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes irrasa Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 18. 1994. Neooreophilus irrasus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 80. 2009 [2011]. Oreophilus irrasus (Luer & R.Escobar) Archila, Selbyana 29(2): 204. 2008 [2009]. Penducella irrasa (Luer & R.Escobar) Luer, Orchid Digest 74(2): 70. 2010; Karremans & Vieira-Uribe 2020, p. 112.

Illustration: Luer 1994, pl. 9.

Distribution: *COL.

Type: Colombia, Luer et al. 10341 (holotype, MO).

Andinia longiserpens (C.Schweinf.) Karremans & Mark Wilson, Phytotaxa 295(2): 126. 2017. Masdevalliantha longiserpens (C.Schweinf.) Szlach. & Marg., Polish Bot. J. 46(2): 117. 2001. Pleurothallis longiserpens C.Schweinf., Bot. Mus. Leafl. 10: 183, t. 20. 1942.

Illustration: Luer 2006, f. 47; Karremans & Vieira-Uribe 2020, p. 103.

Distribution: COL PER.

Type: Peru, Weberbauer 7510 (holotype, AMES; isotype, F, US, W).

Voucher: Colombia, Ortiz-Valdivieso 97 (COL).

Andinia lueriS.Vieira-Uribe & Karremans, Orquideología 33(2): 116. 2016.

Illustration: Karremans & Vieira-Uribe 2016, f. 1, f. 3(b), f. 4; Karremans & Vieira-Uribe 2020, p. 115.

Distribution: *COL.

Type: Colombia, Luer et al. 8034 (holotype, SEL).

Andinia lunaris (Luer) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium lunare (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes lunarisLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 21. 1994. Neooreophilus lunaris (Luer) Archila, Revista Guatemal. 12(2): 81. 2009 [2011]. Oreophilus lunaris (Luer) Archila, Selbyana 29(2): 205. 2008 [2009]. Penducella lunaris (Luer) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer 1994, pl. 10; Karremans & Vieira-Uribe 2020, p. 116.

Distribution: COL ECU.

Type: Colombia, Lehmann 277 (holotype, G).

Andinia lupula (Luer & Hirtz) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium lupulum (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes lupula Luer & Hirtz, Amer. Orchid Soc. Bull. 53(11): 1162. 1984. Neooreophilus lupulus (Luer & Hirtz) Archila, Revista Guatemal. 12(2): 81. 2009 [2011]. Oreophilus lupulus (Luer & Hirtz) Archila, Selbyana 29(2): 205. 2008 [2009]. Penducella lupula (Luer & Hirtz) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer 1994, pl. 11; Karremans & Vieira-Uribe 2020, p. 116. Fig. 3G.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9817 (holotype, SEL).

Voucher: Colombia, Vieira et al. 391 (JAUM).

Andinia macrotica (Luer & Dalström) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 126. 2017. Lepanthes macroticaLuer & Dalström, Monogr. Syst. Bot. Missouri Bot. Gard. 61(4): 3. 1996.Neooreophilus macrotica (Luer & Dalström) Archila, Revista Guatemal. 12(2): 82. 2009 [2011]. Oreophilus macroticus (Luer & Dalström) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 257. 2009. Penducella macrotica (Luer & Dalström) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer 1996b, pl. 26; Karremans & Vieira-Uribe 2020, p. 118.

Distribution: COL ECU.

Type: Ecuador, Dalström et al. 2132 (holotype, MO).

Voucher: Colombia, Vieira 390 (JAUM).

Andinia micropetala (L.O.Williams) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium micropetalum (L.O.Williams) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes micropetala L.O.Williams, Bot. Mus. Leafl. 9: 4. 1940. Neooreophilus micropetalus (L.O.Williams) Archila, Revista Guatemal. 12(2): 82. 2009 [2011]. Oreophilus micropetalus (L.O.Williams) Archila, Selbyana 29(2): 205. 2008 [2009]. Penducella micropetala (L.O.Williams) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer 1994, pl. 12; Karremans & Vieira-Uribe 2020, p. 111.

Distribution: COL ECU.

Type: Colombia, Pennell & Killip 7331 (holotype, AMES).

Andinia monilia (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium monilium (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes moniliaLuer & R.Escobar, Orquideología 16(1): 12. 1983. Neooreophilus monilius (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 83. 2009 [2011]. Oreophilus monilius (Luer & R.Escobar) Archila, Selbyana 29(2): 205. 2008 [2009]. Penducella monilia (Luer & R.Escobar) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer & Escobar 1983, p. 13; Luer 1994, pl. 13; Karremans & Vieira-Uribe 2020, p. 114.

Distribution: *COL.

Type: Colombia, Luer et al. 8036 (holotype, SEL).

Andinia montis-rotundi (P.Ortiz) Karremans & S. Vieira-Uribe, Phytotaxa 295(2): 124. 2017.

Lepanthes montis-rotundiP.Ortiz, Orquideología 20: 318. 1997. Oreophilus montis-rotundi (P.Ortiz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 257. 2009. Penducella montis-rotundi (P.Ortiz) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Ortiz 1997, p. 317 (top center), p. 322 (top left); Karremans & Vieira-Uribe 2020, p. 118.

Distribution: *COL.

Type: Colombia, Ortiz 705 (holotype, HUJ).

Andinia nummularia (Rchb.f.) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124. 2017. Brachycladium nummularium (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes nummularia Rchb.f., Xenia Orchid. 1: 142. 1856. Neooreophilus nummularius (Rchb.f.) Archila, Revista Guatemal. 12(2): 83. 2009 [2011]. Oreophilus nummularius (Rchb.f.) Archila, Selbyana 29(2): 205. 2008 [2009]. Penducella nummularia (Rchb.f.) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Luer 1994, pl. 14, pl. 15; Karremans & Vieira-Uribe 2020, p. 109, p. 110.

Distribution: COL ECU PER.

Type: Ecuador, Jameson 97 (holotype, W).

Voucher: Colombia, Luer 7554 (MO).

Andinia obesaS.Vieira-Uribe & Karremans, Lankesteriana 17(2): 311. 2017.

Illustration: Vieira-Uribe & Karremans 2017a, f. 1, 2; Karremans & Vieira-Uribe 2020, p. 110.

Distribution: *COL.

Type: Colombia, Vieira 0022 (holotype, JAUM).

Andinia ortiziana (S.Vieira-Uribe & Thoerle) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 124-125. 2017. Lepanthes ortizii J.M.H. Shaw, Orchid Review Suppl., 124(1314): 37. 2016. Neooreophilus ortizianusS.Vieira-Uribe & Thoerle, Orquideología 28(2): 135. 2011.

Illustration: Vieira-Uribe & Thoerle 2011, f. 1, f. 2, p. 138; Karremans & Vieira-Uribe 2020, p. 126.

Distribution: *COL.

Type: Colombia, Vieira 0001 (holotype, HPUJ).

Andinia panica (Luer & Dalström) Pridgeon & M.W.Chase, Lindleyana 16(4): 251. 2001. Pleurothallis panica Luer & Dalström, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 6. 1996.

Illustration: Luer 1996b, pl. 9; Karremans & Vieira-Uribe 2020, p. 108.

Distribution: COL ECU.

Type: Ecuador, Dalström et al. 2113 (holotype, MO).

Voucher: Colombia, Medina T.1124 (JAUM).

Andinia pendens (Garay) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium pendens (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes pendensGaray, Canad. J. Bot. 34: 252. 1956. Neooreophilus pendens (Garay) Archila, Revista Guatemal. 12(2): 84. 2009 [2011]. Oreophilus pendens (Garay) Archila, Selbyana 29(2): 206. 2008 [2009]. Penducella pendens (Garay) Luer, Orchid Digest 74(2): 70. 2010.

Illustration: Garay 1956a, f. 10D; Luer 1994, pl. 16; Karremans & Vieira-Uribe 2020, p. 126.

Distribution: *COL.

Type: Colombia, Grant 9770 (holotype, US; isotypes, AMES, WIS).

Andinia pilosella (Rchb.f.) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium pilosellum (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 307. 2005. Lepanthes pilosella Rchb.f., Flora 69: 556. 1886. Neooreophilus pilosellus (Rchb.f.) Archila, Revista Guatemal. 12(2): 85. 2009 [2011]. Oreophilus pilosellus (Rchb.f.) Archila, Selbyana 29(2): 206. 2008 [2009]. Penducella pilosella (Rchb.f.) Luer, Orchid Digest 74(2): 71. 2010.

Illustration: Garay 1956a, f. 10C; Luer 1994, pl. 18; Vásquez & Ibisch 2000, pl. 43; Ocupa Horna et al. 2021, f. 6d; Karremans & Vieira-Uribe 2020, p. 121, p. 122.

Distribution: BOL COL ECU.

Type: Colombia, Lehmann 3217 (holotype, W).

Andinia platysepala (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium platysepalum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005. Lepanthes platysepalaLuer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 36. 1994. Neooreophilus platysepalus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 85-86. 2009 [2011]. Oreophilus platysepalus (Luer & R.Escobar) Archila, Selbyana 29(2): 206. 2008 [2009]. Penducella platysepala (Luer & R.Escobar) Luer, Orchid Digest 74(2): 71. 2010.

Illustration: Luer 1994, pl. 19; Karremans & Vieira-Uribe 2020, p. 123, p. 124.

Distribution: *COL.

Type: Colombia, Luer et al. 11354 (holotype, MO).

Andinia pseudocaulescens (L.B. Sm. & S.K. Harris) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium pseudocaulescens (L.B. Sm. & S.K. Harris) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005. Lepanthes pseudocaulescens L.B. Sm. & S.K. Harris, Bot. Mus. Leafl. 2(3): 33. 1934. Neooreophilus pseudocaulescens (L.B. Sm. & S.K. Harris) Archila, Revista Guatemal. 12(2): 86. 2009 [2011]. Oreophilus pseudocaulescens (L.B. Sm. & S.K. Harris) Archila, Selbyana 29(2): 206. 2008 [2009]. Penducella pseudocaulescens (L.B. Sm. & S.K. Harris) Luer, Orchid Digest 74(2): 71. 2010.

Illustration: Luer 1994, pl. 20; Vieira-Uribe & Karremans 2018b, f. 14; Karremans & Vieira-Uribe 2020, p. 125.

Distribution: COL ECU.

Type: Colombia, Lehmann B.T. 352 (holotype, GH; isotype, AMES, G).

Andinia sibundoyensis (Kolan.) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Lepanthes sibundoyensis (Kolan.) J.M.H. Shaw, Orchid Review Suppl., 124(1314): 37. 2016. Neooreophilus sibundoyensis Kolan., Ann. Bot. Fennici 50: 169. 2014.

llustration: Karremans & Vieira-Uribe 2020, p. 113.

Distribution: *COL.

Type: Colombia, Kolanowska & Barrera s.n. (holotype, UGDA; isotype, COL).

Andinia spiralis (Ruiz & Pav.) Karremans & Mark Wilson, Phytotaxa 295(2): 126. 2017. Humboldtia spiralis Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 237. 1798. Pleurothallis spiralis (Ruiz & Pav.) Lindl., Gen. Sp. Orchid. Pl. 7. 1830. Stelis spiralis (Ruiz & Pav.) Pers., Syn. Pl. 2: 524. 1807. Xenosia spiralis (Ruiz & Pav.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 2006, f. 185; Luer 1975f, f. 82; Karremans & Vieira-Uribe 2020, p. 101, p. 102.

Distribution: COL ECU PER.

Type: Peru, Pavón s.n. (holotype, MA-814351).

Voucher: Colombia, Lehmann 4625 (W).

Andinia triangularis (Luer) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium triangulare (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005. Lepanthes triangularisLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 43. 1994. Neooreophilus triangularis (Luer) Archila, Revista Guatemal. 12(2): 87-88. 2009 [2011]. Oreophilus triangularis (Luer) Archila, Selbyana 29(2): 207. 2008 [2009]. Penducella triangularis (Luer) Luer, Orchid Digest 74(2): 71. 2010.

Illustration: Luer 1994, pl. 23.

Distribution: *COL.

Type: Colombia, Barclay & Juajibioy 6809 (holotype, MO).

Andinia trimytera (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 252. 2001. Pleurothallis trimyteraLuer & R.Escobar, Orquideología 16: 34. 1983.

Illustration: Luer & Escobar 1983, p. 35; Luer 1994, pl. 7 (p. 65); Karremans & Vieira-Uribe 2020, p. 108.

Distribution: COL ECU.

Type: Colombia, Luer et al. 8033 (holotype, SEL).

Andinia ursula (Luer & R.Escobar) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Brachycladium ursulum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005. Lepanthes ursula Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 46. 1994. Neooreophilus ursulus (Luer & R.Escobar) Archila, Revista Guatemal. 12(2): 88. 2009 [2011]. Oreophilus ursulus (Luer & R.Escobar) Archila, Selbyana 29(2): 207. 2008 [2009]. Penducella ursula (Luer & R.Escobar) Luer, Orchid Digest 74(2): 71. 2010.

Illustration: Luer 1994, pl. 24; Karremans & Vieira-Uribe 2020, p. 120.

Distribution: *COL.

Type: Colombia, Luer et al. 8035 (holotype, SEL).

Note: The type was cited as Luer et al. 8034 in the protologue, but the type specimen at SEL is labeled Luer et al. 8035. This is a correctable error and thus the number is emended here.

Andinia vestigipetala (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 252. 2001. Lueranthos vestigipetalus (Luer) Szlach. & Marg., Polish Bot. J. 46(2): 117. 2001 [2002]. Pleurothallis vestigipetala Luer, Selbyana 3(3,4): 404. 1977.

Illustration: Luer 1986, pl. 9; Luer 1994, pl. 8 (p. 67); Karremans & Vieira-Uribe 2020, p. 108.

Distribution: BOL COL ECU PER.

Type: Ecuador, Luer et al. 1496 (holotype, SEL).

Voucher: Colombia, Lehmann 7087 (K).

Andinia vieira-pereziana (P.Ortiz) Karremans & S.Vieira-Uribe, Phytotaxa 295(2): 125. 2017. Lepanthes vieira-pereziana (P.Ortiz) Pfahl, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 1 Feb. 2014. 2014. Neooreophilus vieira-perezianusP.Ortiz, Orquideología 28(1): 7. 2011.

Illustration: Ortiz 2011, p. 7(top), p. 10; Karremans & Vieira-Uribe 2020, p. 115.

Distribution: *COL.

Type: Colombia, Ortiz 1398 (holotype, HPUJ).

Note: Vieira & Pérez 003 indicated as isotype, but is a paratype.

Andinia xenion (Luer & R.Escobar) Karremans & Mark Wilson, Phytotaxa 295(2): 126. 2017. Pleurothallis xenionLuer & R.Escobar, Orquideología 16(1): 38. 1983. Xenosia xenion (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer & Escobar 1983, p. 39; Luer 1986, pl. 52; Karremans & Vieira-Uribe 2020, p. 103.

Distribution: *COL.

Type: Colombia, Portillo 7981 (holotype, SEL; isotypes, COL, JAUM).

Andreettaea Luer

Andreettaea barberiana (Rchb.f.) Karremans, comb. nov. Bas.: Pleurothallis barberiana Rchb.f., Gard. Chron., n.s. 16(392): 6. 1881. Humboldtia barberiana (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Muscarella barberiana (Rchb.f.) Szlach. & Sawicka, Orchids French Guiana: 195. 2012. Verapazia barberiana (Rchb.f.) Chiron & Szlach., Richardiana, N.S. 2: 167. 2018.

Illustration: Luer & Escobar 1996, p. 87; Luer 2006, f. 63, f. 90; Karremans & Vieira 2020, p. 179.

Distribution: COL ECU.

Type: Unknown country, Barber sub Low s.n. (holotype, W-R-25696).

Voucher: Colombia, Luer 16763 (MO).

Andreettaea bota-caucanensis (Uribe Vélez & Sauleda) Karremans, comb. nov. Bas.: Muscarella bota-caucanensisUribe Vélez & Sauleda, New World Orchid. Nomencl. Notes 101: 2. 2021.

Illustration: Uribe-Velez & Sauleda 2021, p. 2-5.

Distribution: *COL.

Type: Colombia, Ordoñez s.n. (holotype, HPUJ).

Note: The holotype of this taxon, like others described in this journal seem not to have been deposited at HPUJ. They may be invalid if it is confirmed that the type specimens were never prepared.

Andreettaea cabellensis (Rchb.f.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 2. 2022. Humboldtia cabellensis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Muscarella cabellensis (Rchb.f.) Karremans, Phytotaxa 272(1): 29. 2016. Pleurothallis cabellensis Rchb.f., Linnaea 22: 832. 1849. Specklinia cabellensis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 256-257. 2001. Sylphia cabellensis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 227. 2006. Verapazia cabellensis (Rchb.f.) Chiron & Szlach., Richardiana, N.S. 2: 167. 2018.

Illustration: Luer 2006, f. 179; Karremans & Vieira 2020, p. 172.

Distribution: COL VEN.

Type: Venezuela, Funk & Schlim 578 (holotype, W-R-0025568).

Voucher: Colombia, Schlim 1021 (W-R-0025568).

Andreettaea cestrochila (Garay) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 2. 2022. Muscarella cestrochila (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 97-98. 2006. Pleurothallis cestrochila Garay, Orquideología 8: 180. 1973. Specklinia cestrochila (Garay) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001.

Illustration: Luer 2006, f. 51; Karremans & Vieira 2020, p. 170.

Distribution: COL ECU.

Type: Colombia, Escobar 715 (holotype, AMES).

Andreettaea claviculata (Luer & Hirtz) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 3. 2022. Muscarella claviculata (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 98. 2006. Pleurothallis claviculata Luer & Hirtz, Lindleyana 11(3): 153-154. 1996. Specklinia claviculata (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004.

Illustration: Luer 2006, f. 52; Luer 1996c, f. 16; Karremans & Vieira 2020, p. 169; Karremans et al. 2021, f. 2b.

Distribution: COL ECU.

Type: Ecuador, Hirtz et al. 4635 (holotype, MO5259574).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 169.

Andreettaea clavigera (Luer) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 3. 2022. Muscarella clavigera (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 99. 2006. Pleurothallis clavigera Luer, Selbyana 3(3,4): 268. 1977. Specklinia clavigera (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001.

Illustration: Luer 2006, f. 53; Misas Urreta 2005, p. 519; Karremans & Vieira 2020, p. 178.

Distribution: COL ECU.

Type: Ecuador, Luer 1468 (holotype, SEL).

Voucher: Colombia, Misas Urreta 114 (HPUJ).

Andreettaea corynetes (Luer & R. Vásquez) A. Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 3. 2022. Muscarella corynetes (Luer & R.Vásquez) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 100. 2006. Pleurothallis corynetes Luer & R.Vásquez, Phytologia 46(6): 363. 1980. Specklinia corynetes (Luer & R.Vásquez) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001.

Illustration: Luer 2006, f. 55; Ortiz & Medina 2012, f. 11.

Distribution: BOL COL.

Type: Bolivia, Luer et al. 5131 (holotype, SEL).

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz & Medina (2012).

Andreettaea cryptophyta (Barb.Rodr.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 3. 2022. Lepanthes cryptophyta Barb.Rodr., Gen. Sp. Orchid. 2: 65. 1881. Muscarella cryptophyta (Barb.Rodr.) Bogarín & Karremans, Pleurothallids Neotropical Jewels I: 175. 2020.

Illustration: Cogniaux 1896, f. 4 (p. 116); Luer 2006, f. 96; Misas Urreta 2005, p. 549; Karremans & Vieira-Uribe 2020, p. 176.

Distribution: BOL BRA COL COS ECU VEN.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype, IOB (3) t. 116 (Karremans & Vieira-Uribe 2020)].

Voucher: Colombia, Luer 2187 (SEL).

Andreettaea divexa (Ames) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 4. 2022. Muscarella divexa (Ames) Karremans & S.Vieira-Uribe, Pleurothallids Neotropical Jewels I: 178. 2020. Pleurothallis dichotoma Ames, Sched. Orch. 6: 58-59. 1923, nom. illeg. Pleurothallis divexa Ames, Sched. Orch. 7: 20, t. 5. 1924. Non Pleurothallis dichotoma (Poepp. & Endl.) Schltr. [1921].

Illustration: Misas Urreta 2005, p. 566, p. 567; Karremans & Vieira-Uribe 2020, p. 177.

Distribution: COL COS.

Type: Costa Rica, Lankester 464 (holotype, AMES).

Voucher: Colombia, Misas Urreta 97 (HPUJ).

Andreettaea echinodes (Luer & R.Escobar) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 4. 2022. Muscarella echinodes (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 101-102. 2006. Pleurothallis echinodes Luer & R.Escobar, Orquideología 16(1): 24. 1983. Specklinia echinodes (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001.

Illustration: Luer & Escobar 1983, p. 25; Luer 2006, f. 58.

Distribution: *COL.

Type: Colombia, Luer et al. 7675 (holotype, SEL; isotypes, AAU, COL, JAUM).

Andreettaea ichthyonekys (Luer) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 5. 2022. Muscarella ichthyonekys (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 106-107. 2006. Pleurothallis ichthyonekys Luer, Selbyana 3(3-4): 316-318). 1977. Specklinia ichthyonekys (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001.

Illustration: Luer 2006, f. 66.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 1571 (holotype, MO).

Voucher: Colombia, Morales & Giraldo 6008 (HUA, MO).

Andreettaea intonsa (Luer & R.Escobar) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 5. 2022. Muscarella intonsa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 107-108. 2006. Pleurothallis intonsa Luer & R.Escobar, Orquideología 14(2): 156. 1981. Specklinia intonsa (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001.

Illustration: Luer & Escobar 1981a, p. 157; Luer 2006, f. 68; Karremans & Vieira-Uribe 2020, p. 175.

Distribution: *COL.

Type: Colombia, Escobar et al. 1121 (holotype, JAUM).

Andreettaea latilabris (Foldats) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 5. 2022. Muscarella latilabris (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 108-109. 2006. Pleurothallis latilabris Foldats, Acta Bot. Venez. 3: 376. 1968. Specklinia latilabris (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004, nom. superfl. Specklinia latilabris (Foldats) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001.

Illustration: Foldats 1968, f. 24; Luer 2006, f. 70; Karremans & Vieira-Uribe 2020, p. 169.

Distribution: COL VEN.

Type: Venezuela, Steryermark & Dunsterville 10881 (holotype, VEN).

Voucher: Colombia, De Wilde 4437 (MO).

Andreettaea macroblepharis (Rchb.f.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 6. 2022. Muscarella macroblepharis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 110-111. 2006. Pleurothallis macroblepharis Rchb.f., Gard. Chron., n.s. 2: 772. 1874. Specklinia macroblepharis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001.

Illustration: Luer 2006, f. 74; Karremans & VieiraUribe 2020, p. 180.

Distribution: COL ECU PER.

Type: Peru, Kirchhoff s.n. (holotype, W; isotype, AMES).

Voucher: Colomba, Luer 13084 (MO).

Andreettaea oblonga (Luer & Hirtz) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 6. 2022. Muscarella oblonga (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 113. 2006. Pleurothallis oblonga Luer & Hirtz, Lindleyana 11(3): 174. 1996. Specklinia oblonga (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 2006, f. 77; Luer 1996c, f. 42; Karremans & Vieira-Uribe 2020, p. 172.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12479 (holotype, MO).

Voucher: Colomba, González YGO2070 (COL).

Andreettaea quinqueseta (Ames) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 7. 2022. Pleurothallis quinqueseta Ames, Sched. Orch. 9: 35-37, f. 6. 1925. Muscarella quinqueseta (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 114. 2006. Specklinia quinqueseta (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer 2006, f. 79; Karremans et al. 2016, f. 8h; Karremans & Vieira-Uribe 2020, p. 182.

Distribution: COL COS PAN.

Type: Costa Rica, Standley 36283 (holotype, AMES; isotype, US).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 182.

Andreettaea samacensis (Ames) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 7. 2022. Muscarella samacensis (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 115-116. 2006. Pleurothallis samacensis Ames, Sched. Orch. 2: 22-24. 1923. Specklinia samacensis (Ames) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001.

Illustration: Luer 1975f, f. 74; Luer 2006, f. 81; Karremans & Vieira-Uribe 2020, p. 168.

Distribution: COL COS ECU GUA VEN.

Type: Guatemala, Ames 765 (holotype, AMES).

Voucher: Colombia, Luer 2311 (SEL).

Andreettaea semperflorens (Lindl.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 7. 2022. Pleurothallis acuminata H.Focke, Tijdschr. WisNatuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 2: 195-196. 1849, nom. illeg. Humboldtia semperflorens (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Muscarella semperflorens (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 117. 2006. Pleurothallis semperflorens Lindl., Fol. Orchid. ~Pleurothallis~ 40. 1859. Specklinia semperflorens (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001.

Illustration: Luer 1999a, f. 1 (p. 126); Luer 2006, f. 83; Karremans & Vieira-Uribe 2020, p. 174.

Distribution: BOL BRA COL ECU FRE GUY PER SUR VEN.

Type: Suriname, Focke 72 [lectotype (selected here), K-L, type illustr.; isolectotype, W-R-53801, type illustr. bottom left].

Voucher: Colombia, Lehmann 8204 (K).

Andreettaea sibatensis (F.Lehm. & Kraenzl.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 7. 2022. Muscarella sibatensis (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 117-118. 2006. Pleurothallis sibatensis F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 445. 1899. Specklinia sibatensis (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer 2006, f. 84.

Distribution: *COL.

Type: Colombia, Lehmann 2553 (holotype, K-000584285; isotypes, AMES-00074747, BM000082446, BR-0000006572587; LE-00006521).

Andreettaea tamboensis (Luer & R.Escobar) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 8. 2022. Muscarella tamboensis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 119-120. 2006. Pleurothallis tamboensis Luer & R.Escobar, Orquideología 14(2): 174. 1981. Specklinia tamboensis (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001.

Illustration: Luer & Escobar 1981a, p. 175; Luer 2006, f. 87; Karremans & Vieira-Uribe 2020, p. 171, p. 172.

Distribution: *COL.

Type: Colombia, Escobar et al. 1828 (holotype, JAUM).

Andreettaea villosilabia (Luer & Hirtz) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 8. 2022. Muscarella villosilabia (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 122, f. 91. 2006. Pleurothallis villosilabia Luer & Hirtz, Selbyana 23(1): 41, f. 30. 2002. Specklinia villosilabia (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Luer 2006, f. 91; Karremans & VieiraUribe 2020, p. 172.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 19622 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 172.

Andreettaea zephyrina (Rchb.f.) A.Doucette, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 8(1): 9. 2022. Humboldtia zephyrina (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Muscarella zephyrina (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 124-125. 2006. Pleurothallis zephyrinaRchb.f., Bonplandia (Hannover) 3: 71. 1855. Specklinia zephyrina (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004. Trichosalpinx zephyrina (Rchb.f.) Luer, Phytologia 54(5): 398. 1983.

Illustration: Luer 1975f, f. 79; Luer 2006, f. 95a, f. 95b, f. 95c; Karremans & Vieira-Uribe 2020, p. 171, p. 172. Fig. 3H.

Distribution: BOL COL ECU VEN.

Type: Colombia, Wagener s.n. [L556] (holotype, W-R-0025666; isotype, K-L).

Barbosella Schltr.

Barbosella cucullata (Lindl.) Schltr., Repert. Spec. Nov. Regni Veg. 15: 261. 1918. Pleurothallis angustisegmenta C.Schweinf., Bot. Mus. Leafl. 10: 139. 1942. Restrepia cucullata Lindl., Ann. Mag. Nat. Hist. 15: 108. 1845. Non Pleurothallis cucullata Ames [1923].

Illustration: Luer 2000a, f. 6 , f. 7, f. 8 (p. 31); Vásquez & Ibisch 2000, f. 2, pl. 1. Fig. 3I.

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Hartweg 1430 (holotype, K; isotypes, G, LD, W).

Barbosella dolichorhiza Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 117. 1920.

Illustration: Luer 2000a, f. 9 (p. 32).

Distribution: COL COS ECU PAN PER.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 41, No. 157)]. Ecuador, Luer et al. 9726 [neotype, MO (Luer 2000a), here rejected and selected as epitype].

Note: We follow Pupulin et al. (2022) in recognizing Schlechter’s floral analyses reproduced by Mansfeld as original material and therefore reject Luer’s neotype in favor of a lectotype.

Barbosella fuscataGaray, Orquideología 4: 153. 1969.

Illustration: Luer 2000a, f. 11, f. 12 (p. 32).

Distribution: COL ECU.

Type: Colombia, Escobar 417 (holotype, AMES).

Barbosella prorepens (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. 15(427/433): 263. 1918. Pleurothallis prorepens (Rchb.f.) Ames, F.T.Hubb. & C.Schweinf., Bot. Mus. Leafl. 3(2): 39-40. 1934. Restrepia prorepens Rchb.f., Gard. Chron., n.s. 7(183): 810. 1877.

Illustration: Luer 2000a, f. 17, f. 18, f. 19 (p. 34); Stenzel 2007, f. 7.

Distribution: BOL COL COS DOM ECU GUA L.A. NIC PAN PER VEN.

Type: Costa Rica, Endrés 2618 (holotype, W).

Voucher: Colombia, Lehmann 3305 (G, K, W).

Barbosella schistaLuer & R.Escobar, Orquideología 16(1): 7. 1983.

Illustration: Luer & Escobar 1983, p. 9; Luer 2000a, f. 21 (p. 35).

Distribution: *COL.

Type: Colombia, Luer et al. 7751 (holotype, SEL; isotypes, COL, JAUM).

Brachionidium Lindl.

Brachionidium brachycladum Luer & R.Escobar, Orquideología 16(3): 33-34. 1986.

Illustration: Luer & Escobar 1986, p. 35; Luer 1995, pl. 5.

Distribution: *COL.

Type: Colombia, Luer et al. 8946 (holotype, SEL).

Brachionidium dorisiae Kolan., Rykacz. Medina Tr. & Szlach., Nordic J. Bot. 2018: e01546: 3. 2017.

Illustration: Kolanowska et al. 2017, f. 2, 3.

Distribution: *COL.

Type: Colombia, Medina et al. S16/17 (holotype, JAUM-spirit).

Brachionidium ecuadorense Garay, Canad. J. Bot. 34: 731. 1956.

Illustration: Luer 1995, pl. 16; Ortiz & Medina 2012, f. 1.

Distribution: COL ECU.

Type: Ecuador, Lehmann 7057 (holotype, NY; isotype, K).

Voucher: Colombia, Delgado-V. 26 (PSO).

Brachionidium elzbietae Kolan., Rykacz. Medina Tr. & Szlach., Nordic J. Bot. 2018: e01546: 2. 2017.

Illustration: Kolanowska et al. 2017, f. 1.

Distribution: *COL.

Type: Colombia, Garay 837 (holotype, AMES; isotypes AMES, UGDA).

Brachionidium escobarii Luer, Lindleyana 1: 172. 1986.

Illustration: Luer 1995, pl. 19.

Distribution: *COL.

Type: Colombia, Luer et al. 3059 (holotype, MO; isotype, SEL).

Brachionidium filamentosum Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 57: 50, pl. 20. 1995.

Illustration: Luer 1995, pl. 20.

Distribution: COL ECU PAN.

Type: Ecuador, Luer et al. 15563 (holotype, MO).

Voucher: Colombia, Hartman 12434 (MO).

Brachionidium folsomiiDressler, Orquideología 15(2-3): 154-157. 1982.

Illustration: Dressler 1982, p. 155-157; Luer 1995, pl. 21. Fig. 3J.

Distribution: COL COS GUA PAN.

Type: Panama, Dressler 6032 (holotype, US; isotype, PMA).

Voucher: Colombia, Vieira 386 (JAUM).

Brachionidium fornicatum Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 57: 54-55, pl. 22. 1995.

Illustration: Luer 1995, pl. 22.

Distribution: COL ECU.

Type: Ecuador, Hirtz 6030 (holotype, MO).

Voucher: Colombia, Luer et al. 4667 (SEL).

Brachionidium imperiale Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 57: 64-65, pl. 27. 1995.

Illustration: Luer 1995, pl. 27.

Distribution: COL ECU.

Type: Colombia, De Wilde & Julian 16799 (holotype, MO).

Brachionidium lehmanniiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 57: 72-73, pl. 31. 1995.

Illustration: Luer 1995, pl. 31

Distribution: COL ECU.

Type: Ecuador, Lehmann s.n. [HK316] (holotype, K-000583667).

Voucher: Colombia, Arias 952 (HUAZ).

Brachionidium meridense Garay, Canad. J. Bot. 34: 736. 1956.

Illustration: Luer 1995, pl. 35.

Distribution: COL VEN.

Type: Venezuela, Renz 7507 (holotype, Herb. Renz).

Voucher: Colombia, Luer et al. 10194 (MO).

Brachionidium parvum Cogn., Repert. Spec. Nov. Regni Veg. 6: 307. 1909. Brachionidium sherringii var. parvum (Cogn.) Stehlé, Fl. Descript. Antilles Franç. 1: 219. 1939.

Illustration: Luer 1995, pl. 40; Stenzel 2007, f. 8. Fig. 3K.

Distribution: COL ECU HAI PUE L.A. VEN.

Type: Grenada, Broadway 268 [lectotype, K, isolectotype, BR (Luer 1995)].

Voucher: Colombia, Vieira 387 (JAUM).

Brachionidium parvifolium (Lindl.) Lindl., Fol. Orchid. Brachionidium 1. 1859. Restrepia parvifolia Lindl., Orchid. Linden. 4. 1846.

Illustration: Luer 1995, pl. 39.

Distribution: COL ECU.

Type: Colombia, Linden 1278 (holotype, K-L000079979; isotype, BR-0000009972827, W).

Brachionidium phalangiferum Garay, Canad. J. Bot. 34: 738, pl. 3. 1956.

Illustration: Luer 1995, pl. 42.

Distribution: COL VEN.

Type: Venezuela, Renz 6007 (holotype, herb. Renz).

Voucher: Colombia, Luer et al. 7968 (SEL).

Brachionidium puraceenseLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 57: 104-105, pl. 47. 1995.

Illustration: Luer 1995, pl. 47.

Distribution: *COL.

Type: Colombia, Lozano et al. 4697 (holotype, COL).

Brachionidium tetrapetalum (F.Lehm. & Kraenzl.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 241. 1920. Pleurothallis tetrapetala F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 446. 1899.

Illustration: Luer 1995, pl. 57.

Distribution: COL ECU.

Type: Colombia, Lehmann 347 (holotype, K-000583666; isotypes, AMES-00271853, BR0000009894860, W).

Brachionidium tuberculatum Lindl., Fol. Orchid. Brachionidium 1. 1859.

Illustration: Luer 1995, pl. 58, pl. 59; Vásquez & Ibisch 2000, pl. 6.

Distribution: BOL COL ECU VEN.

Type: Venezuela, Schlim 1558 (holotype, K-L000079980; isotype, G-00168549, LD, MPU- 017566, W).

Voucher: Colombia, Ospina Hernández 79 (AMES, COL).

Diodonopsis Pridgeon & M.W.Chase

Diodonopsis anachaeta (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001. Masdevallia anachaeta Rchb.f., Otia Bot. Hamburg. 1: 17. 1878.

Illustration: Luer 2000c, pl. 217, pl. 218; Vásquez & Ibisch 2000, f. 25, pl. 58. Fig. 3L.

Distribution: BOL COL ECU PER.

Type: Ecuador, Lehmann XXXII (holotype, W-R50752).

Voucher: Colombia, Luer & Escobar 7942 (SEL).

Diodonopsis erinacea (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001.

Masdevallia erinacea Rchb.f., Linnaea 41: 11-12. 1877. Scaphosepalum erinaceum (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 219. 1920.

Illustration: Luer 2000c, pl. 226, pl. 227.

Distribution: COL COS ECU PAN.

Type: Colombia, Roezl s.n. (holotype, W-R50791).

Diodonopsis pygmaea (Kraenzl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001.

Masdevallia pygmaea Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 92. 1925.

Illustration: Luer 2000c, pl. 242.

Distribution: COL COS ECU.

Type: Costa Rica, Endres 289 ([lectotype (selected here), W-R-41429; isolectotypes, W-R-14514, W-R-14536; type illustr., W-R-38559, left]).

Voucher: Colombia, Lehmann 2734 (K, G).

Diodonopsis ramiromedinaeThoerle, Lankesteriana 17(2): 223. 2017.

Illustration: Thoerle & Medina Trejo 2017, f. 1, 2.

Distribution: *COL.

Type: Colombia, Medina 98 (holotype, JAUM).

Draconanthes (Luer) Luer

Draconanthes aberrans (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 61(1): 3. 1996.

Lepanthes aberrans Schltr., Repert. Spec. Nov. Regni Veg. 14: 125. 1915.

Illustration: Luer 1996b, f. 1, f. 2, f. 3; Vásquez & Ibisch 2000, pl. 9; Mansfeld 1929, pl. 34 (n. 130); Kolanowska et al. 2020, f. 1-9. Fig. 4A.

Distribution: BOL COL COS ECU PER VEN.

Type: Ecuador, Sodiro 21 (holotype, B, destroyed). Ecuador, Hirtz 1056 [MO, neotype (Luer 1996b); SEL, isoneotype].

Voucher: Colombia, Luer et al. 12520 (MO).

Dracula Luer

Dracula adrianae Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 103. 1998.

Illustration: Luer 1998b, pl. 122 (p. 105).

Distribution: *COL.

Type: Colombia, Sijm 20898 (holotype, MO).

Dracula alcithoë Luer & R.Escobar, Orquideología 15: 3. 1981.

Illustration: Luer & Escobar 1981b, p. 4, p. 5; Luer 1993, pl. 1.

Distribution: COL ECU.

Type: Colombia, Ospina s.n. (holotype, JAUM; isotype, SEL).

Dracula amaliae Luer & R.Escobar, Orquideología 13: 5. 1978.

Illustration: Luer & Escobar 1978, p. 6, p. 7, p. 58 (bottom right); Luer 1993, pl. 2.

Distribution: *COL.

Type: Colombia, Escobar 1746 (holotype, JAUM).

Dracula andreettae (Luer) Luer, Selbyana 2(2-3): 193. 1978. Masdevallia andreettae Luer, Phytologia 39: 186-187. 1978.

Illustration: Luer 1993, pl. 3. Fig. 4B.

Distribution: COL ECU.

Type: Ecuador, Luer 1687 (holotype, SEL).

Voucher: Colombia, Escobar 1926 (JAUM).

Dracula anicula Luer & R.Escobar, Orquideología 15: 6. 1981.

Illustration: Luer & Escobar 1981b, p. 7-9; Luer 1993, pl. 4.

Distribution: *COL.

Type: Colombia, Escobar 1279 (holotype, JAUM; isotype, SEL).

Dracula anthracina Luer & R.Escobar, Selbyana 7: 64. 1982.

Illustration: Luer 1993, pl. 5.

Distribution: *COL.

Type: Colombia, Luer 3457 (holotype, SEL; isotype, JAUM).

Dracula antonii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 98. 2002.

Illustration: Luer 2002a, f. 2 (p. 115).

Distribution: *COL.

Type: Colombia, Sijm s.n. (holotype, MO).

Dracula aphrodes Luer & R.Escobar, Amer. Orchid Soc. Bull. 58: 1002. 1989.

Illustration: Luer 1993, pl. 6.

Distribution: *COL.

Type: Colombia, Escobar 3978 (holotype, JAUM; isotype, MO).

Dracula bella (Rchb.f.) Luer, Selbyana 2: 194. 1978. Masdevallia bella Rchb.f., Gard. Chron. 1: 725. 1878.

Illustration: Luer 1993, pl. 8.

Distribution: *COL.

Type: Colombia, Wallis 218 (holotype, W).

Dracula bellerophon Luer & R.Escobar, Orquideología 13: 9. 1978.

Illustration: Luer & Escobar 1978, p. 10, p. 11, p. 63 (top); Luer 1993, pl. 9.

Distribution: *COL.

Type: Colombia, Luer 2894 (holotype, JAUM; isotype, SEL).

Dracula benedictii (Rchb.f.) Luer, Selbyana 2: 194. 1978. Masdevallia benedictii Rchb.f., Xenia Orchid. 2: 197. 1874.

Illustration: Luer 1993, pl. 10.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W).

Dracula berthae Luer & R.Escobar, Orquideología 13: 115. 1979.

Illustration: Luer & Escobar 1979, p. 116, p. 117, p. 167 (bottom right); Luer 1993, pl. 11.

Distribution: *COL.

Type: Colombia, Escobar 1817 (holotype, JAUM).

Dracula brangeri Luer, Orchideeën 48: 47. 1986.

Illustration: Luer 1993, pl. 12.

Distribution: *COL.

Type: Colombia, Luer 11430 (holotype, MO).

Dracula callithrix N.Peláez, Buitr.-Delg. & Gary Mey., Orquideología 26(1): 38. 2009.

Illustration: Peláez et al. 2009, f. 8 (p. 39), f. 7 (p. 40).

Distribution: *COL.

Type: Colombia, Peláez NP0283 (holotype, JAUM-44187).

Note: Peláez NP0284 cited as isotype is a paratype.

Dracula carcinopsis Luer & R.Escobar, Amer. Orchid Soc. Bull. 58(10): 1003-1004. 1989.

Illustration: Luer 1993, pl. 13.

Distribution: *COL.

Type: Colombia, Escobar 4004 (holotype, JAUM; isotype, MO).

Dracula chestertonii (Rchb.f.) Luer, Selbyana 2: 194. 1978. Masdevallia chestertonii Rchb.f., Gard. Chron. 1: 532. 1883.

Illustration: Luer & Escobar 1978, p. 61(top right); Luer 1993, pl. 14.

Distribution: *COL.

Type: Colombia, Chesterton s.n. (holotype, W).

Dracula chimaera (Rchb.f.) Luer, Selbyana 2: 194. 1978. Masdevallia chimaera Rchb.f., Gard. Chron. 463. 1872.

Illustration: Luer 1993, pl. 15.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W).

Dracula chiroptera Luer & Malo, Selbyana 2: 367. 1978.

Illustration: Luer 1993, pl. 16.

Distribution: COL ECU.

Type: Ecuador, Luer 1759 (holotype, SEL).

Voucher: Colombia, Luer 6028 (SEL).

Dracula circe Luer & R.Escobar, Orquideología 15: 11. 1981.

Illustration: Luer & Escobar 1981b, p. 11-13; Luer 1993, pl. 17.

Distribution: *COL.

Type: Colombia, Escobar 1883 (holotype, JAUM; isotype, SEL).

Dracula citrina Luer & R.Escobar, Selbyana 7: 64. 1982.

Illustration: Luer 1993, pl. 18.

Distribution: *COL.

Type: Colombia, Luer 6545 (holotype, SEL; isotype, JAUM).

Dracula cochliops Luer & R.Escobar, Orquideología 13: 119. 1979.

Illustration: Luer & Escobar 1979, p. 121; Luer 1993, pl. 19. Fig. 4C.

Distribution: *COL.

Type: Colombia, Escobar 2011 (holotype, JAUM).

Note: The holotype is Escobar 2011 according to the protologue, but Luer (1993) cites Luer et al. 3127 (holotype, JAUM; isotype, SEL) as type, which is the number of his illustration based on that specimen.

Dracula cutis-bufonis Luer & R.Escobar, Orquideología 13: 13. 1978.

Illustration: Luer & Escobar 1978, p. 14, p. 15, p. 61(top left); Luer 1993, pl. 21.

Distribution: *COL.

Type: Colombia, Escobar et al. 1377 (holotype, JAUM).

Dracula decussata Luer & R.Escobar, Orquideología 13: 18. 1978.

Illustration: Luer & Escobar 1978, p. 20, p. 21, p. 62(top); Luer 1993, pl. 24; Luer 1996d, p. 11.

Distribution: *COL.

Type: Colombia, Escobar 1818 (holotype, JAUM).

Dracula dens-canis N.Peláez, Orchid Rev. 114: 344. 2006.

Illustration: Peláez 2006, p. 344-347.

Distribution: COL

Type: Colombia, Peláez NP0058 (holotype, HUA145622; isotypes, JAUM-038840; HUA-145623)

Dracula diabola Luer & R.Escobar, Orquideología 13: 124. 1979.

Illustration: Luer & Escobar 1979, p. 126, p. 127, p. 167(top); Luer 1993, pl. 26.

Distribution: *COL.

Type: Colombia, Escobar 1864 (holotype, JAUM).

Dracula diana Luer & R.Escobar, Orquideología 15: 15. 1981.

Illustration: Luer & Escobar 1981b, p. 15-17; Luer 1993, pl. 27.

Distribution: *COL.

Type: Colombia, Escobar 1882 (holotype, JAUM; isotype, SEL).

Dracula exasperata Luer & R.Escobar, Amer. Orchid Soc. Bull. 58(10): 1005. 1989.

Illustration: Luer 1993, pl. 30.

Distribution: *COL.

Type: Colombia, Luer 13185 (holotype, JAUM, not found; isotype, MO).

Dracula felix (Luer) Luer, Selbyana 2(2,3): 195. 1978. Masdevallia felix Luer, Phytologia 39: 201. 1978.

Illustration: Luer 1993, pl. 32.

Distribution: COL ECU.

Type: Colombia, Luer 2285 (holotype, SEL; isotype, JAUM).

Dracula gerhardii Luer & Sijm, Harvard Pap. Bot. 16(2): 324, 17. 2011.

Illustration: Luer 2011, f. 17.

Distribution: *COL.

Type: Colombia, Sijm 291122 (holotype, MO).

Dracula gigas (Luer) Luer, Selbyana 2(2,3): 195. 1978. Masdevallia gigas Luer, Phytologia 39: 204-205). 1978.

Illustration: Luer 1993, pl. 35

Distribution: COL ECU.

Type: Ecuador, Luer 1690 (holotype, SEL).

Voucher: Colombia, Luer 13011A (MO).

Dracula gorgona (H.J.Veitch) Luer & R.Escobar, Selbyana 2: 195. 1978. Masdevallia chimaera var. gorgona H.J.Veitch, Man. Orchid. Pl. 5: 31. 1889.

Illustration: Luer & Escobar 1978, p. 24, p. 25, p. 63(bottom right); Luer 1993, pl. 36.

Distribution: *COL.

Type: Colombia, Luer 13162 (neotype, MO).

Dracula gorgonella Luer & R.Escobar, Orquideología 15: 19. 1981.

Illustration: Luer & Escobar 1981b, p. 19-21; Luer 1993, pl. 37.

Distribution: *COL.

Type: Colombia, Escobar 1824 (holotype, JAUM).

Note: The holotype is Escobar 1824 according to the protologue, but Luer (1993) indicates that the type is Escobar 1975 (holotype, JAUM).

Dracula hirtzii Luer, Orchid Digest 43(1): 38. 1979.

Illustration: Luer 1993, pl. 40.

Distribution: COL ECU.

Type: Ecuador, Luer, 2596 (holotype, SEL).

Voucher: Colombia, Luer 12513 (MO).

Dracula houtteana (Rchb.f.) Luer, Selbyana 2: 195. 1978. Masdevallia houtteana Rchb.f., Gard. Chron. 2: 98. 1874.

Illustration: Luer 1993, pl. 41.

Distribution: *COL.

Type: Colombia, Wallis 314 (holotype, W).

Dracula inaequalis (Rchb.f.) Luer & R.Escobar, Selbyana 7: 65. 1982. Masdevallia inaequalis Rchb.f., Gard. Chron. 1: 372. 1874.

Illustration: Luer 1993, pl. 42.

Distribution: *COL.

Type: Colombia, Williams s.n. (holotype, W).

Dracula incognita Luer & R.Escobar, Amer. Orchid Soc. Bull. 58(10): 1006-1007. 1989.

Illustration: Luer 1993, pl. 43.

Distribution: *COL.

Type: Colombia, Escobar 4008 (holotype, JAUM, not found; isotype, MO).

Dracula insolita Luer & R.Escobar, Amer. Orchid Soc. Bull. 58: 1007. 1989.

Illustration: Luer 1993, pl. 44.

Distribution: *COL.

Type: Colombia, Escobar 3992 (holotype, JAUM, not found; isotype, MO).

Dracula irmelinae N.Peláez, Gary Mey. & L.Mazariegos, Lankesteriana 20(2): 138. 2020.

Illustration: Peláez et al. 2020, f. 1-4, f. 5A-C. Fig. 4D.

Distribution: *COL.

Type: Colombia, JDF-078 (holotype, JAUM).

Dracula lehmanniana Luer & R.Escobar, Selbyana 7(1): 66. 1982.

Illustration: Luer 1993, pl. 48.

Distribution: *COL.

Type: Colombia, Luer 2982 (holotype, SEL).

Dracula lemurella Luer & R.Escobar, Orquideología 15: 23. 1981.

Illustration: Luer & Escobar 1981b, p. 23-25; Luer 1993, pl. 49; Gil & Vieira-Uribe 2019, f. 26.

Distribution: *COL.

Type: Colombia, Escobar 1540 (holotype, JAUM; isotype, SEL).

Dracula levii Luer, Selbyana 2(4): 369. 1978.

Illustration: Luer 1993, pl. 50.

Distribution: COL ECU.

Type: Ecuador, Luer 3455 (holotype, SEL).

Voucher: Colombia, Escobar s.n. (JAUM).

Dracula ligiae Luer & R.Escobar, Amer. Orchid Soc. Bull. 58: 1008. 1989.

Illustration: Luer 1993, pl. 51.

Distribution: *COL.

Type: Colombia, Escobar 3979 (holotype, JAUM, not found; isotype, MO).

Dracula mantissa Luer & R.Escobar, Amer. Orchid Soc. Bull. 48(7): 709. 1979.

Illustration: Luer 1993, pl. 54.

Distribution: COL ECU.

Type: Ecuador, Luer 3966 (holotype, SEL; isotype, JAUM).

Voucher: Colombia, Escobar 1946 (JAUM).

Dracula minax Luer & R.Escobar, Orquideología 15: 27. 1981.

Illustration: Luer & Escobar 1981b, p. 28-30; Luer 1993, pl. 56.

Distribution: *COL.

Type: Colombia, Escobar 682 (holotype, JAUM; isotype, SEL).

Dracula niesseniae P.Ortiz, Orquideología 19(3): 3-4. 1994.

Illustration: Ortiz 1994, p. 5, p. 7 (bottom); Luer 1996d, p. 12; Luer 1994, pl. 108.

Distribution: *COL.

Type: Colombia, Ortiz 1059 (holotype, HPUJ; isotype, MO).

Dracula nosferatu Luer & R.Escobar, Amer. Orchid Soc. Bull. 58(10): 1009-1010. 1989.

Illustration: Luer 1993, pl. 60.

Distribution: *COL.

Type: Colombia, Escobar 3977 [Luer ill. 13013] (holotype, JAUM; isotypes, K-000815825, MO149020)

Dracula nycterina (Rchb.f.) Luer, Selbyana 2: 196. 1978. Masdevallia chimaera André, Ill. Hort. 20: 25, t. 117, 118. 1873, nom. illeg. Masdevallia nycterina Rchb.f., Gard. Chron. 1238. 1873. Non Masdevallia chimaera Rchb.f. [1872].

Illustration: Luer 1993, pl. 61.

Distribution: *COL.

Type: Colombia, Wallis s.n. (holotype, W).

Dracula octavioi Luer & R.Escobar, Orquideología 13: 128. 1979.

Illustration: Luer & Escobar 1979, p. 131; Luer 1993, pl. 62.

Distribution: *COL.

Type: Colombia, Escobar 2001 (holotype, JAUM).

Note: The protologue indicates an isotype at SEL, but apparently that bears a different collection number (Luer 3092) and therefore would be a paratype.

Dracula ophioceps Luer & R.Escobar, Orquideología 13(1): 26. 1978.

Illustration: Luer & Escobar 1978, p. 28, p. 29, p. 58(top right); Luer 1993, pl. 63.

Distribution: *COL.

Type: Colombia, Escobar 1298 (holotype, JAUM).

Dracula orientalis Luer & R.Escobar, Selbyana 7: 67. 1982.

Illustration: Escobar 1983, p. 88; Luer 1993, pl. 64.

Distribution: *COL.

Type: Colombia, Escobar & Portillo 7935 (holotype, SEL; isotype, COL).

Dracula ortiziana Luer & R.Escobar, Orquideología 19(1): 40. 1993.

Illustration: Luer & Escobar 1993b, p. 41, p. 42; Luer 1993, pl. 65.

Distribution: *COL.

Type: Colombia, Escobar 4085 (holotype, JAUM, not found; isotype, MO).

Dracula pholeodytes Luer & R.Escobar, Selbyana 7: 68. 1982.

Illustration: Luer 1993, pl. 67.

Distribution: *COL.

Type: Colombia, Luer 1979 (holotype, SEL, isotype, JAUM, not found).

Dracula pileus Luer & R.Escobar, Orquideología 13: 132. 1979.

Illustration: Luer & Escobar 1979, p. 134, p. 135, p. 170 (bottom right); Luer 1993, pl. 68.

Distribution: *COL.

Type: Colombia, Escobar 1880 (holotype, JAUM).

Note: The protologue indicates an isotype at SEL, but apparently that bears a different collection number (Luer 2930) and therefore would be a paratype.

Dracula posadarum Luer & R.Escobar, Orquideología 15: 31. 1981.

Illustration: Luer & Escobar 1981b, p. 32-34; Luer 1993, pl. 72.

Distribution: *COL.

Type: Colombia, Escobar 1904 (holotype, JAUM; isotype, SEL).

Dracula presbys Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 46: 172. 1993.

Illustration: Luer 1993, pl. 73.

Distribution: *COL.

Type: Colombia, Luer 14092 (holotype, MO).

Dracula psittacina (Rchb.f.) Luer & R.Escobar, Thes. Dracul. 6: 10. 1993. Masdevallia psittacina Rchb.f., Gard. Chron., n.s. 5: 817. 1876.

Illustration: Luer 1993, pl. 74.

Distribution: *COL.

Type: Colombia, Lawrence s.n. (holotype, W).

Dracula × radio-syndactyla Luer, Selbyana 5: 389. 1981.

Distribution: *COL.

Type: Colombia, Luer et al. 6314 (holotype, SEL).

Note: A putative hybrid between D. radiosa and D. syndactyla.

Dracula radiosa (Rchb.f.) Luer, Selbyana 2: 197. 1978. Masdevallia radiosa Rchb.f., Gard. Chron. 1: 684. 1877.

Illustration: Luer 1993, pl. 79.

Distribution: *COL.

Type: Colombia, Wallis 652 (holotype, W).

Dracula rojasii N.Peláez, Buitr.-Delg. & Gary Mey., Orquideología 26(1): 25. 2009.

Illustration: Peláez et al. 2009, f. 1, f. 2.

Distribution: *COL.

Type: Colombia, Peláez NP0285 (holotype, JAUM).

Note: Peláez NP0286 cited as isotype is a paratype.

Dracula robledorum (P.Ortiz) Luer & R.Escobar, Selbyana 2: 197. 1978. Masdevallia chimaera var. robledorum P.Ortiz, Orquideología 10: 222. 1975.

Illustration: Luer & Escobar 1978, p. 32, p. 33, p. 63 (bottom left); Luer 1993, pl. 82.

Distribution: *COL.

Type: Colombia, Ortiz V 715 (holotype, COL; isotype, HPUJ).

Dracula roezlii (Rchb.f.) Luer, Selbyana 2: 197. 1978. Masdevallia chimaera var. roezlii (Rchb.f.) H.J.Veitch, Man. Orchid. Pl. 5: 31. 1889. Masdevallia roezlii Rchb.f., Xenia Orchid. 2: 196, t. 186, II. 1874.

Illustration: Luer 1993, pl. 83.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W).

Dracula senex-furens N.Peláez, Buitr.-Delg. & Gary Mey., Orquideología 26(1): 30. 2009.

Illustration: Peláez et al. 2009, f. 3-5, f. 6 (right).

Distribution: *COL.

Type: Colombia, Peláez NP0287 (holotype, JAUM-44183).

Note: Peláez NP0288 cited as isotype is a paratype.

Dracula sergioi Luer & R.Escobar, Orquideología 13: 35. 1978.

Illustration: Luer & Escobar 1978, p. 36, p. 37, p. 58 (top left); Luer 1993, pl. 84.

Distribution: *COL.

Type: Colombia, Escobar 1402 (holotype, JAUM).

Dracula severa (Rchb.f.) Luer, Selbyana 2: 197. 1978. Masdevallia chimaera var. severa (Rchb.f.) H.J. Veitch, Man. Orchid. Pl. 5: 31. 1889. Masdevallia severa Rchb.f., Gard. Chron. 1: 170. 1875.

Illustration: Luer 1993, pl. 85.

Distribution: *COL.

Type: Colombia, Patin 10 (holotype, W).

Dracula sibundoyensis Luer & R.Escobar, Orquideología 13: 137. 1979.

Illustration: Luer & Escobar 1979, p. 139; Luer 1993, pl. 86.

Distribution: *COL.

Type: Colombia, Escobar 2002 (holotype, JAUM).

Note: The protologue indicates an isotype at SEL, but apparently that bears a different collection number (Luer 3091) and therefore would be a paratype.

Dracula spectrum (Rchb.f.) A.Doucette, Phytotaxa 57: 59. 2012. Masdevallia spectrum Rchb.f., Gard. Chron. 1: 429. 1875.

Illustration: Williams & Williams 1894, p. 496; Luer 1993, pl. 68.

Distribution: *COL.

Type: Colombia, Shuttleworth s.n. (holotype, W).

Dracula syndactyla Luer, Selbyana 5(3-4) 389. 1981.

Illustration: Luer 1993, pl. 89.

Distribution: *COL.

Type: Colombia, Luer 6313 (holotype, SEL).

Dracula trichroma (Schltr.) Hermans, Orchid Rev. 105(1217): 298. 1997. Masdevallia trichroma Schltr., Repert. Spec. Nov. Regni Veg. 6: 57. 1919. Masdevallia tricolor Rchb.f., Gard. Chron. n.s., 18: 102. 1882, nom. illeg. Dracula tricolor (Rchb.f.) Luer & R.Escobar, Selbyana 7: 65. 1982, nom. illeg. Non Masdevallia tricolor Rchb.f. [1850].

Illustration: Luer 1993, pl. 45. Fig. 4E.

Distribution: COL ECU.

Type: Colombia, Roezl s.n. (holotype, W).

Note: For a clarification of the use of Dracula trichroma for what has been mistakenly named Dracula iricolor see Karremans et al. (2019).

Dracula tsubotae Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 102. 2002.

Illustration: Luer 2002a, f. 8 (p. 116).

Distribution: *COL.

Type: Colombia, Barrow RO8 (holotype, MO).

Dracula veleziana Luer & V.N.M.Rao, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 276. 2005.

Illustration: Luer 2005, f. 3 (p. 280).

Distribution: *COL.

Type: Colombia, Rao 171 (holotype, MO).

Dracula velutina (Rchb.f.) Luer, Selbyana 2: 198. 1978. Masdevallia velutina Rchb.f., Gard. Chron. 2: 420. 1875.

Illustration: Luer 1993, pl. 94.

Distribution: *COL.

Type: Colombia, Wallis s.n. (holotype, W).

Dracula venefica Luer & R.Escobar, Orquideología 15: 39. 1981.

Illustration: Luer & Escobar 1981b, p. 40-42; Luer 1993, pl. 95.

Distribution: *COL.

Type: Colombia, Escobar 1903 (holotype, JAUM; isotype, SEL).

Dracula venosa (Rolfe) Luer, Selbyana 2: 198. 1978. Masdevallia venosa Rolfe, Bull. Misc. Inform. Kew 146. 1901.

Illustration: Luer 1993, pl. 96

Distribution: COL ECU.

Type: Colombia, Sander s.n. (holotype, K).

Dracula verticulosa Luer & R.Escobar, Amer. Orchid Soc. Bull. 58(10): 1010-1011. 1989.

Illustration: Luer 1993, pl. 97; Peláez et al. 2020, f. 5D-F.

Distribution: *COL.

Type: Colombia, Escobar 4000 (holotype, JAUM, not found; isotype, MO).

Dracula vespertilio (Rchb.f.) Luer, Selbyana 2(2-3): 198. 1978. Masdevallia vespertilio Rchb.f., Bot. Zeitung (Berlin) 31(25): 390-391. 1873.

Illustration: Luer 1993, pl. 98.

Distribution: COL COS NIC.

Type: Colombia, Wallis s.n. [lectotype (selected here), W-R-0039187].

Dracula villegasiiKöniger, Arcula 8: 218. 1999.

Illustration: Königer 1999, p. 219.

Distribution: *COL.

Type: Colombia, Königer WK-91 (holotype, M; isotypes, K, JAUM, Herb. Königer).

Dracula vinacea Luer & R.Escobar, Orquideología 13: 39. 1978.

Illustration: Luer & Escobar 1978, p. 40, p. 41, p. 58 (bottom left); Luer 1993, pl. 99. Fig. 4F.

Distribution: *COL.

Type: Colombia, Escobar 968 (holotype, JAUM; isotype, SEL).

Dracula vlad-tepes Luer & R.Escobar, Orquideología 15: 43. 1981.

Illustration: Luer & Escobar 1981b, p. 44-47; Luer 1993, pl. 100.

Distribution: *COL.

Type: Colombia, Luer 2180 (holotype, JAUM; isotype, SEL).

Dracula wallisii (Rchb.f.) Luer, Selbyana 2: 198. 1979. Masdevallia chimaera var. wallisii (Rchb.f.) H.J.Veitch, Man. Orchid. Pl. 5: 31. 1889. Masdevallia wallisii Rchb.f., Gard. Chron. 2: 258. 1875.

Illustration: Luer 1993, pl. 101.

Distribution: *COL.

Type: Colombia, Wallis s.n. (holotype, W).

Dracula xenos Luer & R.Escobar, Amer. Orchid Soc. Bull. 58: 1011. 1989.

Illustration: Luer 1993, pl. 103.

Distribution: *COL.

Type: Colombia, Luer 13012 (holotype, MO).

Dresslerella Luer

Dresslerella hispida (L.O.Williams) Luer, Selbyana 3(1): 4-6, f. 105. 1976. Pleurothallis hispida L.O.Williams, Ann. Missouri Bot. Gard. 27(3): 277-278, t. 33, f. 1-4. 1940.

Illustration: Luer 1988, pl. 4 (p. 11). Fig. 4G.

Distribution: COL COS NIC PAN.

Type: Panama, Allen 782 (holotype, AMES).

Voucher: Colombia, Luer 2662 (SEL).

Dresslerella stellaris Luer & R.Escobar, Selbyana 2(2-3): 188, f. 305. 1978.

Illustration: Luer 1988, pl. 8 (p. 19).

Distribution: COL COS.

Type: Colombia, Luer 2261 (holotype, JAUM; isotype, SEL).

Dryadella Luer

Dryadella crassicaudataLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 14, f. 9. 2005.

Illustration: Luer 2005, f. 9.

Distribution: COL PER.

Type: Peru, Luer 11650 (holotype, MO).

Voucher: Colombia, Luer 6519 (SEL).

Dryadella cristata Luer & R.Escobar, Orquideología 15: 126. 1982.

Illustration: Luer & Escobar 1982, p. 126, p. 127; Luer 2005, f. 11.

Distribution: *COL.

Type: Colombia, Escobar 1981 (holotype, JAUM; isotype, SEL).

Dryadella cuspidata Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 160-161. 1999.

Illustration: Luer 2005, f. 12.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 13977 (holotype, MO).

Voucher: Colombia, Luer 2957 (SEL).

Dryadella dodsonii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 161-162. 1999.

Illustration: Luer 2005, f. 13.

Distribution: COL ECU.

Type: Ecuador, Dodson 18798 (holotype, MO).

Voucher: Colombia, Otero & Angulo 1003 (CUVC, MO).

Dryadella gnoma (Luer) Luer, Selbyana 2(2-3): 208. 1978. Pleurothallis gnoma Luer, Selbyana 3(1-2): 110. 1976.

Illustration: Luer & Escobar 1979, p. 142, p. 143, p. 169 (bottom); Luer 2005, f. 19a, f. 19b.

Distribution: COL COS ECU PAN PER.

Type: Ecuador, Luer et al. 698 (holotype, SEL).

Voucher: Colombia, Escobar 1451 (JAUM).

Dryadella guatemalensis (Schltr.) Luer, Selbyana 2(2- 3): 208. 1978. Masdevallia guatemalensis Schltr., Repert. Spec. Nov. Regni Veg. 15(427/433): 201. 1918. Masdevallia simula var. guatemalensis (Schltr.) Kraenzl., Repert. Spec. Nov. Regni Veg. 34: 191. 1925.

Illustration: Luer 2005, f. 22; Karremans et al. 2016, f. 8k.

Distribution: BRA COL COS GUA MEX.

Type: Guatemala, von Tuerkheim II 490 ‘Aug. 1903’ (holotype, B, destroyed. Guatemala, von Tuerkheim II 490 ‘Sept. 1906’ [neotype (selected here), W-R-0012244].

Voucher: Colombia, Luer 1873 (SEL).

Note: Luer (2005) suggested the holotype of this taxon was at W-R, with isotypes at AMES, BR and G. These specimens indeed represent this taxon and have the same collection number, but all have different preparation dates. The actual holotype was destroyed at B and strictly speaking BR0000009973565, AMES-00101256, G-00168643 and W-R-0012244 are not isotypes and cannot be used as lectotypes. The latter is selected as neotype.

Dryadella meiracyllium (Rchb.f.) Luer, Selbyana 2: 209. 1978. Masdevallia meiracyllium Rchb.f., Flora 69: 558. 1886.

Illustration: Luer 2005, f. 29.

Distribution: COL ECU.

Type: Colombia, Lehmann 3367 (holotype, W).

Dryadella minuscula Luer & R.Escobar, Orquideología 13: 43. 1978.

Illustration: Luer & Escobar 1978, p. 44, p. 45, p. 60 (bottom); Luer 2005, f. 30.

Distribution: *COL.

Type: Colombia, Escobar 1516 (holotype, JAUM).

Dryadella mocoana Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 28, f. 31. 2005.

Illustration: Luer 2005, f. 31; Jiménez et al. 2021b, f. 1.

Distribution: COL ECU.

Type: Colombia, Luer 2957 (holotype, SEL).

Dryadella odontostele Luer, Lindleyana 11(2): 54, f. 1. 1996.

Illustration: Luer 2005, f. 34.

Distribution: COL COS PAN.

Type: Panama, Gentry 6178 (holotype, MO).

Voucher: Colombia, Luer 1874 (SEL).

Dryadella pusiola (Rchb.f.) Luer, Selbyana 2(2/3): 209. 1978. Masdevallia pusiola Rchb.f., Gard. Chron. 1: 140. 1887.

Illustration: Luer 2005, f. 36.

Distribution: COL ECU.

Type: Colombia, Wendland 85 (holotype, W).

Dryadella rodrigoi Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 163-164. 1999.

Illustration: Luer 2005, f. 37.

Distribution: *COL.

Type: Colombia, Luer 1920 (holotype, SEL).

Dryadella simula (Rchb.f.) Luer, Selbyana 2: 209. 1978. Masdevallia simula Rchb.f., Gard. Chron. n.s., 3: 8. 1875.

Illustration: Luer 2005, f. 38a, f. 38b.

Distribution: COL ECU PER.

Type: Colombia, Chesterton sub Veitch 342 (holotype, W-R-0050982; illustr., W-R-0050981).

Dryadella verrucosa Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 164. 1999.

Illustration: Luer 2005, f. 45. Fig. 4H.

Distribution: *COL.

Type: Colombia, Luer et al. 10091 (holotype, MO).

Dryadella zebrina (Porsch) Luer, Selbyana 2: 209. 1978. Masdevallia zebrina Porsch, Oesterr. Bot. Z. 154. 1905.

Illustration: Luer 2005, f. 49a, f. 49b, f. 49c.

Distribution: BOL BRA COL PER.

Type: Brazil, von Wettstein & Schiffner s.n. [holotype, B, destroyed; lectotype (selected here), WU0074776; isolectotype, WU-0074777].

Voucher: Colombia, Ortiz s.n. (SEL).

Echinosepala Pridgeon & M.W.Chase

Echinosepala aspasicensis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 17(2): 101. 2002. Brenesia aspasicensis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004. Echinella aspasicensis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001. Humboldtia aspasicensis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Myoxanthus aspasicensis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 15: 38. 1986. Pleurothallis aspasicensis Rchb.f., Bonplandia (Hannover) 3: 73. 1855.

Illustration: Luer 1992, pl. 45.

Distribution: BOL BRA COL ECU GUY PAN VEN.

Type: Colombia, Wagener 618 (holotype, W; isotype, K-L, a single flower which appears to be lost).

Echinosepala isthmica Pupulin, Harvard Pap. Bot. 25: 172. 2020.

Illustration: Luer 1992, pl. 47; Pupulin et al. 2017b, f. 4p-s, 6; Misas Urreta 2005, p. 438, p. 439; Karremans & Vieira-Uribe 2020, p. 131, p. 132.

Distribution: COL COS NIC PAN.

Type: Costa Rica, Bogarín et al. 8571 (holotype, JBL-spirit).

Voucher: Colombia, Misas Urreta 100 (HPUJ).

Echinosepala lappiformis (A.H.Heller & L.O.Williams) Pridgeon & M.W.Chase, Lindleyana 17(2): 101. 2002. Brenesia lappiformis (A.H.Heller & L.O.Williams) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004. Echinella lappiformis (A.H.Heller & L.O.Williams) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001, nom. illeg. Myoxanthus lappiformis (A.H.Heller & L.O.Williams) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 15: 38. 1986. Pleurothallis lappiformis A.H.Heller & L.O.Williams, Fieldiana, Bot. 31(2): 42, f. 8. 1964.

Illustration: Luer 1992, pl. 42; Pupulin et al. 2017b, f. 1; Misas Urreta 2005, p. 434, p. 435. Fig. 4I; Karremans & Vieira-Uribe 2020, p. 144.

Distribution: COL COS ECU NIC PAN VEN.

Type: Nicaragua, Heller 6620 (holotype, F).

Voucher: Colombia, Misas Urreta 244 (HPUJ).

Frondaria Luer

Frondaria colombiana Szlach., Kolan. & Rykacz., Turkish J. Bot. 42(6): 774. 2018.

Illustration: Szlachetko et al. 2018, f. 3. Fig. 5.

Distribution: *COL.

Type: Colombia, Pennell 9270 (holotype, AMES; drawing, UGDA).

Note: We agree with Szlachetko et al. (2018) that the Colombian specimens previously determined as Frondaria caulescens represent a different species.

Gravendeelia Bogarín & Karremans

Gravendeelia chamaelepanthes (Rchb.f.) Bogarín & Karremans, Phytotaxa 340(2): 130. 2018. Humboldtia chamaelepanthes (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis chamaelepanthes Rchb.f., Bonplandia (Hannover) 3: 240. 1855. Trichosalpinx chamaelepanthes (Rchb.f.) Luer, Phytologia 54(5): 394. 1983. Tubella chamaelepanthes (Rchb.f.) Archila, Revista Guatemal. 3(1): 52. 2000 [2009]. Tubella chamaelepanthes (Rchb.f.) Archila, Revista Guatemal. 17(1): 50. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 35, f. 36; Karremans & Vieira-Uribe 2020, p. 151, p. 152. Fig. 4J.

Distribution: COL ECU PER.

Type: Ecuador, Jameson 313 (holotype, W-R0025570; isotypes, BM-000082422, BR-0000006594534, G-00169107, G-00169108, K-000583708, LE-00001409, P-00485473; tracing of type, K-L).

Voucher: Colombia, Cuatrecasas 18821 (AMES; COL).

Karma Karremans

Karma acremona (Luer) Karremans, comb. nov. Bas.: Pleurothallis acremona Luer, Selbyana 5(2): 157. 1979. Trichosalpinx acremona (Luer) Luer, Phytologia 54(5): 394. 1983. Tubella acremona (Luer) Archila, Revista Guatemal. 3(1): 46, 48. 2000 [2009]. Tubella acremona (Luer) Archila, Revista Guatemal. 17(1): 48. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 25.

Distribution: BOL COL ECU.

Type: Ecuador, Luer 596 (holotype, SEL).

Voucher: Colombia, Luer 14111 (MO).

Karma alabastra (Luer & R.Escobar) Karremans, Harvard Pap. Bot. 28(1): 62. 2023. Pleurothallis alabastra Luer & R.Escobar, Orquideología 16: 17. 1983. Trichosalpinx alabastra (Luer & R.Escobar) Luer, Phytologia 54(5): 394. 1983. Tubella alabastra (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 49. 2000 [2009]. Tubella alabastra (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 48. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1983, p. 19; Luer 1997, f. 26.

Distribution: *COL.

Type: Colombia, Luer et al. 7614 (holotype, SEL).

Karma amygdalodora (Kraenzl.) Karremans, Harvard Pap. Bot. 28(1): 62. 2023. Pleurothallis amygdalodora Kraenzl., Bot. Jahrb. Syst. 37: 521. 1906. Trichosalpinx amygdalodora (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 43. 1997. Tubella amygdalodora (Kraenzl.) Archila, Revista Guatemal. 3(1): 49. 2000 [2009]. Tubella amygdalodora (Kraenzl.) Archila, Revista Guatemal. 17(1): 48. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1981a, p. 169; Luer 1997, f. 27.

Distribution: COL ECU PER.

Type: Peru, Weberbauer 1824 (holotype, B, destroyed). Peru, Hutchison & von Bismarck 6425 [neotype, AMES; isoneotypes, F, K, M, MICH, MO, NY, P, UC, US, USM (Luer 1997)].

Voucher: Colombia, Luer 3047 (SEL, JAUM).

Karma arbuscula (Lindl.) Karremans, Harvard Pap. Bot. 28(1): 63. 2023. Humboldtia arbuscula (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis arbuscula Lindl., Edwards’s Bot. Reg. 28: Misc. 72-73. 1842. Trichosalpinx arbuscula (Lindl.) Luer, Phytologia 54(5): 394. 1983. Tubella arbuscula (Lindl.) Archila, Revista Guatemal. 3(1): 49. 2000 [2009]. Tubella arbuscula (Lindl.) Archila, Revista Guatemal. 17(1): 49. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 28, f. 29.

Distribution: BOL COL COS ECU PAN PER VEN.

Type: Ecuador, Hartweg 1408 (holotype, K; isotypes: AMES, LD, W).

Voucher: Colombia, Luer 14150 (MO).

Karma ballatrix (Luer & R.Escobar) Karremans, Harvard Pap. Bot. 28(1): 63. 2023. Trichosalpinx ballatrix Luer & R.Escobar, Orquideología 16(2): 178. 1984. Tubella ballatrix (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 50. 2000 [2009]. Tubella ballatrix (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 49. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1984b, p. 179, p. 180; Luer 1997, f. 103.

Distribution: *COL.

Type: Colombia, Luer et al. 8748 (holotype, SEL).

Karma dura (Lindl.) Karremans, Harvard Pap. Bot. 28(1): 64. 2023. Humboldtia dura (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis dura Lindl., Fol. Orchid. ~Pleurothallis~ (9): 32. 1859. Trichosalpinx dura (Lindl.) Luer, Phytologia 54(5): 395. 1983. Tubella dura (Lindl.) Archila, Revista Guatemal. 3(1): 54. 2000 [2009]. Tubella dura (Lindl.) Archila, Revista Guatemal. 17(1): 51. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 44, f. 45; Stenzel 2007, f. 12.

Distribution: BOL BRA COL COS CUB DOM ECU ELS GUA HAI HON JAM L.A. MEX PAN PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K-L; isotypes, AMES-00074225, W-R-45903).

Voucher: Colombia, Luer 13194 (MO).

Karma escobarii (Luer) Karremans, Harvard Pap. Bot. 28(1): 65. 2023. Trichosalpinx escobarii Luer, Orquideología 18(2): 168-170. 1991. Trichosalpinx escobarii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 114-115. 1997, nom. superfl. Tubella escobarii (Luer) Archila, Revista Guatemal. 3(1): 55. 2000 [2009]. Tubella escobarii (Luer) Archila, Revista Guatemal. 17(1): 52. 2014 [2015], nom. superfl.

Illustration: Luer 1991f, p. 169; Luer 1997, f. 104.

Distribution: *COL.

Type: Colombia, Munera & Escobar 607 (holotype, JAUM; isotype, MO).

Karma hypocrita (Garay & Dunst.) Karremans, Harvard Pap. Bot. 28(1): 65. 2023. Pleurothallis hypocrita Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 344. 1976. Trichosalpinx hypocrita (Garay & Dunst.) Luer, Phytologia 54(5): 395. 1983. Tubella hypocrita (Garay & Dunst.) Archila, Revista Guatemal. 3(1): 57. 2000 [2009]. Tubella hypocrita (Garay & Dunst.) Archila, Revista Guatemal. 17(1): 53. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 105.

Distribution: COL VEN.

Type: Venezuela, Tillett 746-682 (holotype, AMES).

Voucher: Colombia, Cuatrecasas & Romero 25183 (US).

Karma intricata (Lindl.) Karremans, Harvard Pap. Bot. 28(1): 65. 2023. Humboldtia intricata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis intricata Lindl., Orchid. Linden. 1. 1846. Trichosalpinx intricata (Lindl.) Luer, Phytologia 54(5): 396. 1983. Tubella intricata (Luer) Archila, Revista Guatemal. 3(1): 57. 2000 [2009]. Tubella intricata (Luer) Archila, Revista Guatemal. 17(1): 53. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 53.

Distribution: COL ECU PER VEN.

Type: Venezuela, Linden 668 (holotype, K-L; isotypes, AMES-00074356, BR-0000008434869, G-00169076, P-00485638, W-R-12712).

Voucher: Colombia, Luer Silverstone-Sopkin et al. 1740 (CUVC, MO).

Karma multicuspidata (Rchb.f.) Karremans, Harvard Pap. Bot. 28(1): 66. 2023. Pleurothallis multicuspidata Rchb.f., Linnaea 41: 117. 1877. Trichosalpinx multicuspidata (Rchb.f.) Luer, Phytologia 54(5): 396. 1983. Tubella multicuspidata (Rchb.f.) Archila, Revista Guatemal. 3(1): 60. 2000 [2009]. Tubella multicuspidata (Rchb.f.) Archila, Revista Guatemal. 17(1): 54. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 62.

Distribution: COL ECU VEN.

Type: Colombia, Wallis s.n. (holotype, W-R-25694; isotypes, AMES-00074481, AMES-00074482).

Karma pusilla (Kunth) Karremans, Harvard Pap. Bot. 28(1): 67. 2023. Humboldtia pusilla (Kunth) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pusilla (Kunth) Lindl., Edwards’s Bot. Reg. 28: Misc. 82-83. 1842. Specklinia pusilla (Kunth) Lindl., Edwards’s Bot. Reg. 21: sub. t. 1797. 1835. Trichosalpinx pusilla (Kunth) Luer, Phytologia 54(5): 397. 1983. Tubella pusilla (Kunth) Archila, Revista Guatemal. 3(1): 62. 2000 [2009]. Tubella pusilla (Kunth) Archila, Revista Guatemal. 17(1): 56. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 72.

Distribution: COL COS ECU GUY PER VEN.

Type: Ecuador, Bonpland s.n. (holotype, P-00669678; isotype, W-R-30981; photo of type, AMES-00085705).

Voucher: Colombia, Luer 14347 (MO).

Karma robledorum (Luer) Karremans, Harvard Pap. Bot. 28(1): 67. 2023. Pleurothallis robledorum Luer, Orquideología 14: 170. 1981. Trichosalpinx robledorum (Luer) Luer, Phytologia 54(5): 397. 1983. Tubella robledorum (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 63. 2000 [2009]. Tubella robledorum (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 56. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1981a, p. 171; Luer 1997, f. 74.

Distribution: COL ECU.

Type: Colombia, Luer 2293 (holotype, SEL; isotype, JAUM).

Karma scabridula (Rolfe) Karremans, Harvard Pap. Bot. 28(1): 67. 2023. Pleurothallis scabridula Rolfe, Mem. Torrey Bot. Club 4: 260. 1895. Trichosalpinx scabridula (Rolfe) Luer, Phytologia 54(5): 397. 1983. Tubella scabridula (Rolfe) Archila, Revista Guatemal. 3(1): 64. 2000 [2009]. Tubella scabridula (Rolfe) Archila, Revista Guatemal. 17(1): 56. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 75, f. 76.

Distribution: BOL COL PER.

Type: Bolivia, Bang 907 (holotype, K-000583703; isotypes, AMES-00074724, BR-0000006572570, G-00169106, GH-00074722, GH-00074723, LE-00001429, MO-176782, NDG-16158, NY00009238, PH-00023399, US-00446960, US-00093690, W-R-9266).

Voucher: Colombia, Ortiz-Valdivieso 829 (HPUJ).

Karma tenuis (C.Schweinf.) Karremans, Harvard Pap. Bot. 28(1): 68. 2023. Pleurothallis tenuis C.Schweinf., Bot. Mus. Leafl. 10: 190, t. 18, f. 8-14. 1942. Trichosalpinx tenuis (C.Schweinf.) Luer, Phytologia 54(5): 397. 1983. Tubella tenuis (C.Schwweinf.) Archila, Revista Guatemal. 3(1): 66. 2000 [2009]. Tubella tenuis (C.Schwweinf.) Archila, Revista Guatemal. 17(1): 57. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1984b, p. 187; Luer 1997, f. 84.

Distribution: BOL COL ECU PER.

Type: Peru, Herrera 2157 (holotype, US).

Voucher: Colombia, Luer & Escobar 8488 (SEL, COL, JAUM).

Karma vagans (Garay & Dunst.) Karremans, Harvard Pap. Bot. 28(1): 68. 2023. Pleurothallis vagans Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 372. 1976. Trichosalpinx vagans (Garay & Dunst.) Luer, Phytologia 54(5): 398. 1983. Tubella vagans (Garay & Dunsterville) Archila, Revista Guatemal. 3(1): 67. 2000 [2009]. Tubella vagans (Garay & Dunsterville) Archila, Revista Guatemal. 17(1): 58. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 89.

Distribution: *COL.

Type: Colombia, Garay & Sweet 1066 (holotype, AMES).

Karma webbiae (Luer & R.Escobar) Karremans, Harvard Pap. Bot. 28(1): 68. 2023. Trichosalpinx webbiae Luer & R.Escobar, Orquideología 16(2): 188. 1984. Tubella webbiae (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 67. 2000 [2009]. Tubella webbiae (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 58. 2014 [2015], nom. superfl.

Illustration: Luer & Escobar 1984b, p. 189; Luer 1997, f. 90.

Distribution: *COL.

Type: Colombia, Luer et al. 8765 (holotype, SEL).

Lankesteriana Karremans

Lankesteriana barbulata (Lindl.) Karremans, Lankesteriana 13(3): 326. 2014. Anathallis barbulata (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 247. 2001. Humboldtia barbulata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Panmorphia barbulata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 146-148. 2006. Pleurothallis barbata H.Focke, Bot. Zeitung (Berlin) 11(13): 227. 1853, nom. illeg. Pleurothallis barbulata Lindl., Fol. Orchid. ~Pleurothallis~ 40. 1859. Specklinia barbulata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004. Non Pleurothallis barbata Westc. [1841].

Illustration: Foldats 1968, f. 26; Luer 1975a, f. 2 (p. 61); Barringer 1986, f. 13; Luer 2006, f. 102a; Misas Urreta 2005, p. 521; Karremans & Vieira-Uribe 2020, p. 153-155, p. 159, p. 160. Fig. 4K.

Distribution: BOL COL COS ECU FRE GUA GUY HON MEX PER SUR VEN.

Type: Suriname, Focke s.n. (holotype, K-L00079871, illustration).

Voucher: Colombia, Misas Urreta 105 (HPUJ).

Lankesteriana caudatipetala (C.Schweinf.) Karremans, Lankesteriana 13(3): 327. 2014. Anathallis caudatipetala (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 257. 2009. Panmorphia caudatipetala (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 150. 2006. Pleurothallis caudatipetala C.Schweinf., Bot. Mus. Leafl. 10(7): 175-176, t. 22. 1942. Specklinia caudatipetala (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004.

Illustration: Luer 2006, f. 106; Karremans & Vieira-Uribe 2020, p. 158. Fig. 4L.

Distribution: BOL COL COS ECU PER.

Type: Peru, Schunke 1307 (holotype, F).

Voucher: Colombia, Gutierrez & Moreno 74 (JBB).

Lankesteriana cuspidata (Luer) Karremans, Lankesteriana 13(3): 327. 2014. Anathallis cuspidata (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 248. 2001. Panmorphia cuspidata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 152. 2006. Pleurothallis cuspidata Luer, Selbyana 3(3-4): 282-283, f. 241. 1977. Specklinia cuspidata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004.

Illustration: Luer 2006, f. 109; Karremans & Vieira-Uribe 2020, p. 158, p. 160.

Distribution: COL COS ECU PAN.

Type: Panama, Williams & Stevenson 1335 (holotype, SEL).

Voucher: Colombia, Ospina & Idrobo 152 (AMES, COL).

Lankesteriana millipeda (Luer) Karremans, Lankesteriana 13(3): 327. 2014. Anathallis millipeda (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 259. 2009. Panmorphia millipeda (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 166-167. 2006. Pleurothallis millipeda Luer, Orquideología 20: 216-217. 1996. Specklinia millipeda (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 1996f, 215; Luer 2006, f. 132.

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K).

Lankesteriana minutissima (Luer) Karremans, Orchids (West Palm Beach) 84: 310. 2015. Pleurothallis minutissima Luer, Orquideología 20: 218. 1996. Specklinia minutissima (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 1996f, 219; Luer 2006, f. 102b.

Distribution: *COL.

Type: Colombia, Lehmann 8204 (holotype, K).

Lankesteriana muricaudata (Luer) Karremans, Lankesteriana 13(3): 327. 2014. Anathallis muricaudata (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 249. 2001. Panmorphia muricaudata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 168-169. 2006. Pleurothallis muricaudata Luer, Selbyana 7(1): 119. 1982. Specklinia muricaudata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 2006, f. 135a, f. 135b; Kolanowska et al. 2011, p. 33, p. 34; Karremans & Vieira-Uribe 2020, p. 158.

Distribution: BOL COL ECU PAN.

Type: Ecuador, Hirtz & Leon s.n. (holotype, SEL).

Voucher: Colombia, Luer 14747 (MO).

Lepanthes Sw.

Lepanthes abortiva Luer & R.Escobar, Orquideología 18(1): 22-24. 1991.

Illustration: Luer & Escobar 1991a, p. 23, p. 31(top); Luer & Thoerle 2012a, f. 1.

Distribution: *COL.

Type: Colombia? [uncertain origin], Luer 14221 (holotype, MO).

Note: Described from a cultivated specimen without collection data. Probably a synonym of Lepanthes rafaeliana Pupulin.

Lepanthes acarina Luer, Phytologia 54(5): 326. 1983.

Illustration: Luer 1996b, f. 2 (p. 185); Vásquez & Ibisch 2000, pl. 12; Luer & Thoerle 2012a, f. 2.

Distribution: BOL COL ECU PER.

Type: Ecuador, Luer et al. 4399 (holotype, SEL).

Voucher: Colombia, Luer 8810 (SEL).

Lepanthes aciculifolia Luer, Orquideología 20(2): 194-196. 1996.

Illustration: Luer 1996f, 195; Luer & Thoerle 2012a, f. 3.

Distribution: *COL.

Type: Colombia, Luer 15536 (holotype, MO).

Lepanthes acrogenia Luer & R.Escobar, Orquideología 18(1): 24. 1991.

Illustration: Luer & Escobar 1991a, p. 25; Luer & Thoerle 2012a, f. 4; Baquero et al. 2019, f. 4c.

Distribution: *COL.

Type: Colombia, Luer et al. 4624 (holotype, SEL).

Lepanthes aculeata Luer, Phytologia 54: 327. 1983.

Illustration: Luer 1996b, f. 2. Fig. 6A.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3203 (holotype, SEL).

Voucher: Colombia, Moreno 363 (CAUP).

Lepanthes acutissimaLuer & R.Escobar, Orquideología 19(2): 76-77. 1994.

Illustration: Luer & Escobar 1994, f. 81 (p. 79), f. 81 (p. 83); Luer & Thoerle 2012a, f. 5.

Distribution: *COL.

Type: Colombia, Luer et al. 16831 (holotype, MO).

Lepanthes aduncata Luer & R.Escobar, Orquideología 19(1): 31-32. 1993.

Illustration: Luer & Escobar 1993a, p. 17 (left), p. 20 (top left); Luer & Thoerle 2012a, f. 6.

Distribution: *COL.

Type: Colombia, Escobar 4060 (holotype, JAUM; isotype, MO).

Lepanthes affinis Luer & R.Escobar, Orquideología 17(3): 145-148. 1988.

Illustration: Luer & Escobar 1988, p. 147; Luer & Thoerle 2012a, f. 7.

Distribution: *COL.

Type: Colombia, Luer et al. 10331 (holotype, MO).

Lepanthes agatarumJ.S. Moreno, N. Gut. & S.Vieira-Uribe, Phytotaxa 376(2): 92. 2018.

Illustration: Moreno et al. 2018, f. 1, f. 2, f. 3a, f. 4a. Fig. 6B.

Distribution: *COL.

Type: Colombia, Gutiérrez Morales 012 (holotype, JBB).

Lepanthes aggeris Luer & R.Escobar, Orquideología 19(1): 32-33. 1993.

Illustration: Luer & Escobar 1993a, p. 17 (right), p. 20 (top middle); Luer & Thoerle 2012a, f. 8.

Distribution: *COL.

Type: Colombia, Escobar 5244 (holotype, JAUM; isotype, MO).

Lepanthes agglutinata Luer, Phytologia 54(5): 327. 1983.

Illustration: Luer 1996b, f. 7 (p. 186); Luer & Thoerle 2012a, f. 9.

Distribution: COL ECU.

Type: Ecuador, D Alessandro 165 (holotype, SEL).

Voucher: Colombia, Luer 10071 (MO, SEL).

Lepanthes aguirrei Luer, Selbyana 30: 4. 2009.

Illustration: Luer & Thoerle 2012a, f. 10; Baquero et al. 2018, f. 3c.

Distribution: *COL.

Type: Colombia, J&L 1296-536 (holotype, MO).

Lepanthes alcicornis Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(4): 368. 1984.

Illustration: Luer & Thoerle 2012a, f. 11; VieiraUribe & Larsen 2014a, f. 4 (left), Restrepo et al. 2022, f. 3c.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2646 (holotype, SEL).

Lepanthes alkaia Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(5): 580-582. 1985.

Illustration: Luer & Thoerle 2012a, f. 12.

Distribution: *COL.

Type: Colombia, Luer et al. 10142 (holotype, MO).

Lepanthes alvarezii P.Ortiz, Orquideología 24: 115. 2006.

Illustration: Ortiz 2006, p. 117 (below), p. 119; Luer & Thoerle 2012a, f. 13.

Distribution: *COL.

Type: Colombia, Álvarez & Ortiz 1247 (holotype, HPUJ).

Lepanthes amplior Luer & R.Escobar, Orquideología 18(1): 26-2f8. 1991.

Illustration: Luer & Escobar 1991a, p. 27, p. 31(bottom); Luer & Thoerle 2012a, f. 14.

Distribution: *COL.

Type: Colombia, Luer et al. 10267 (holotype, MO).

Lepanthes amplisepalaLuer & R.Escobar, Orquideología 19(2): 77-78. 1994.

Illustration: Luer & Escobar 1994, f. 82 (p. 83); Luer & Thoerle 2012a, f. 15.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8430 (holotype, SEL).

Lepanthes anatinaLuer & R.Escobar, Orquideología 20(3): 281-283, f. 111. 1997.

Illustration: Luer & Escobar 1997, f. 111 (p. 282); Luer & Thoerle 2012a, f. 16.

Distribution: *COL.

Type: Colombia, Luer et al. 16847 (holotype, MO).

Lepanthes anchicayae J.S.Moreno & S.Vieira-Uribe, Harvard Pap. Bot. 25(1): 100. 2020.

Illustration: Moreno et al. 2020b, f. 2, 3, 4a, 5a. Fig. 6C.

Distribution: *COL.

Type: Colombia, Moreno & Erazo 429 (holotype, CAUP).

Lepanthes anchorifera Luer, Revista Soc. Boliv. Bot. 4(1): 3. 2003.

Illustration: Luer 2003a, f. 1, Luer & Thoerle 2012a, f. 17.

Distribution: *COL.

Type: Colombia, Niessen & Uribe 087 (holotype, MO).

Lepanthes andrenoglossa Rchb.f., Bonplandia (Hannover) 3: 241. 1855.

Illustration: Luer 1996b, f. 13 (p. 188); Luer & Thoerle 2012a, f. 18.

Distribution: COL ECU.

Type: Ecuador, Jameson 66 (holotype, W-R0054544; isotype, BM-000084579).

Voucher: Colombia, Luer 12532 (MO).

Lepanthes anemica Luer & Thoerle, Harvard Pap. Bot. 16(2): 326, 20. 2011.

Illustration: Luer & Thoerle 2012a, f. 19.

Distribution: *COL.

Type: Colombia, Thoerle 142 (holotype, MO).

Lepanthes ankistra Luer & Dressler, Orquideología 16(3): 12 (1986).

Illustration: Luer & Dressler 1986a, p. 13, p. 14; Fernández et al. 2014, f. 2, f. 14b; Moreno et al. 2022b, f. 6a. Fig. 6D.

Distribution: COL COS PAN.

Type: Panama, Luer et al. 10534 (holotype, MO).

Voucher: Colombia, Vieira 395 (JAUM).

Lepanthes anserina Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(3): 258. 1984.

Illustration: Luer & Thoerle 2012a, f. 20.

Distribution: *COL.

Type: Colombia, Luer et al. 8920 (holotype, SEL; isotypes, COL, JAUM).

Lepanthes antennataLuer & R.Escobar, Orquideología 19(2): 81. 1994.

Illustration: Luer & Escobar 1994, f. 83 (p. 79), f. 83 (p. 83); Luer & Thoerle 2012a, f. 21.

Distribution: *COL.

Type: Colombia, Escobar 5064 (holotype, JAUM; isotype, MO).

Lepanthes antennifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(1): 52-54. 1985.

Illustration: Luer & Thoerle 2012a, f. 22.

Distribution: *COL.

Type: Colombia, Luer et al. 10288 (holotype, MO; isotype JAUM).

Lepanthes antioquiensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 95. 1920.

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 32, No. 122)].

Note: An obscure taxon excluded by Luer.

Lepanthes aperta Luer, Selbyana 30: 5. 2009.

Illustration: Luer 2009b, f. 8; Luer & Thoerle 2012a, f. 23.

Distribution: *COL.

Type: Colombia, Luer 19668 (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes aquila-borussiae Rchb.f., Bonplandia (Hannover) 2: 22. 1854.

Illustration: Luer & Thoerle 2012a, f. 24.

Distribution: COL VEN.

Type: Venezuela, Wagener s.n. (holotype, W-R0054546).

Voucher: Colombia, Luer 10202 (MO).

Lepanthes arbuscula Luer & R.Escobar, Orquideología 19(1): 33-34. 1993.

Illustration: Luer & Escobar 1993a, p. 22 (left), p. 20 (top right); Luer & Thoerle 2012a, f. 25.

Distribution: *COL.

Type: Colombia, Escobar 5009 (holotype, JAUM; isotype, MO).

Lepanthes argentataLuer & R.Escobar, Orquideología 17(3): 150-153. 1988.

Illustration: Luer & Escobar 1988, p. 149 (bottom), p. 151; Luer & Thoerle 2012a, f. 26.

Distribution: *COL.

Type: Colombia, Escobar 3274 (holotype, MO).

Lepanthes aries Luer, Phytologia 54(5): 329. 1983.

Illustration: Luer 1996b, f. 16 (p. 188); Luer & Thoerle 2012a, f. 27.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6044 (holotype, SEL).

Voucher: Colombia, Thoerle 138 (MO).

Lepanthes aristata Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(5): 582-584. 1985.

Illustration: Luer & Thoerle 2012a, f. 28.

Distribution: *COL.

Type: Colombia, Luer et al. 10332 (holotype, JAUM; isotype, MO).

Lepanthes astraea S.Vieira-Uribe & B.T.Larsen, Orquideología 31(1): 5-6. 2014.

Illustration: Vieira-Uribe & Larsen 2014a, f. 1, f. 3, f. 4 (right). Fig. 6E.

Distribution: *COL.

Type: Colombia, Vieira 0017 (holotype, JAUM).

Lepanthes athena Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 38, f. 18. 1996.

Illustration: Luer 1996b, f. 18 (p. 189); Luer & Thoerle 2012a, f. 29.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9933 (holotype, MO).

Voucher: Colombia, Dodson 18050 (MO).

Lepanthes atomifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(1): 26. 1984.

Illustration: Luer & Thoerle 2012a, f. 30.

Distribution: *COL.

Type: Colombia, Luer et al. 8965 (holotype, SEL).

Lepanthes auditorLuer & R.Escobar, Orquideología 17(3): 154. 1988.

Illustration: Luer & Escobar 1988, p. 149(top), p. 155; Luer & Thoerle 2012a, f. 31.

Distribution: *COL.

Type: Colombia, Escobar 3417 (holotype, JAUM; isotype, MO).

Lepanthes aures-asini Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(2): 158. 1984.

Illustration: Luer & Thoerle 2012a, f. 32.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2540 (holotype, SEL).

Lepanthes auriculata Luer, Phytologia 54(5): 330. 1983.

Illustration: Luer 1996b, f. 19 (p. 189); Luer & Thoerle 2012a, f. 33.

Distribution: BOL COL ECU.

Type: Ecuador, D’Alessandro 164 (holotype, SEL).

Voucher: Colombia, Silverstone-Sopkin 3786 (CUVC).

Lepanthes bachue S.Vieira-Uribe, J.S.Moreno & E.Parra, Lankesteriana 23(1):129. 2023.

Illustration: Vieira-Uribe et al. [Lankesteriana, in review], f. 2-5.

Distribution: *COL.

Type: Colombia, Parra-Sánchez et al. 2521 (holotype, VALLE).

Lepanthes ballatrix Luer, Phytologia 54(5): 331. 1983.

Illustration: Luer 1996b, f. 22 (p. 190); Luer & Thoerle 2012a, f. 34; Tobar Suarez et al. 2021, f. 12d.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 10699 (holotype, MO).

Voucher: Colombia, Luer 14102 (MO).

Lepanthes barclaya Luer & Thoerle, Harvard Pap. Bot. 16(2): 327. 2011.

Illustration: Luer & Thoerle 2012a, f. 35.

Distribution: *COL.

Type: Colombia, Barclay & Juajibioy 6070 (holotype, MO; isotype US).

Lepanthes beatrizaeLuer & R.Escobar, Orquideología 20(3): 283-284. 1997.

Illustration: Luer & Escobar 1997, f. 112 (p. 282), p. 300 (top right); Luer & Thoerle 2012a, f. 36.

Distribution: *COL.

Type: Colombia, Escobar 5382 (holotype, MO).

Lepanthes biglomerisLuer & R.Escobar, Orquideología 19(1): 34-35. 1993.

Illustration: Luer & Escobar 1993a, p. 22 (right), p. 20 (bottom left); Luer & Thoerle 2012a, f. 37.

Distribution: *COL.

Type: Colombia, Escobar 5122 (holotype, JAUM; isotype, MO).

Lepanthes biloba Lindl., Companion Bot. Mag. 2: 356. 1836.

Illustration: Luer 1996b, f. 26 (p. 191), f. 156 (p. 223); Luer & Thoerle 2012a, f. 38.

Distribution: COL ECU.

Type: Ecuador, Hall s.n. (holotype, K-L).

Voucher: Colombia, Luer 14224 (MO).

Lepanthes binaria Luer & Hirtz, Orchidee (Hamburg) 38(6): 291. 1987.

Illustration: Luer 1996b, f. 27 (p. 191); Luer & Thoerle 2012a, f. 39.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11080 (holotype, MO).

Voucher: Colombia, Luer 4651 (SEL).

Lepanthes bipinnatulaLuer & R.Escobar, Orquideología 20(3): 284-286. 1997.

Illustration: Luer & Escobar 1997, f. 113 (p. 282); Luer & Thoerle 2012a, f. 40.

Distribution: *COL.

Type: Colombia, Luer et al. 17653 (holotype, MO).

Lepanthes bitriangularis Luer & R.Escobar, Selbyana 30: 5. 2009.

Illustration: Luer & Thoerle 2012a, f. 41.

Distribution: *COL.

Type: Colombia, Escobar 5220 (holotype, MO).

Lepanthes botivana .S.Vieira-Uribe & J.SMoreno, Orquideología 37(1): 18. 2020.

Illustration: Vieira-Uribe & Moreno 2020, f. 1-3. Fig. 6F.

Distribution: *COL.

Type: Colombia, Vieira & Alfonso 051 (holotype, JAUM).

Lepanthes boyacensis Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(8): 820. 1984.

Illustration: Luer & Thoerle 2012a, f. 42.

Distribution: *COL.

Type: Colombia, Luer 8777 (holotype, SEL).

Lepanthes brevipedicellata Luer, R.Escobar & Thoerle, Harvard Pap. Bot. 16(2): 323. 2011.

Illustration: Luer & Thoerle 2012a, f. 43.

Distribution: *COL.

Type: Colombia, Luer 10041 (holotype, MO).

Lepanthes cacique-toneLuer & R.Escobar, Orquideología 20(3): 286-287. 1997.

Illustration: Luer & Escobar 1997, f. 114 (p. 282), p. 301 (bottom left); Luer & Thoerle 2012a, f. 44.

Distribution: *COL.

Type: Colombia, Escobar s.n. (holotype, JAUM; isotype, MO).

Note: The holotype is Escobar s.n. according to the protologue, but Luer & Thoerle (2012a) indicate the type is Luer et al. 17641 (holotype, JAUM; isotype, MO).

Lepanthes cactoura Luer & R.Escobar, Orquideología 18(1): 28. 1991.

Illustration: Luer & Escobar 1991a, p. 29; Luer & Thoerle 2012a, f. 45; Vieira-Uribe & Moreno 2019, f. 9e.

Distribution: *COL.

Type: Colombia, Escobar 3399 (holotype, MO).

Lepanthes caesariataLuer & R.Escobar, Orquideología 19(2): 82. 1994.

Illustration: Luer & Escobar 1994, f. 84 (p. 83); Luer & Thoerle 2012a, f. 46; Vieira-Uribe & Moreno 2022, f. 5b.

Distribution: *COL.

Type: Colombia, Luer 16733 (holotype, MO).

Lepanthes calimae P.Ortiz, Orquideología 21: 69. 1998.

Illustration: Ortiz 1998, p. 69, p. 70; Luer & Thoerle 2012a, f. 47; Moreno 2019a, f. 30.

Distribution: *COL.

Type: Colombia, Ortiz et al. 1090 (holotype, COL).

Lepanthes callisto Luer & Hirtz, Orchidee (Hamburg) 38(6): 287-289. 1987.

Illustration: Luer 1996b, f. 32 (p. 192); Luer & Thoerle 2012a, f. 48.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11076 (holotype, SEL).

Voucher: Colombia, Luer 17949 (MO).

Lepanthes calochromaThoerle & Hirtz, Phytotaxa 201: 52. 2015

Illustration: Thoerle & Hirtz 2015, f. 1, f. 2. Fig. 6G.

Distribution: COL ECU.

Type: Ecuador, Hirtz 10397 (holotype, MO).

Voucher: Colombia, Moreno 389 (CAUP).

Note: Described from a cultivated specimen without collection data.

Lepanthes calodictyon Hook., Bot. Mag. t. 5259. 1861.

Illustration: Luer 1996b, f. 33 (p. 193); Luer & Thoerle 2012a, f. 49.

Distribution: COL ECU.

Type: Ecuador, Spruce s.n. (holotype, K).

Voucher: Colombia, Escobar 5166 (MO).

Lepanthes campodostele Luer & Hirtz, Novon 3: 444. 1993.

Illustration: Luer 1996b, f. 36 (p. 193). Fig. 6H.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 13472 (holotype, MO).

Voucher: Colombia, Moreno 530 (CUVC).

Lepanthes camptica Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 50. 1996.

Illustration: Luer 1996b, f. 37 (p. 194). Fig. 6I.

Distribution: COL ECU.

Type: Ecuador, Hirtz et al. 4662 (holotype, MO).

Voucher: Colombia, Gutierrez 076 (JBB).

Lepanthes canaliculataLuer & R.Escobar, Orquideología 20(3): 287-289. 1997.

Illustration: Luer & Escobar 1997, f. 115 (p. 290); Luer & Thoerle 2012a, f. 50.

Distribution: *COL.

Type: Colombia, Luer 16739 (holotype, MO).

Lepanthes cardenasii J.S.Moreno, Gal.-Tar. & Zuluaga, Lankesteriana 22(2): 86. 2022.

Illustration: Moreno et al. 2022b, f. 1, f. 2, f. 3a. Fig. 6K.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona et al. 1485 (holotype, CUVC-spirit).

Lepanthes carinata Luer & Hirtz, Orchidee (Hamburg) 41: 163. 1990.

Illustration: Luer 1996b, f. 39 (p. 194). Fig. 6J.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11833 (holotype, MO).

Voucher: A photo of a Colombian specimen is reproduced herein (Fig. 6J).

Lepanthes carrilloi Luer & Thoerle, Harvard Pap. Bot. 16(2): 329, 23. 2011.

Illustration: Luer & Thoerle 2012a, f. 51.

Distribution: *COL.

Type: Colombia, Thoerle 104 (holotype, MO).

Lepanthes carrizosana Gal.-Tar., Zuluaga & J.S.Moreno, Harvard Pap. Bot. 28(1): 89-92 2023.

Illustration: Moreno et al. 2023b, f. 1, f. 2.

Distribution: *COL.

Type: Colombia, R. Galindo-Tarazona et al. 1586 (holotype, CUVC).

Lepanthes carunculigera Rchb.f., Flora 69: 557. 1886.

Illustration: Luer & Thoerle 2012a, f. 52.

Distribution: *COL.

Type: Colombia, Lehmann 3213 (holotype, W-R0054517).

Lepanthes caudata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(3): 260. 1984.

Illustration: Luer 1996b, f. 42 (p. 195); Luer & Thoerle 2012a, f. 53.

Distribution: COL ECU.

Type: Colombia, Luer & Escobar 8345 (holotype, SEL).

Lepanthes caudatisepala C.Schweinf., Bot. Mus. Leafl. 10: 123, t. 15. 1942.

Illustration: Luer 1996b, f. 43 (p. 195); Luer & Thoerle 2012a, f. 54.

Distribution: BOL COL ECU PER.

Type: Peru, Macbride 4273 (holotype, F).

Voucher: Colombia, Luer 14101 (MO).

Lepanthes celox Luer & Hirtz, Lindleyana 2: 36. 1987.

Illustration: Luer 1996b, f. 44 (p. 195). Fig. 6L.

Distribution: COL ECU.

Type: Ecuador, Hirtz 1461 (holotype, MO).

Voucher: Colombia, Moreno 347 (CAUP).

Lepanthes cerambyx Luer & R.Escobar, Orquideología 18(1): 32-34. 1991.

Illustration: Luer & Escobar 1991a, p. 33, p. 39(top); Luer & Thoerle 2012a, f. 55.

Distribution: *COL.

Type: Colombia, Escobar et al. 3382 (holotype, JAUM; isotypes, COL, MO).

Lepanthes cercion Luer & R.Escobar, Orquideología 17(3): 158-161. 1987.

Illustration: Luer & Escobar 1988, p. 159; Luer & Thoerle 2012a, f. 56.

Distribution: COL VEN.

Type: Colombia, Luer et al. 11355 (holotype, MO).

Note: holotype originally SEL, corrected to MO (Luer 2012).

Lepanthes chelonion Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(1): 54-56. 1985.

Illustration: Moreno et al. 2018, f. 3c; Luer & Thoerle 2012a, f. 57.

Distribution: *COL.

Type: Colombia, Luer et al. 10215 (holotype, MO).

Lepanthes chimaera Luer & R.Escobar, Orquideología 16(2): 130. 1984.

Illustration: Luer & Escobar 1984a, p. 131; Luer 1996b, f. 45 (p. 196); Luer & Thoerle 2012a, f. 58.

Distribution: COL ECU.

Type: Colombia, Luer et al. 4633 (holotype, SEL).

Lepanthes chocoensis Luer & Thoerle, Harvard Pap. Bot. 16(2): 330, f. 24. 2011.

Illustration: Luer & Thoerle 2012a, f. 59.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 1636 (holotype, MO; isotype CUVC).

Lepanthes cincinnata Luer & R.Escobar, Orquideología 19(1): 35-36. 1993.

Illustration: Luer & Escobar 1993a, p. 23 (left), p. 20 (middle left); Luer & Thoerle 2012a, f. 60.

Distribution: *COL.

Type: Colombia, Escobar 5081 (holotype, JAUM; isotype, MO).

Lepanthes cingensLuer & R.Escobar, Orquideología 19(2): 87-88. 1994.

Illustration: Luer & Escobar 1994, f. 86 (p. 79), f. 86 (p. 84); Luer & Thoerle 2012a, f. 61.

Distribution: *COL.

Type: Colombia, Luer et al. 16783 (holotype, MO).

Lepanthes cissyana S.Vieira-Uribe & J.S.Moreno, Lankesteriana 19(2): 68. 2019.

Illustration: Vieira-Uribe & Moreno 2019, f. 5, f. 6. Fig. 7B.

Distribution: *COL.

Type: Colombia, Moreno et al. 520 (holotype, JAUM; isotype CAUP).

Lepanthes clausa Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(6): 728. 1985.

Illustration: Gutiérrez Morales et al. 2021, f. 5a; Luer & Thoerle 2012a, f. 62.

Distribution: *COL.

Type: Colombia, Luer et al. 10274 (holotype, MO; isotype JAUM).

Lepanthes climaxLuer & R.Escobar, Orquideología 17(3): 162. 1988.

Illustration: Luer & Escobar 1988, p. 163; Luer & Thoerle 2012a, f. 63.

Distribution: *COL.

Type: Colombia, Luer & Escobar 7870 (holotype, SEL).

Lepanthes cocculifera Luer & R.Escobar, Orquideología 16(2): 133. 1984.

Illustration: Luer & Escobar 1984a, p. 134; Luer & Thoerle 2012a, f. 64.

Distribution: *COL.

Type: Colombia, Luer & Escobar 7866 (holotype, SEL).

Lepanthes cogolloi Luer & R.Escobar, Orquideología 18(1): 34,36. 1991.

Illustration: Luer & Escobar 1991a, p. 35; Luer & Thoerle 2012a, f. 65.

Distribution: *COL.

Type: Colombia, Cogollo et al. 3539 (holotype, JAUM; isotype, MO).

Lepanthes conchyliata Luer, Selbyana 30(1): 6. 2009.

Illustration: Luer & Thoerle 2012a, f. 66.

Distribution: *COL.

Type: Colombia, Luer 19658 (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes confusa Ames & C.Schweinf., Sched. Orch. 10: 40-41. 1930.

Illustration: Luer 1996b, f. 53 (p. 198); Luer & Thoerle 2012a, f. 67; Bogarín et al. 2020. f. 8, f. 9.

Distribution: COL COS ECU HON PAN.

Type: Costa Rica, Standley 48709 (holotype, AMES).

Voucher: Colombia, Escobar 5084 (MO).

Lepanthes contingens Luer, Phytologia 54(5): 335. 1983.

Illustration: Luer 1996b, f. 55 (p. 198); Luer & Thoerle 2012a, f. 68;

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7152 (holotype, SEL).

Voucher: Colombia, Luer 4637 (SEL).

Lepanthes cordataLuer & R.Escobar, Orquideología 17(3): 165-168. 1988.

Illustration: Luer & Escobar 1988, p. 167; Luer & Thoerle 2012a, f. 69; Vieira-Uribe et al. 2021, f. 4b.

Distribution: *COL.

Type: Colombia, Luer et al. 11371 (holotype, MO).

Lepanthes cordillerana E.Restrepo, J.S.Moreno & Gal.-Tar., Phytotaxa 603(1): 71. 2023.

Illustration: Parra-Sánchez et al. 2023, f. 1-2.

Distribution: *COL.

Type: Galindo-Tarazona et al. 1454 (holotype, CUVC).

Lepanthes cornualis Luer & R.Escobar, Orquideología 18(1): 36, 79. 1991.

Illustration: Luer & Escobar 1991a, p. 37; Luer 1996b, f. 57 (p. 199); Luer & Thoerle 2012a, f. 70; Moreno et al. 2020a, f. 9A.

Distribution: COL ECU.

Type: Colombia, Luer et al. 12525 (holotype, MO).

Lepanthes corrugata Luer & Dalström, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 63, f. 58. 1996.

Illustration: Luer 1996b, f. 58 (p. 199) Luer & Thoerle 2012a, f. 71.

Distribution: COL ECU.

Type: Ecuador, Dalström et al. 2153 (holotype, MO).

Voucher: Colombia, Ortiz 1414 (HPUJ).

Lepanthes costata Rchb.f., Flora 69: 557. 1886.

Distribution: *COL.

Type: Colombia, unknown collector (holotype, ?).

Note: An obscure name.

Lepanthes cuatrecasasiiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 109, f. 3. 2007.

Illustration: Luer & Thoerle 2012a, f. 72.

Distribution: *COL.

Type: Colombia, Cuatrecasas 20601 (holotype, AMES).

Lepanthes cucullataLuer & R.Escobar, Orquideología 19(2): 89-90. 1994.

Illustration: Luer & Escobar 1994, f. 88 (p. 84); Luer & Thoerle 2012a, f. 73; Gutiérrez Morales et al. 2021, f. 5b.

Distribution: *COL.

Type: Colombia, Luer et al. 10282 (holotype, MO).

Lepanthes culex Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(7): 716. 1984.

Illustration: Restrepo et al. 2022, f. 3d; Luer & Thoerle 2012a, f. 74.

Distribution: *COL.

Type: Colombia, Escobar 3189 (holotype, MO; isotypes, JAUM, SEL).

Lepanthes cuneiformis Luer & R.Escobar, Orquideología 16(2): 135. 1984.

Illustration: Luer & Escobar 1984a, p. 136; Luer & Thoerle 2012a, f. 75.

Distribution: *COL.

Type: Colombia, Luer et al. 7943 (holotype, SEL).

Lepanthes cunicularisLuer & R.Escobar, Orquideología 20(3): 289, 291. 1997.

Illustration: Luer & Escobar 1997, f. 116 (p. 290), p. 300 (bottom left); Luer & Thoerle 2012a, f. 76.

Distribution: *COL.

Type: Colombia, Escobar s.n. (holotype, JAUM; isotype, MO).

Lepanthes cyclochilaLuer & R.Escobar, Orquideología 19(2): 90-91. 1994.

Illustration: Luer & Escobar 1994, f. 89 (p. 79), f. 89 (p. 93); Luer & Thoerle 2012a, f. 77.

Distribution: *COL.

Type: Colombia, Escobar 5241 (holotype, JAUM; isotype, MO).

Lepanthes cymbiumLuer & R.Escobar, Orquideología 19(2): 91. 1994.

Illustration: Luer & Escobar 1994, f. 90 (p. 80), f. 90 (p. 93); Luer & Thoerle 2012a, f. 78; Luer 1996b, f. 65 (p. 201); Moreno et al. 2018, f. 3e.

Distribution: COL ECU.

Type: Colombia, Escobar et al. 4016 (holotype, MO).

Lepanthes cyrtostele Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 102. 1998.

Illustration: Dodson & Luer 2011, f. 5B. Fig. 7A.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 18748 (holotype, MO).

Voucher: Colombia, Cortes 81 (CUVC).

Lepanthes darioiLuer & R.Escobar, Orquideología 20(3): 291-292, f. 117. 1997.

Illustration: Luer & Escobar 1997, f. 117 (p. 290), p. 300 (bottom right); Luer & Thoerle 2012a, f. 79.

Distribution: *COL.

Type: Colombia, Luer et al. 17638 (holotype, MO; isotype JAUM).

Lepanthes dasyura Luer & R.Escobar, Orquideología 18(1): 40-42. 1991.

Illustration: Luer & Escobar 1991a, p. 41; Luer & Thoerle 2012a, f. 80.

Distribution: *COL.

Type: Colombia, Cogollo et al. 3365 (holotype, JAUM; isotype, MO).

Lepanthes davidii Gal.-Tar., Zuluaga & J.S.Moreno, Lankesteriana 22(2): 86. 2022.

Illustration: Moreno et al. 2022b, f. 4, f. 5, f. 6c. Fig. 7C.

Distribution: *COL.

Type: Colombia, Zuluaga et al. 5077 (holotype, CUVC-spirit).

Lepanthes decurva Luer & Hirtz, Orchidee (Hamburg) 39(3): 103. 1988.

Illustration: Luer 1996b, f. 67 (p. 201). Fig. 7D.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11810 (holotype, MO).

Voucher: Colombia, Moreno & Erazo 277 (CAUP).

Lepanthes debedoutii P.Ortiz, Orquideología 24: 116. 2006.

Illustration: Ortiz 2006, p. 117 (above), p. 120; Luer & Thoerle 2012a, f. 81.

Distribution: *COL.

Type: Colombia, Ortiz et al. 1248 (holotype, HPUJ).

Lepanthes debilis Luer & R.Escobar, Orquideología 17(3): 169-172. 1988.

Illustration: Luer & Escobar 1988, p. 171, p. 173(bottom); Luer & Thoerle 2012a, f. 82.

Distribution: *COL.

Type: Colombia, Escobar et al. 3465 (holotype, JAUM; isotype, MO).

Lepanthes declivisLuer & R.Escobar, Orquideología 21(2): 142-145. 1999.

Illustration: Luer & Escobar 1999, p. 143; Pérez-Escobar et al. 2010c, p. 48, p. 49; Luer & Thoerle 2012a, f. 83; Moreno et al. 2022b, f. 12b.

Distribution: *COL.

Type: Colombia, Luer 17713 (holotype, MO).

Lepanthes deficiens Luer & R.Escobar, Orquideología 17(3): 174-178. 1988.

Illustration: Luer & Escobar 1988, p. 173(top), p. 177; Luer & Thoerle 2012a, f. 84.

Distribution: *COL.

Type: Colombia, Escobar 3270 (holotype, MO).

Lepanthes deformis Luer & Hirtz, Orchidee (Hamburg) 38(1): 38. 1987.

Illustration: Luer 1996b, f. 68 (p. 201); Luer & Thoerle 2012a, f. 85.

Distribution: COL COS ECU PAN.

Type: Ecuador, Luer et al. 9857 (holotype, MO).

Voucher: Colombia, Luer 18421 (MO).

Lepanthes delhierroi Luer & Hirtz, Novon 3: 445. 1993.

Illustration: Luer 1996b, f. 70 (p. 202). Fig. 7E.

Distribution: COL ECU.

Type: Ecuador, Hirtz et al. 4965 (holotype, MO).

Voucher: Colombia, Vieira 396 (JAUM).

Lepanthes deliciasensisLuer & R.Escobar, Orquideología 19(2): 95-96. 1994.

Illustration: Luer & Escobar 1994, f. 91 (p. 93); Luer & Thoerle 2012a, f. 86.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8473 (holotype, SEL).

Lepanthes deutera Luer & Thoerle, Selbyana 30(1): 6. 2009.

Illustration: Luer & Thoerle 2012a, f. 87.

Distribution: *COL.

Type: Colombia, Escobar 5347 (holotype, MO).

Lepanthes dewildeiLuer & R.Escobar, Orquideología 20(3): 292-294. 1997.

Illustration: Luer & Escobar 1997, f. 118 (p. 290); Luer & Thoerle 2012a, f. 88.

Distribution: *COL.

Type: Colombia, Luer et al. 16848 (holotype, MO).

Lepanthes diabolica Luer & R.Escobar, Orquideología 18(1): 42. 1991.

Illustration: Luer & Escobar 1991a, p. 43; Luer & Thoerle 2012a, f. 89.

Distribution: *COL.

Type: Colombia, Luer et al. 10158 (holotype, MO).

Lepanthes dicyrtopetala Luer & Hirtz, Selbyana 30(1): 8. 2009.

Illustration: Luer & Thoerle 2012a, f. 90.

Distribution: *COL.

Type: Colombia, Hirtz 8670 (holotype, MO).

Note: Without collection data.

Lepanthes didactyla Luer & R.Escobar, Orquideología 16(2): 137. 1984.

Illustration: Luer & Escobar 1984a, p. 138; Luer & Thoerle 2012a, f. 91.

Distribution: *COL.

Type: Colombia, Luer et al. 7715 (holotype, SEL).

Lepanthes discolorLuer & R.Escobar, Orquideología 17(3): 183-187. 1988.

Illustration: Luer & Escobar 1988, p. 187; Luer & Thoerle 2012a, f. 92.

Distribution: COL PAN.

Type: Colombia, Escobar 3359 (holotype, JAUM; isotypes, COL, MO).

Lepanthes dodsonii Luer, Phytologia 54(5): 340. 1983.

Illustration: Luer 1996b, f. 75 (p. 203); Luer & Thoerle 2012a, f. 93.

Distribution: COL ECU.

Type: Ecuador, Dodson et al. 8475 (holotype, SEL).

Voucher: Colombia, Cogollo et al. 3870 (MO).

Lepanthes dolabrataLuer & R.Escobar, Orquideología 20(3): 294-295. 1997.

Illustration: Luer & Escobar 1997, f. 119 (p. 296); Luer & Thoerle 2012a, f. 94.

Distribution: *COL.

Type: Colombia, Luer et al. 17642 (holotype, MO).

Lepanthes dolichopus Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 96. 1920.

Illustration: Mansfeld 1929, pl. 32, (n. 123).

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 32, No. 123)].

Note: An obscure taxon not monographed by Luer.

Lepanthes dorae J.S.Moreno, Gal.-Tar. & Zuluaga, Lankesteriana 22(2): 93. 2022.

Illustration: Moreno et al. 2022b, f. 7, f. 8, f. 9a. Fig. 7F.

Distribution: *COL.

Type: Colombia, Zuluaga et al. 5078 (holotype, CUVC-spirit).

Lepanthes dougdarlingii S.Vieira-Uribe & J.S.Moreno, Lankesteriana 19(2): 71. 2019.

Illustration: Vieira-Uribe & Moreno 2019, f. 7, f. 8, f. 9a, f. 10a. Fig. 7G.

Distribution: *COL.

Type: Colombia, Moreno et al. 521 (holotype, JAUM; isotype CAUP).

Lepanthes dryades Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(9): 918. 1984.

Illustration: Luer & Thoerle 2012a, f. 95.

Distribution: *COL.

Type: Colombia, Luer et al. 10137 (holotype, MO; isotype SEL).

Lepanthes dumbo Luer, Selbyana 30(1): 9. 2009.

Illustration: Luer & Thoerle 2012a, f. 96.

Distribution: *COL.

Type: Colombia, J&L s.n. (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes dunstervilleorumFoldats, Acta Bot. Venez. 3: 333. 1968.

Illustration: Foldats 1968, f. 9; Luer & Escobar 1984a, p. 148, p. 149; Luer 1996b, f. 76 (p. 203); Luer & Thoerle 2012a, f. 97a, f. 97b.

Distribution: COL ECU VEN.

Type: Venezuela, Steyermark et al. 98525 (holotype, VEN).

Voucher: Colombia, Escobar 5189 (MO).

Lepanthes echidion Luer & R.Escobar, Orquideología 16(2): 139. 1984.

Illustration: Luer & Escobar 1984a, p. 140; Luer & Thoerle 2012a, f. 98.

Distribution: *COL.

Type: Colombia, Luer et al. 7953 (holotype, SEL).

Lepanthes ectopa Luer, Selbyana 30(1): 9. 2009.

Illustration: Luer 2009b, f. 17; Luer & Thoerle 2012a, f. 99.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 7967 (holotype, MO).

Lepanthes edwardsii Ames, Bot. Mus. Leafl. 1(4): 4-8. 1933.

Illustration: Luer & Thoerle 2012a, f. 100; Pupulin 2021, f. 1, f. 2a, f. 3, f. 4.

Distribution: BEL COL COS GUA HON NIC.

Type: Honduras, Edwards 96 (holotype, AMES). Voucher: Colombia, Misas 74 (HPUJ).

Note: Colombian materials are unlikely to fit L. edwardsii sensu Pupulin (2021).

Lepanthes effusa Schltr., Repert. Spec. Nov. Regni Veg. 14: 126. 1915.

Illustration: Luer 1996b, f. 78 (p. 204); Luer & Thoerle 2012a, f. 101.

Distribution: COL ECU.

Type: Ecuador, Sodiro 25 (holotype, B, destroyed). Ecuador, Hirtz 2057 [neotype, MO, Luer (1996)].

Voucher: Colombia, Callejas 7795 (HUA).

Lepanthes elata Rchb.f., Beitr. Orchid.-K. C. Amer. 90-91. 1866.

Illustration: Luer 2001b, p. 43; Luer & Thoerle 2012a, f. 102a, f. 102b.

Distribution: COL COS PAN.

Type: Costa Rica, Wendland 844 (holotype, W-R0054548).

Voucher: Colombia, Otero 274 (CUVC, MO).

Lepanthes elegantula Schltr. Repert. Spec. Nov. Regni Veg. 14: 127. 1915.

Illustration: Luer 1996b, f. 86 (p. 205). Fig. 7H.

Distribution: COL ECU.

Type: Ecuador, Sodiroi 24 [holotype, B, destroyed; lectotype (selected here), QPLS-211080]. Ecuador, Hirtz 2063 [neotype, MO (Luer 1996b), here rejected].

Voucher: Colombia, Moreno 415 (CAUP).

Note: Luer (1996b) mistakenly selected a lectotype for this species based on a specimen collected by Hirtz. He probably intended to select a neotype instead. However, an isotype is available and should be selected as lectotype.

Lepanthes elephantina Luer & R.Escobar, Orquideología 16(2): 141. 1984.

Illustration: Luer & Escobar 1984a, p. 142; Luer 1996b, f. 83 (p. 205); Luer & Thoerle 2012a, f. 103.

Distribution: COL ECU.

Type: Colombia, Luer & Luer 3470 (holotype, SEL).

Lepanthes elongata Luer & Hirtz, Lindleyana 2(2): 99. 1987.

Illustration: Luer 1996b, f. 84 (p. 205); Luer & Thoerle 2012a, f. 104.

Distribution: COL ECU.

Type: Ecuador, Hirtz 1811 (holotype, MO).

Voucher: Colombia, Luer 14225 (MO).

Lepanthes equicalceolata Luer & R.Escobar, Orquideología 19(1): 24. 1993.

Illustration: Luer & Escobar 1993a, p. 23 (right), p. 20 (middle below); Luer 1996b, f. 3 (p. 10); Luer & Thoerle 2012a, f. 105.

Distribution: *COL.

Type: Colombia, Escobar 4059 (holotype, JAUM; isotype, MO).

Lepanthes eros Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(2): 160. 1984.

Illustration: Luer & Thoerle 2012a, f. 106.

Distribution: *COL.

Type: Colombia, Luer et al. 8869 (holotype, SEL).

Lepanthes erythrocles Luer & R.Escobar, Orquideología 18(1): 44. 1991.

Illustration: Luer & Escobar 1991a, p. 45; Luer & Thoerle 2012a, f. 107.

Distribution: *COL.

Type: Colombia, Luer 12808 (holotype, MO).

Lepanthes escifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(1): 28. 1984.

Illustration: Luer & Thoerle 2012a, f. 108.

Distribution: *COL.

Type: Colombia, Luer et al. 8956 (holotype, SEL; isotype COL).

Lepanthes escobarianaGaray, Orquideología 4: 76. 1969.

Illustration: Garay 1969, p. 81 (top right); Luer & Thoerle 2012a, f. 109.

Distribution: *COL.

Type: Colombia, Escobar 531 (holotype, AMES).

Lepanthes eucerca Luer & Thoerle, Harvard Pap. Bot. 16: 332. 2011.

Illustration: Luer & Thoerle 2012a, f. 110.

Distribution: *COL.

Type: Colombia, Luer et al. 10232 (holotype, MO).

Lepanthes exaltata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(4): 370. 1984.

Illustration: Luer & Thoerle 2012a, f. 111.

Distribution: *COL.

Type: Colombia, Luer et al. 6640 (holotype, SEL).

Lepanthes farallonensis Haelterman, Gal-Tar. & Zuluaga, Lankesteriana 21(3): 342. 2021.

Illustration: Baquero et al. 2021, f. 1a, f. 2, f. 3, f. 4a, f. 4c. Fig. 7I.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona & Espitia 1531 (holotype, CUVC-72981).

Lepanthes felis Luer & R.Escobar, Amer. Orchid Soc. Bull. 52(12): 1264. 1983.

Illustration: Luer 1996b, f. 4 (p. 10), f. 5 (p. 11); Luer & Thoerle 2012a, f. 112; Moreno & Vieira-Uribe 2020, f. 43.

Distribution: *COL.

Type: Colombia, Valencia 2605 (holotype, SEL).

Lepanthes ferax Luer & R.Escobar, Orquideología 18(1): 48. 1991.

Illustration: Luer & Escobar 1991a, p. 39(bottom), p. 47; Luer & Thoerle 2012a, f. 113.

Distribution: *COL.

Type: Colombia, Luer et al. 10376 (holotype, MO).

Lepanthes fibulifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(1): 29. 1984.

Illustration: Luer & Thoerle 2012a, f. 114.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2604 (holotype, SEL).

Lepanthes filamentosa Luer & Hirtz, Novon 3(4): 448, f. 8. 1993.

Illustration: Luer 1996b, f. 93 (p. 208); ; Luer & Thoerle 2012a, f. 115; Baquero et al. 2018, f. 4e.

Distribution: COL ECU.

Type: Ecuador, Luer at al. 12378 (holotype, MO).

Voucher: Colombia, Thoerle 141 (MO).

Lepanthes fimbriatilabia J.S.Moreno, Gal.-Tar., Zuluaga, Harvard Pap. Bot. 26(1): 178, 181. 2021.

Illustration: Moreno et al. 2021, f. 4, f. 5, f. 6a. Fig. 7J.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona & Mora 1548 (holotype, CUVC).

Lepanthes florenciana J.S.Moreno & D.Hoyos, Phytotaxa 559(1): 56. 2022.

Illustration: Moreno et al. 2022a, f. 2, f. 3, f. 6, f. 7A. Fig. 7K.

Distribution: *COL.

Type: Colombia, Hoyos et al. 945 (holotype, COAH; isotype HUAZ).

Lepanthes focalis Luer, Phytologia 54(5) 343. 1983.

Illustration: Luer 1996b, f. 99 (p. 209); Luer & Thoerle 2012a, f. 116.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 5524 (holotype, SEL).

Voucher: Colombia, Luer 16830 (MO).

Lepanthes fonnegrae Luer & R.Escobar, Orquideología 19(1): 25. 1993.

Illustration: Luer & Escobar 1993a, p. 20 (middle right), p. 26 (left); Luer & Thoerle 2012a, f. 117.

Distribution: *COL.

Type: Colombia, Escobar 5181 (holotype, MO).

Lepanthes forceps Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(2): 161. 1984.

Illustration: Luer & Thoerle 2012a, f. 118.

Distribution: *COL.

Type: Colombia, Luer et al. 6680 (holotype, SEL).

Lepanthes foreroi P.Ortiz, O.Pérez & E.Parra, Orquideología 26: 137. 2009.

Illustration: Ortiz et al. 2009, p. 138; Luer & Thoerle 2012a, f. 119.

Distribution: *COL.

Type: Colombia, Canizales et al. 244 (holotype, VALLE).

Note: Ortiz et al. 1439 (HPUJ) is cited as isotype but is actually a paratype prepared from the same cultivated plant as the holotype but on a different date.

Lepanthes foveata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(10): 1044. 1984.

Illustration: Luer & Thoerle 2012a, f. 120.

Distribution: *COL.

Type: Colombia, Luer et al. 10308 (holotype, MO; isotype SEL).

Lepanthes furcata Luer & R.Escobar, Orquideología 19(1): 27. 1993.

Illustration: Luer & Escobar 1993a, p. 20 (bottom center), p. 26 (right); Luer & Thoerle 2012a, f. 121.

Distribution: *COL.

Type: Colombia, Escobar 5055 (holotype, JAUM; isotype, MO).

Lepanthes gargantua Rchb.f., Xenia Orchid. 1: 150. 1856.

Illustration: Luer 1996b, f. 103 (p. 210); Luer & Thoerle 2012a, f. 122.

Distribution: COL ECU.

Type: Ecuador, Jameson 110 (holotype, W).

Voucher: Colombia, Luer 14135 (holotype, MO).

Lepanthes gelataLuer & R.Escobar, Orquideología 20: 295. 1997.

Illustration: Luer & Escobar 1997, f. 120 (p. 296); Luer & Thoerle 2012a, f. 123.

Distribution: *COL.

Type: Colombia, Luer et al. 16837 (holotype, MO).

Lepanthes gemina Luer & Hirtz, Amer. Orchid Soc. Bull. 54(3): 314-316. 1985.

Illustration: Luer & Thoerle 2012a, f. 124.

Distribution: *COL.

Type: Colombia, Luer et al. 10340 (holotype, JAUM; isotype, MO).

Lepanthes georgiiLuer & R.Escobar, Orquideología 19(2): 96. 1994.

Illustration: Luer & Escobar 1994, f. 92 (p. 79), f. 92 (p. 93); Luer & Escobar 1999, p. 141; Luer & Thoerle 2012a, f. 125; Restrepo et al. 2022, f. 3e.

Distribution: *COL.

Type: Colombia, Luer 16639 (holotype, JAUM; isotype, MO).

Lepanthes giraldoi Luer, Orquideología 22(1): 37. 2001.

Illustration: Luer 2001b, p. 39; Luer & Thoerle 2012a, f. 126.

Distribution: *COL.

Type: Colombia, Giraldo-Gensini 169 (holotype, TULV; isotype, MO).

Lepanthes gloriaeS.Vieira-Uribe, J.S. Moreno & Pérez-Arcila, Orquideología 38(1): 6. 2021.

Illustration: Vieira-Uribe et al. 2021, f. 1-3, f. 4a. Fig. 7L.

Distribution: *COL.

Type: Colombia, Vieira 055 (holotype, JAUMspirit; isotype HUA).

Lepanthes golondrina Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(7): 718. 1984.

Illustration: Luer & Thoerle 2012a, f. 127.

Distribution: *COL.

Type: Colombia, Escobar 1171 (holotype, JAUM; isotype SEL).

Lepanthes grildrig Luer & R.Escobar, Orquideología 18(1): 49. 1991.

Illustration: Luer & Escobar 1991a, p. 51; Luer & Thoerle 2012a, f. 128.

Distribution: *COL.

Type: Colombia, Luer et al. 10356 (holotype, MO).

Lepanthes guanacasensisLuer & R.Escobar, Orquideología 19(2): 97. 1994.

Illustration: Luer & Escobar 1994, f. 93 (p. 94); Luer & Thoerle 2012a, f. 129.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8419 (holotype, SEL).

Lepanthes gustavoiLuer & R.Escobar, Orquideología 20(3): 298-299. 1997.

Illustration: Luer & Escobar 1997, f. 121 (p. 296); Luer & Thoerle 2012a, f. 130.

Distribution: *COL.

Type: Colombia, Luer et al. 17677 (holotype, MO).

Lepanthes guttula-sanguinisLuer & R.Escobar, Orquideología 17(3): 188-192. 1988.

Illustration: Luer & Escobar 1988, p. 189, p. 193(top); Luer & Thoerle 2012a, f. 131.

Distribution: *COL.

Type: Colombia, Escobar et al. 3460 (holotype, JAUM; isotype, MO).

Note: Originally described as Lepanthes gutulasanguinis.

Lepanthes habenifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(4): 371. 1984.

Illustration: Luer & Thoerle 2012a, f. 132; Vieira-Uribe & Moreno 2019, f. 9f.

Distribution: *COL.

Type: Colombia, Luer et al. 8966 (holotype, SEL).

Lepanthes helgae Luer & R.Escobar, Orquideología 18(1): 52. 1991.

Illustration: Luer & Escobar 1991a, p. 53; Luer & Thoerle 2012a, f. 133.

Distribution: *COL.

Type: Colombia, Luer 14261 (holotype, MO).

Lepanthes hemirhoda Garay, Canad. J. Bot. 34: 250. 1956.

Illustration: Luer & Thoerle 2012a, f. 135.

Distribution: *COL.

Type: Colombia, Cuatrecasas 12458 (holotype, US; isotype COL).

Lepanthes heptapus Luer & R.Escobar, Amer. Orchid Soc. Bull. 52(12): 1266. 1983.

Illustration: Luer & Thoerle 2012a, f. 136.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2601 (holotype, SEL).

Lepanthes heteroloba Luer, R.Escobar & Thoerle, Harvard Pap. Bot. 16: 332. 2011.

Illustration: Luer & Thoerle 2012a, f. 137.

Distribution: *COL.

Type: Colombia, Escobar 3421 (holotype, MO).

Lepanthes hexapus Luer & R.Escobar, Orquideología 16(2): 144. 1984.

Illustration: Luer & Escobar 1984a, p. 145; Luer 1996b, f. 114 (p. 213); Luer & Thoerle 2012a, f. 138; Baquero et al. 2018, f. 3b.

Distribution: COL ECU.

Type: Colombia, Luer et al. 4619 (holotype, SEL).

Lepanthes hirpex Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(5): 584. 1985.

Illustration: Luer & Thoerle 2012a, f. 139.

Distribution: *COL.

Type: Colombia, Escobar 3264 (holotype, JAUM; isotypes, COL, MO).

Lepanthes hirsutula Luer & Hirtz, Lindleyana 2(3): 135. 1987.

Illustration: Luer 1996b, f. 115 (p. 213); ; Luer & Thoerle 2012a, f. 140; Baquero et al. 2019, f. 4e.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12365 (holotype, MO).

Voucher: Colombia, Thoerle 130 (MO).

Lepanthes hirtzii Luer, Phytologia 54(1): 345. 1983.

Illustration: Luer 1996b, f. 116 (p. 213); Luer & Thoerle 2012a, f. 141.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 4394 (holotype, SEL).

Voucher: Colombia, Luer 3118 (SEL).

Lepanthes hortensisLuer & R.Escobar, Orquideología 17(3): 194. 1988.

Illustration: Luer & Escobar 1988, p. 193(bottom), p. 195; Luer & Thoerle 2012a, f. 142; Vieira-Uribe & Moreno 2019, f. 9b, 10b.

Distribution: *COL.

Type: Colombia, Escobar et al. 2727 (holotype, MO).

Lepanthes hubeinii Luer, Orchideen 38(2): 58. 1987.

Illustration: Luer & Thoerle 2012a, f. 143.

Distribution: *COL.

Type: Colombia, Luer 11422 (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes hurgo Luer & Béhar, Revista Soc. Boliv. Bot. 4(1): 7. 2003.

Illustration: Luer 2003a, f. 3; Luer & Thoerle 2012a, f. 144.

Distribution: *COL.

Type: Colombia, Behar 154 (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes hwangiaeJ.S.Moreno & S. Vieira-Uribe, Phytotaxa 567 (2): 149-161. 2022.

Illustration: Vieira et al. 2022, f. 4, 5. Fig. 8A.

Distribution: *COL.

Type: Colombia, Vieira 30 (holotype, JAUM).

Lepanthes hymenoptera Luer & Hirtz, Orchidee (Hamburg) 39: 101 (1988).

Illustration: Luer 1996b, f. 120 (p. 214); VieiraUribe & Moreno 2022, f. 5c. Fig. 10J.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11002 (holotype, MO)

Voucher: Colombia, Perdomo et al. 554 (LAMUA).

Lepanthes hyphosaLuer & R.Escobar, Orquideología 17(3): 199. 1988.

Illustration: Luer & Escobar 1988, p. 201, p. 203(top); Luer & Thoerle 2012a, f. 145.

Distribution: *COL.

Type: Colombia, Escobar 3272 (holotype, MO).

Lepanthes ictalurus Luer, Phytologia 54(5): 346. 1983.

Illustration: Luer 1996b, f. 122 (p. 215); Luer & Thoerle 2012a, f. 146.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7151 (holotype, SEL).

Voucher: Colombia, Luer 12539 (MO).

Note: The Colombian voucher may be L. tetrachaeta instead.

Lepanthes illex Luer, Phytologia 54(5): 347. 1983.

Illustration: Luer 1996b, f. 125 (p. 216); Luer & Thoerle 2012a, f. 147.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6263 (holotype, SEL).

Voucher: Colombia, Moreno 272 (CAUP).

Lepanthes imbricansLuer & R.Escobar, Orquideología 21(2): 147-149. 1999.

Illustration: Luer & Escobar 1999, p. 148; Luer & Thoerle 2012a, f. 148.

Distribution: *COL.

Type: Colombia, Luer 17855 (holotype, MO).

Lepanthes imposita Luer & R.Escobar, Amer. Orchid Soc. Bull. 54: 56. 1985.

Illustration: Luer & Thoerle 2012a, f. 149; Thoerle & Hirtz 2015, f. 4.

Distribution: *COL.

Type: Colombia, Luer et al. 7671 (holotype, SEL).

Lepanthes impotensLuer & R.Escobar, Orquideología 20(3): 299, 302. 1997.

Illustration: Luer & Escobar 1997, f. 122 (p. 296); Luer & Thoerle 2012a, f. 150; Moreno et al. 2021, f. 7b.

Distribution: *COL.

Type: Colombia, Luer et al. 16842 (holotype, MO).

Lepanthes insolitaLuer & R.Escobar, Orquideología 17(3): 204. 1988.

Illustration: Luer & Escobar 1988, p. 203(bottom), p. 207; Luer & Thoerle 2012a, f. 151; Restrepo et al. 2022, f. 3f.

Distribution: *COL.

Type: Colombia, Escobar 3354 (holotype, JAUM; isotype, MO).

Lepanthes intonsa Luer, Phytologia 54(5): 348. 1983.

Illustration: Luer 1996b, f. 129 (p. 217); Luer & Thoerle 2012a, f. 152.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6212 (holotype, SEL).

Voucher: Colombia, Escobar 5205 (MO).

Lepanthes ionoptera Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Illustration: Vásquez & Ibisch 2000, pl. 26; Luer & Thoerle 2012a, f. 153.

Distribution: BOL COL PER VEN.

Type: Colombia, Wagener s.n. (holotype, W).

Lepanthes irmae Gal.-Tar., Zuluaga, J.S.Moreno & Mora-Aguilar, Harvard Pap. Bot. 26(1): 178. 2021.

Illustration: Moreno et al. 2021, f. 1, f. 2. Fig. 8B.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona & Mora 1504 (holotype, CUVC).

Lepanthes isochila Luer, Orquideología 20(2): 196-198. 1996.

Illustration: Luer 1996f, 197; Luer & Thoerle 2012a, f. 154.

Distribution: *COL.

Type: Colombia, Behar 36 (holotype, MO).

Lepanthes isosceles Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(4): 372. 1984.

Illustration: Luer & Thoerle 2012a, f. 155.

Distribution: *COL.

Type: Colombia, Luer 8740 (holotype, SEL).

Lepanthes janitor Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(5): 476. 1984.

Illustration: Luer 1996b, f. 135 (p. 218); Luer & Thoerle 2012a, f. 156.

Distribution: COL ECU.

Type: Colombia, Escobar & Valencia 3197 (holotype, MO; isotypes, COL, JAUM, SEL).

Lepanthes jardinensis Luer & R.Escobar, Orquideología 19(1): 38-39. 1993.

Illustration: Luer & Escobar 1993a, p. 20 (bottom right), p. 29 (left); Luer & Thoerle 2012a, f. 157.

Distribution: *COL.

Type: Colombia, Escobar 2696 (holotype, JAUM; isotypes, MO, SEL).

Lepanthes jostii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 142. 1999.

Illustration: Dodson & Luer 2011, f. 50a. Fig. 8C.

Distribution: COL ECU.

Type: Ecuador, Jost 1137 (holotype, MO).

Voucher: Colombia, Moreno 278 (CAUP).

Lepanthes juan-felipei Luer & Thoerle, Harvard Pap. Bot. 16(2): 333, 28. 2011.

Illustration: Luer & Thoerle 2012a, f. 158.

Distribution: *COL.

Type: Colombia, Thoerle 136 (holotype, MO).

Lepanthes jubata Luer, Phytologia 54(5): 350. 1983.

Illustration: Luer 1996b, f. 138 (p. 219); Luer & Thoerle 2012a, f. 159.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 4447 (holotype, SEL).

Voucher: Colombia, Luer 21050 (MO).

Lepanthes jucas J.S.Moreno & S.Vieira-Uribe, Lankesteriana 20(2): 233. 2020.

Illustration: Moreno et al. 2020a, f. 7, f. 8, f. 9D. Fig. 8D.

Distribution: *COL.

Type: Colombia, Moreno et al. 526 (holotype, JAUM; isotype CAUP).

Lepanthes kokonuko J.S.Moreno & Pisso-Florez, Lankesteriana 20(2): 230. 2020.

Illustration: Moreno et al. 2020a, f. 2-3, f. 6A. Fig. 8E.

Distribution: *COL.

Type: Colombia, Moreno et al. 523 (holotype, JAUM; isotype CAUP).

Lepanthes labiata Luer, Orquideología 22(1): 38-42. 2001.

Illustration: Luer 2001b, p. 41; Luer & Thoerle 2012a, f. 160.

Distribution: *COL.

Type: Colombia, Otero 1004 (holotype, CUVC; isotype, MO).

Lepanthes lacera Luer & Thoerle, Harvard Pap. Bot. 16(2): 333. 2011.

Illustration: Luer & Thoerle 2012a, f. 161.

Distribution: *COL.

Type: Colombia, Cogollo et al. 2639 (holotype, MO).

Lepanthes larvina Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(2): 162. 1984.

Illustration: Luer & Thoerle 2012a, f. 162.

Distribution: *COL.

Type: Colombia, Luer et al. 8984 (holotype, SEL).

Lepanthes lasiopetala Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 178. 1976.

Illustration: Luer & Thoerle 2012a, f. 163.

Distribution: *COL.

Type: Venezuela, Dunsterville 1025 (holotype, AMES).

Lepanthes licrophora Luer & B.T.Larsen, Harvard Pap. Bot. 18: 179. 2013.

Illustration: Luer & Thoerle 2013, f. 8. Fig. 8F.

Distribution: COL ECU.

Type: Ecuador, Larsen 2012 0430-02 (holotype, MO).

Voucher: Colombia, Vieira & Jaramillo 269 (JAUM).

Note: Described from a cultivated specimen without collection data.

Lepanthes lehmannii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 160. 1924.

Distribution: *COL.

Type: Colombia, Lehmann CLVII [holotype, B, destroyed; lectotype (selected here), BM-000084588].

Lepanthes ligiae Luer & R.Escobar, Orquideología 18(1): 56. 1991.

Illustration: Luer & Escobar 1991a, p. 57, p. 65(bottom); Luer & Thoerle 2012a, f. 164.

Distribution: *COL.

Type: Colombia, Escobar 3309 (holotype, MO).

Note: In the protologue, the type collection is given the numbers Escobar 3309 and Luer 10389, with the latter indicated as the holotype. Luer & Thoerle (2012a) indicate that Escobar 3309 is the number of the holotype species and Luer 10389 is the number of Luer’s illustration based on that specimen.

Lepanthes lilliputaeLuer & R.Escobar, Orquideología 17(3): 208. 1988.

Illustration: Luer & Escobar 1988, p. 209; Luer 1996b, f. 108 (p. 211); Luer & Thoerle 2012a, f. 165.

Distribution: COL ECU.

Type: Colombia, Luer et al. 10230 (holotype, MO).

Lepanthes limbataLuer & R.Escobar, Orquideología 19(2): 98. 1994.

Illustration: Luer & Escobar 1994, f. 94 (p. 79), f. 94 (p. 94); Luer & Thoerle 2012a, f. 166.

Distribution: *COL.

Type: Colombia, Luer et al. 16807 (holotype, MO).

Lepanthes linealis Luer & R.Escobar, Orquideología 18(1): 58. 1991.

Illustration: Luer & Escobar 1991a, p. 59; Luer & Thoerle 2012a, f. 167.

Distribution: *COL.

Type: Colombia, Luer 14223 (holotype, MO).

Lepanthes lingulosaLuer & R.Escobar, Orquideología 21(2): 149-151. 1999.

Illustration: Luer & Escobar 1999, p. 150; Luer & Thoerle 2012a, f. 168.

Distribution: *COL.

Type: Colombia, Escobar 5481 (holotype, MO).

Lepanthes lycocephala Luer & R.Escobar, Orquideología 16(2): 147. 1984.

Illustration: Luer & Escobar 1984a, p. 150; Luer 1996b, f. 143 (p. 220); Luer & Thoerle 2012a, f. 169; Vieira-Uribe & Moreno 2019, f. 4b.

Distribution: COL ECU.

Type: Colombia, Luer et al. 4626 (holotype, SEL).

Lepanthes macdougalii Luer & Thoerle, Harvard Pap. Bot. 16(2): 335. 2011.

Illustration: Luer & Thoerle 2012a, f. 170.

Distribution: *COL.

Type: Colombia, Macdougal 4389 (holotype, MO).

Lepanthes macrantha Garay, Canad. J. Bot. 34: 250. 1956.

Illustration: Luer & Thoerle 2012a, f. 171.

Distribution: *COL.

Type: Colombia, Cuatrecasas 12403 (holotype, US; isotype COL).

Lepanthes macrostylisLuer & R.Escobar, Orquideología 21(2): 151-154. 1999.

Illustration: Luer & Escobar 1999, p. 143, p. 153; Luer & Thoerle 2012a, f. 172.

Distribution: *COL.

Type: Colombia, Luer et al. 17672 (holotype, MO).

Lepanthes magnifica Luer, Phytologia 54: 351 (1983).

Illustration: Luer 1996b, f. 145 (p. 221). Fig. 8G.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7333 (holotype, SEL).

Voucher: Colombia, Bonilla 1190 (VALLE).

Lepanthes manabina Dodson, Icon. Pl. Trop. 2: t. 128. 1980.

Illustration: Luer 1996b, f. 147 (p. 221); ; Luer & Thoerle 2012a, f. 173; Baquero et al. 2021, f. 1c.

Distribution: COL ECU.

Type: Ecuador, Dodson & Thien 1725 (holotype, SEL).

Voucher: Colombia, Luer 17695 (MO).

Lepanthes maria-victoriae Luer & Thoerle, Harvard Pap. Bot. 16: 335. 2011.

Illustration: Luer & Thoerle 2012a, f. 174.

Distribution: *COL.

Type: Colombia, Thoerle 139 (holotype, MO).

Lepanthes marielana N.Gutiérrez, J.S.Moreno & S.Vieira-Uribe, Phytotaxa 484(1): 116-117. 2021.

Illustration: Gutiérrez Morales et al. 2021, f. 1, f. 2, f. 5c. Fig. 8H.

Distribution: *COL.

Type: Colombia, Calderón Hernández & Gutiérrez 156 (holotype, JBB).

Lepanthes marthae Luer & R.Escobar, Orquideología 18(1): 60. 1991.

Illustration: Luer & Escobar 1991a, p. 61; Luer & Thoerle 2012a, f. 175.

Distribution: *COL.

Type: Colombia, Munera & Escobar 160 (holotype, JAUM; isotype, MO).

Lepanthes matisii Luer, Thoerle & P.Ortiz, Orchid Digest 75(1): 24-31. 2011.

Illustration: Luer & Thoerle 2012a, f. 176.

Distribution: *COL.

Type: Colombia, Thoerle 106 (holotype, HPUJ).

Lepanthes medinae Luer & Thoerle, Harvard Pap. Bot. 16(2): 336, 32. 2011.

Illustration: Luer & Thoerle 2012a, f. 177.

Distribution: *COL.

Type: Colombia, Thoerle 114 (holotype, MO).

Lepanthes medusa Luer & R.Escobar, Orquideología 18(1): 64. 1991.

Illustration: Luer & Escobar 1991a, p. 63, p. 65(top); Luer & Thoerle 2012a, f. 178.

Distribution: *COL.

Type: Colombia, Luer 14098 (holotype, MO; isotype JAUM).

Lepanthes mefueënsisLuer & R.Escobar, Orquideología 17(3): 212-215. 1988.

Illustration: Luer & Escobar 1988, p. 213; Luer & Thoerle 2012a, f. 179.

Distribution: *COL.

Type: Colombia, Luer et al. 10296 (holotype, MO).

Lepanthes meleagrisLuer & R.Escobar, Orquideología 19(2): 99. 1994.

Illustration: Luer & Escobar 1994, f. 95 (p. 94); Luer & Thoerle 2012a, f. 180.

Distribution: *COL.

Type: Colombia, Luer et al. 16782 (holotype, JAUM; isotype, MO).

Note: Type originally cited as Luer et al. 167821, corrected in Luer & Thoerle (2012a).

Lepanthes menatoi Luer & R.Vásquez Phytologia 55: 189. 1984.

Illustration: Luer 1996b, f. 153 (p. 223). Fig. 8I.

Distribution: BOL COL ECU PER.

Type: Bolivia, Luer 9428 (holotype, MO).

Voucher: Colombia, Sierra-Ariza 101 (VALLE).

Lepanthes micellilabiaLuer & R.Escobar, Orquideología 19(2): 101-102. 1994.

Illustration: Luer & Escobar 1994, f. 96 (p. 79); f. 96 (p. 94); Luer & Thoerle 2012a, f. 181.

Distribution: *COL.

Type: Colombia, Escobar 5191 (holotype, JAUM; isotype, MO).

Lepanthes microcalodyction J.S.Moreno & Baquero, Harvard Pap. Bot. 25(1): 101. 2020.

Illustration: Moreno et al. 2020b, f. 6, 7, 8B, 9B. Fig. 8J.

Distribution: *COL.

Type: Colombia, Moreno & Erazo 432 (holotype, CAUP).

Lepanthes micronyxLuer & R.Escobar, Orquideología 20(3): 302-304. 1997.

Illustration: Luer & Escobar 1997, f. 123 (p. 306); Luer & Thoerle 2012a, f. 182.

Distribution: *COL.

Type: Colombia, Luer et al. 17648 (holotype, MO).

Lepanthes microscopica Luer & R.Escobar, Amer. Orchid Soc. Bull. 52(12): 1268. 1983.

Illustration: Luer & Thoerle 2012a, f. 183.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2598 (holotype, SEL).

Lepanthes mirabilis Ames, Sched. Orch. 1: 12. 1922.

Illustration: Luer & Thoerle 2012a, f. 184.

Distribution: *COL.

Type: Colombia, Pittier 592 (holotype, US).

Note: Described from a cultivated specimen without collection data.

Lepanthes mirador Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 119, f. 157. 1996.

Illustration: Luer 1996b, f. 157 (p. 224); Luer & Thoerle 2012a, f. 185; Tobar Suarez et al. 2021, f. 8b.

Distribution: COL ECU.

Type: Ecuador, Hirtz et al. 4940 (holotype, MO).

Voucher: Colombia, Thoerle 123 (MO).

Lepanthes monitor Luer, Phytologia 54(5): 354. 1983.

Illustration: Luer 1996b, f. 158 (p. 224); Vásquez & Ibisch 2000, f. 16, pl. 32; Luer & Thoerle 2012a, f. 186.

Distribution: BOL COL ECU PER.

Type: Ecuador, Luer et al. 7096 (holotype, SEL).

Voucher: Colombia, Luer 14335 (MO).

Lepanthes monoptera Lindl., J. Bot. (Hooker) 1: 10. 1834.

Illustration: Luer 1996b, f. 159 (p. 224); Luer & Thoerle 2012a, f. 187; Moreno et al. 2020a, f. 9C.

Distribution: BOL COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K; tracing of type, K-L).

Voucher: Colombia, Luer 10301 (MO).

Lepanthes montezumaeJ.S. Moreno & S.Vieira-Uribe, Orquideología 35(2): 177. 2018.

Illustration: Vieira-Uribe & Moreno 2018, f. 2, f. 3, f. 4b. Fig. 8K.

Distribution: *COL.

Type: Colombia, Vieira et al. 24 (holotype, JAUM).

Lepanthes morae Zuluaga, J.S.Moreno & Gal.-Tar., Lankesteriana 22(2): 96. 2022.

Illustration: Moreno et al. 2022b, f. 10, f. 11, f. 12a. Fig. 8L.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona et al. 1558 (holotype, CUVC-spirit).

Lepanthes mucronata Lindl., Companion Bot. Mag. 2: 356. 1836.

Illustration: Luer 1996b, f. 161 (p. 225); Vásquez & Ibisch 2000, pl. 34; Luer & Thoerle 2012a, f. 188.

Distribution: BOL COL ECU PER.

Type: Ecuador, Hall s.n. (holotype, K).

Voucher: Colombia, Luer 16753 (MO).

Lepanthes muscula Luer, Phytologia 54(5): 354. 1983.

Illustration: Luer 1996b, f. 162 (p. 225); Luer & Thoerle 2012a, f. 189.

Distribution: COL ECU.

Type: Ecuador, Luer & Escobar 8300 (holotype, SEL).

Voucher: Colombia, Pasmiño 43 (MO).

Lepanthes myoxophora Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(3): 261. 1984.

Illustration: Luer & Thoerle 2012a, f. 190.

Distribution: *COL.

Type: Colombia, Escobar et al. 2621 (holotype, SEL).

Lepanthes nautilus Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(10): 1046. 1984.

Illustration: ; Luer & Thoerle 2012a, f. 191; Moreno et al. 2018, f. 3f.

Distribution: *COL.

Type: Colombia, Luer et al. 10203 (holotype, MO; isotype SEL).

Lepanthes nematodes Luer & R.Escobar, Orquideología 18(1): 66-68. 1991.

Illustration: Luer & Escobar 1991a, p. 67; Luer & Thoerle 2012a, f. 192.

Distribution: *COL.

Type: Colombia, Luer 13018 (holotype, MO; isotype, JAUM).

Lepanthes nematostele Luer, Novon 3(4): 449, f. 11. 1993.

Illustration: Luer 1996b, f. 166 (p. 226); Luer & Thoerle 2012a, f. 193.

Distribution: COL ECU.

Type: Ecuador, Dalström & Höijer 1234 (holotype, MO).

Voucher: Colombia, Pérez & Parra s.n. (HPUJ, VALLE).

Lepanthes nicolasiiLuer & R.Escobar, Orquideología 20(3): 304-305. 1997.

Illustration: Luer & Escobar 1997, f. 124 (p. 306), p. 301 (top right); Luer & Thoerle 2012a, f. 194.

Distribution: *COL.

Type: Colombia, Escobar 5452 (holotype, MO).

Lepanthes niesseniae Luer, Revista Soc. Boliv. Bot. 4(1): 7. 2003.

Illustration: Luer 2003a, f. 4; Luer & Thoerle 2012a, f. 195; Baquero & Monteros 2020, f. 1c.

Distribution: *COL.

Type: Colombia, Niessen & Uribe 088 (holotype, MO).

Lepanthes niphas Luer & R.Escobar, Orquideología 18(1): 68-70. 1991.

Illustration: Luer & Escobar 1991a, p. 69; Luer & Thoerle 2012a, f. 196; Moreno et al. 2022, f. 7b.

Distribution: *COL.

Type: Colombia, Luer 13019 (holotype, MO; isotype JAUM).

Lepanthes nontecta Luer, Phytologia 54(5): 356. 1983.

Illustration: Luer 1996b, f. 168 (p. 226).

Distribution: COL ECU.

Type: Ecuador, Luer & Hirtz et al. 6887 (holotype, SEL).

Voucher: A photographic voucher of a Colombian specimen cited in Arias et al. 2023.

Lepanthes norae Foldats, Bol. Soc. Venez. Ci. Nat. 28: 244. 1969.

Illustration: Luer & Thoerle 2012a, f. 197a, f. 197b.

Distribution: *COL.

Type: Venezuela, Dunsterville & Dunsterville 64 (holotype, VEN).

Voucher: Colombia, Luer 10220 (MO).

Lepanthes nubicola Rchb.f., Xenia Orchid. 1: 149. 1858.

Distribution: *COL.

Type: Colombia, Wagener s.n. [601] (holotype, W-R-0054330; isotype, K-L).

Lepanthes nullaLuer & R.Escobar, Orquideología 19(2): 102. 1994.

Illustration: Luer & Escobar 1994, f. 97 (p. 109); Luer & Thoerle 2012a, f. 198.

Distribution: *COL.

Type: Colombia, Luer et al. 16748 (holotype, MO).

Lepanthes obovata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(8): 823. 1984.

Illustration: Luer & Thoerle 2012a, f. 199.

Distribution: *COL.

Type: Colombia, Escobar 3195 (holotype, MO; isotypes, JAUM, SEL).

Lepanthes octavioi Luer & R.Escobar, Orquideología 18(2): 128-130. 1991.

Illustration: Luer & Escobar 1991b, p. 129; Luer & Thoerle 2012a, f. 200.

Distribution: *COL.

Type: Colombia, Luer et al. 12548 (holotype, MO; isotype JAUM).

Lepanthes octopusLuer & R.Escobar, Orquideología 20(3): 305-307. 1997.

Illustration: Luer & Escobar 1997, f. 125 (p. 306), p. 300 (top left); Luer & Thoerle 2012a, f. 201.

Distribution: *COL.

Type: Colombia, Escobar s.n. (holotype, JAUM; isotype, MO).

Note: The holotype is Escobar s.n. according to the protologue, but Luer & Thoerle (2012a) indicate the type as Luer 17531 which is Luer’s illustration number.

Lepanthes ollaris Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(9): 920. 1984.

Illustration: Luer 1996b, f. 171 (p. 227); Luer & Thoerle 2012a, f. 202.

Distribution: COL ECU.

Type: Colombia, Luer et al. 8954 (holotype, SEL).

Lepanthes opetidion Luer & R.Escobar, Orquideología 18(2): 131-133. 1991.

Illustration: Luer & Escobar 1991b, p. 132; Luer & Thoerle 2012a, f. 203.

Distribution: *COL.

Type: Colombia, Luer 12504 (holotype, MO).

Lepanthes ophelma Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(6): 732-734. 1985.

Illustration: Luer & Thoerle 2012a, f. 204; VieiraUribe & Moreno 2019, f. 9c, f. 9d.

Distribution: *COL.

Type: Colombia, Escobar et al. 3380 (holotype, JAUM; isotypes, COL, MED, MO).

Lepanthes oreibates Luer & R.Escobar, Orquideología 18(2): 133, 136. 1991.

Illustration: Luer & Escobar 1991b, p. 134, p. 135(top); Luer & Thoerle 2012a, f. 205.

Distribution: *COL.

Type: Colombia, Luer 14227 (holotype, MO; isotype, JAUM).

Lepanthes orion Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(3): 316. 1985.

Illustration: Luer 1996b, f. 174 (p. 228); Luer & Thoerle 2012a, f. 206.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 13354 (holotype, MO).

Lepanthes ortiziana O.Pérez, E.Parra & Kolan., Orquideologia 27(2): 222. 2011.

Illustration: Pérez-Escobar et al. 2010b, p. 223, p. 224; Luer & Thoerle 2012a, f. 207; Baquero et al. 2021, f. 1f.

Distribution: *COL.

Type: Colombia, Pérez & Parra 752 (holotype, VALLE).

Lepanthes oscillifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(1): 30. 1984.

Illustration: Luer & Thoerle 2012a, f. 208; Moreno et al. 2021, f. 7c.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2669 (holotype, SEL).

Lepanthes osirisLuer & R.Escobar, Orquideología 19(2): 103. 1994.

Illustration: Luer & Escobar 1994, f. 98 (p. 109); Luer & Thoerle 2012a, f. 209.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8470 (holotype, SEL).

Lepanthes ostraconopetala Luer & Hirtz, Selbyana 30(1): 11. 2009.

Illustration: Luer 2009b, f. 20; Luer & Thoerle 2012a, f. 210.

Distribution: *COL.

Type: Colombia, Luer et al. 4630 (holotype, SEL).

Lepanthes otara Luer, Phytologia 54: 359. 1983.

Illustration: Luer 1996b, f. 176 (p. 228). Fig. 9A.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7163 (holotype, SEL).

Voucher: Colombia, Vieira 394 (JAUM).

Lepanthes oxybaphon Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(6): 734-736. 1985.

Illustration: Luer & Thoerle 2012a, f. 211.

Distribution: *COL.

Type: Colombia, Luer et al. 10228 (holotype, JAUM; isotype, MO).

Lepanthes oxypetala Luer & Hirtz, Orchidee (Hamburg) 39(2): 57. 1988.

Illustration: Luer 1996b, f. 177 (p. 229); Luer & Thoerle 2012a, f. 212.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11113 (holotype, MO).

Voucher: Colombia, R.Escobar 5240 (MO).

Lepanthes pachoiLuer & R.Escobar, Orquideología 19(2): 104. 1994.

Illustration: Luer & Escobar 1994, f. 99 (p. 79), f. 99 (p. 109); Luer & Thoerle 2012a, f. 213.

Distribution: *COL.

Type: Colombia, Escobar 5015 (holotype, JAUM; isotype, MO).

Lepanthes palaga Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(6): 604. 1984.

Illustration: Luer & Thoerle 2012a, f. 214.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2651 (holotype, SEL).

Lepanthes panope Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(3): 318-321. 1985.

Illustration: Luer & Thoerle 2012a, f. 215.

Distribution: *COL.

Type: Colombia, Luer et al. 10317 (holotype, MO).

Lepanthes papyrophylla Rchb.f., Xenia Orchid. 1: 151. 1856.

Illustration: Luer 1996b, f. 180 (p. 229).

Distribution: COL ECU.

Type: Ecuador, Jamieson 114 (holotype, W-R0054562; isotype, K-L).

Voucher: Colombia, Killip 20299 (US).

Lepanthes paramosolensis S.Vieira-Uribe & B.T.Larsen, Orquideología 31(2): 96. 2014.

Illustration: Vieira-Uribe & Larsen 2014b, f. 1, f. 2.

Distribution: *COL.

Type: Colombia, Domínguez & Pallandre 59 (holotype, JAUM).

Lepanthes parmata Luer & R.Escobar, Orquideología 18(2): 136-138. 1991.

Illustration: Luer & Escobar 1991b, p. 137; Luer & Thoerle 2012a, f. 216; Vieira-Uribe & Moreno 2018, f. 4a.

Distribution: *COL.

Type: Colombia, Cogollo et al. 4044 (holotype, JAUM; isotype, MO).

Lepanthes pastoënsis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 12. 1924.

Illustration: Luer & Thoerle 2012a, f. 217.

Distribution: *COL.

Type: Colombia, Klaboch s.n. [holotype, B, destroyed; lectotype, tab. 33, n. 125 in Schlechter 1920 (Luer & Thoerle 2012)].

Lepanthes pelyx Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 136-137. 1996.

Illustration: Luer 1996b, f. 183 (p. 230); Vieira-Uribe & Moreno 2018, f. 4d. Fig. 9B.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12431 (holotype, MO).

Voucher: Moreno & Erazo 406 (CAUP).

Lepanthes pembertoniiS. Vieira-Uribe & J.S.Moreno, Phytotaxa 567 (2): 149-161. 2022.

Illustration: Vieira et al. 2022, f. 1. Fig. 9C.

Distribution: *COL.

Type: Colombia, S. Vieira 29 (holotype, JAUM).

Lepanthes penicillifera Luer, Selbyana 30(1): 11. 2009.

Illustration: Luer 2003a, f. 5; Luer 2009b, f. 21; Luer & Thoerle 2012a, f. 218.

Distribution: *COL.

Type: Colombia, Niessen & Uribe 036A (holotype, MO).

Note: Described from a cultivated specimen without collection data.

Lepanthes perdita Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 138. 1996.

Illustration: Luer 1996b, f. 185 (p. 231); Luer & Thoerle 2012a, f. 219.

Distribution: COL ECU.

Type: Ecuador, Hirtz 2053 (holotype, MO).

Voucher: Colombia, Otero 1005 (CUVC, MO).

Lepanthes petalolenta Luer & R.Escobar, Orquideología 18(2): 139-141. 1991.

Illustration: Luer & Escobar 1991b, p. 140; Luer & Thoerle 2012a, f. 220.

Distribution: *COL.

Type: Colombia, Cogollo et al. 3907 (holotype, JAUM; isotype, MO).

Lepanthes petalopteryxLuer & R.Escobar, Orquideología 19(2): 106. 1994.

Illustration: Luer & Escobar 1994, f. 100 (p. 80), f. 100 (p. 109); Luer & Thoerle 2012a, f. 221; Moreno et al. 2020b, f. 4b, 5b.

Distribution: *COL.

Type: Colombia, Escobar 5061 (holotype, JAUM; isotype, MO).

Lepanthes phalloidesLuer & R.Escobar, Orquideología 21(2): 154-156. 1999.

Illustration: Luer & Escobar 1999, p. 155; Luer & Thoerle 2012a, f. 222.

Distribution: *COL.

Type: Colombia, Luer 16728 (holotype, MO).

Lepanthes pictoris Luer & Thoerle, Harvard Pap. Bot. 16(2): 336, 33. 2011.

Illustration: Luer & Thoerle 2012a, f. 223.

Distribution: *COL.

Type: Colombia, Thoerle 122A (holotype, MO).

Lepanthes pilosiaures Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(3): 262. 1984.

Illustration: Luer & Thoerle 2012a, f. 224.

Distribution: *COL.

Type: Colombia, Escobar et al. 2663 (holotype, SEL).

Lepanthes pinnatulaLuer & R.Escobar, Orquideología 20(3): 307-308. 1997.

Illustration: Luer & Escobar 1997, f. 126 (p. 306); Luer & Thoerle 2012a, f. 225.

Distribution: *COL.

Type: Colombia, Luer et al. 17676 (holotype, MO).

Lepanthes planadensis Luer & R.Escobar, Orquideología 18(2): 141-143. 1991.

Illustration: Luer & Escobar 1991b, p. 142; Luer & Thoerle 2012a, f. 226.

Distribution: *COL.

Type: Colombia, Luer 12487 (holotype, MO).

Lepanthes plectilis Luer & Hirtz, Amer. Orchid Soc. Bull. 54(2): 166. 1985.

Illustration: Luer 1996b, f. 187 (p. 231); Luer & Thoerle 2012a, f. 227.

Distribution: COL ECU.

Type: Ecuador, Hirtz 1861 (holotype, MO).

Voucher: Colombia, Luer 16254 (MO).

Note: Without collection data.

Lepanthes pleurorachis Luer, Phytologia 54(5): 363. 1983.

Illustration: Luer 1996b, f. 188 (p. 231); ; Luer & Thoerle 2012a, f. 228; Moreno et al. 2020b, f. 4c, 5c.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7373 (holotype, SEL).

Voucher: Colombia, Luer 13027 (MO).

Lepanthes pleurothallopsisLuer & R.Escobar, Orquideología 19(2): 107. 1994.

Illustration: Luer & Escobar 1994, f. 101 (p. 80), f. 101 (p. 110); Luer & Thoerle 2012a, f. 229.

Distribution: *COL.

Type: Colombia, Escobar 5007 (holotype, MO).

Lepanthes plumifera Luer, Phytologia 55: 190. 1984.

Illustration: Luer 1996b, f. 189 (p. 232). Fig. 9E.

Distribution: COL ECU.

Type: Ecuador, Luer et a. 5542 (holotype, SEL).

Voucher: A photo of a Colombian specimen is reproduced herein (Fig. 9E).

Lepanthes pogonochila Zuluaga, Alegria-Valencia, J.S.Moreno & Gal.-Tar., Harvard Pap. Bot. 26(1): 181-182. 2021.

Illustration: Moreno et al. 2021, f. 7a, f. 8, f. 9. Fig. 9D.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona et al. 1523 (holotype, CUVC).

Lepanthes popayanensisLuer & R.Escobar, Orquideología 21(2): 156-158. 1999.

Illustration: Luer & Escobar 1999, p. 157; Luer & Thoerle 2012a, f. 230.

Distribution: *COL.

Type: Colombia, Luer 16531 (holotype, MO).

Lepanthes porraceaLuer & R.Escobar, Orquideología 20(3): 309-310. 1997.

Illustration: Luer & Escobar 1997, f. 127 (p. 312), p. 301 (middle left); Luer & Thoerle 2012a, f. 231.

Distribution: *COL.

Type: Colombia, Escobar s.n. (holotype, JAUM; isotype, MO).

Note: The holotype is Escobar s.n. according to the protologue, but Luer & Thoerle (2012a) indicate the type as Luer 17721 which is Luer’s illustration number.

Lepanthes posadae Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(5): 479. 1984.

Illustration: Luer 1996b, f. 191 (p. 232); Luer & Thoerle 2012a, f. 232.

Distribution: COL ECU.

Type: Colombia, Luer 8743 (holotype, SEL).

Lepanthes praemorsa Luer & R.Escobar, Orquideología 18(2): 143-145. 1991.

Illustration: Luer & Escobar 1991b, p. 144; Luer & Thoerle 2012a, f. 233; Moreno et al. 2022b, f. 3b.

Distribution: *COL.

Type: Colombia, Luer et al. 14207 (holotype, MO).

Lepanthes profusa Luer & Hirtz, Orchidee (Hamburg) 41(1): 10. 1990.

Illustration: Luer & Thoerle 2012a, f. 234.

Distribution: BOL COL ECU.

Type: Ecuador, Luer et al. 10751 (holotype, MO).

Lepanthes prolifera Foldats, Bol. Soc. Venez. Ci. Nat. 28: 247. 1969.

Illustration: Luer & Thoerle 2012a, f. 235.

Distribution: *COL.

Type: Venezuela, Steryermark & Dunsterville 101198 (holotype, VEN).

Voucher: Colombia, Luer 7996 (SEL).

Lepanthes pteropogon Rchb.f., Xenia Orchid. 1: 146. 1856.

Illustration: Luer 1996b, f. 195 (p. 233).

Distribution: COL ECU.

Type: Ecuador, Jamieson 113 (holotype, W-R54545; isotype, K-L).

Voucher: Colombia, Franco-Rosselli 699 (COL).

Lepanthes pterygion Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(7): 720. 1984.

Illustration: Luer & Thoerle 2012a, f. 236.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2610 (holotype, SEL).

Lepanthes pyramidalis Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(6): 607. 1984.

Illustration: Luer & Thoerle 2012a, f. 237.

Distribution: *COL.

Type: Colombia, Luer et al. 8964 (holotype, SEL).

Lepanthes quadricornisLuer & R.Escobar, Orquideología 19(2): 108. 1994.

Illustration: Luer & Escobar 1994, f. 102 (p. 110); Luer 1996b, f. 7 (p. 11); Luer & Thoerle 2012a, f. 238; Baquero & Monteros 2020, f. 1d.

Distribution: *COL.

Type: Colombia, Dodson et al. 18060 (holotype, MO).

Lepanthes quandi Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(9): 924. 1984.

Illustration: Luer & Thoerle 2012a, f. 239.

Distribution: *COL.

Type: Colombia, Luer et al. 10059 (holotype, MO).

Lepanthes quaternaria Luer, Phytologia 54: 367. 1983.

Illustration: Luer 1996b, f. 198 (p. 234). Fig. 9F.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6307 (holotype, SEL).

Voucher: Colombia, Galindo et al. 1476 (CUVC).

Lepanthes ramosii Luer, Orquideología 22(1): 45-47. 2001.

Illustration: Luer 2001b, p. 47; Luer & Thoerle 2012a, f. 240.

Distribution: *COL.

Type: Colombia, Ramos & Silverstone-Sopkin 1176 (holotype, CUVC; isotype, MO).

Lepanthes reburra Luer & Thoerle, Harvard Pap. Bot. 16(2): 333. 2011.

Illustration: Luer & Thoerle 2012a, f. 241.

Distribution: *COL.

Type: Colombia, Luer et al. 7801 (holotype, SEL).

Lepanthes refracta Luer, Orquideología 22(1): 48-50. 2001.

Illustration: Luer 2001b, p. 49; Luer & Thoerle 2012a, f. 242.

Distribution: *COL.

Type: Colombia, Niessen 071A (holotype, MO).

Note: Without collection data.

Lepanthes renzii Luer, Selbyana 30(1): 13. 2009.

Illustration: Luer & Thoerle 2012a, f. 243.

Distribution: *COL.

Type: Colombia, Renz 3527 (holotype, BAS; isotype, MO).

Lepanthes repens Luer, Phytologia 54(5): 368. 1983.

Illustration: Luer 1996b, f. 199 (p. 234); Luer & Thoerle 2012a, f. 244.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6261 (holotype, SEL).

Lepanthes reticulata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(6): 608. 1984.

Illustration: Luer & Thoerle 2012a, f. 245.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2650 (holotype, SEL).

Lepanthes rhombipetala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 98. 1920.

Illustration: Mansfeld 1929, pl. 33 (n. 127); Luer 1996b, f. 202 (p. 235); Luer & Thoerle 2012a, f. 246.

Distribution: COL ECU.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype, tab. 33, n. 127 in Schlechter 1920 (Luer & Thoerle 2012)].

Lepanthes rhynchion Luer, Phytologia 54(5): 368. 1983.

Illustration: Luer 1996b, f. 203 (p. 235); Luer & Thoerle 2012a, f. 247.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3333 (holotype, MO).

Lepanthes ribes Luer, Selbyana 3(1-2): 14. 1976.

Illustration: Luer 1996b, f. 8 (p. 11); Luer & Thoerle 2012a, f. 248; Baquero & Monteros 2020, f. 1e.

Distribution: COL ECU.

Type: Ecuador, Luer & Kent 233 (holotype, SEL).

Voucher: Colombia, Luer 8075 (JAUM).

Lepanthes ricaurtensisLuer & R.Escobar, Orquideología 19(2): 111. 1994.

Illustration: Luer & Escobar 1994, f. 103 (p. 110); Luer & Thoerle 2012a, f. 249.

Distribution: *COL.

Type: Colombia, Luer et al. 4622 (holotype, SEL).

Lepanthes rodrigoi Luer, Orquideología 20(2): 198-201. 1996.

Illustration: Luer 1996f, 199; Luer & Thoerle 2012a, f. 250.

Distribution: *COL.

Type: Colombia, Escobar et al. s.n. (holotype, JAUM).

Lepanthes roezlianaLuer & R.Escobar, Orquideología 17(3): 216. 1988.

Illustration: Luer & Escobar 1988, p. 217; Luer & Thoerle 2012a, f. 251.

Distribution: *COL.

Type: Colombia, Luer et al. 9093 (holotype, SEL).

Lepanthes rutkisiiFoldats, Acta Bot. Venez. 3: 344. 1968.

Illustration: Foldats 1968, f. 13. Fig. 10K.

Distribution: COL VEN.

Type: Venezuela, Steyermark & Dunsterville 98322 (holotype, VEN).

Voucher: A photographic voucher of a Colombian specimen reproduced here (Fig. 10K).

Lepanthes rutrumLuer & R.Escobar, Orquideología 19(2): 112. 1994.

Illustration: Luer & Escobar 1994, f. 104 (p. 80), f. 104 (p. 110); Luer & Thoerle 2012a, f. 252.

Distribution: *COL.

Type: Colombia, Escobar et al. 4005 (holotype, MO; isotype JAUM).

Lepanthes sabinadaleyana J.S.Moreno & S.VieiraUribe, Lankesteriana 19(2): 64. 2019.

Illustration: Vieira-Uribe & Moreno 2019, f. 1-3, f. 4a. Fig. 9G.

Distribution: *COL.

Type: Colombia, Moreno et al. 519 (holotype, JAUM; isotype CAUP).

Lepanthes saccataLuer & R.Escobar, Orquideología 17(3): 220-223. 1988.

Illustration: Luer & Escobar 1988, p. 221; Luer & Thoerle 2012a, f. 253.

Distribution: *COL.

Type: Colombia, Escobar et al. 3405 (holotype, MO).

Lepanthes saltatrix Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 150. 1996.

Illustration: Luer 1996b, f. 207 (p. 236). Fig. 9H.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12424 (holotype, MO).

Voucher: Colombia, de Benavides 10301 (CAUP).

Lepanthes sanjuanensis Bogarín & Karremans, Lankesteriana 12(2): 113-114, f. 3, 4C. 2012.

Illustration: Bogarín et al. 2012, f. 3, f. 4C. Fig. 9I.

Distribution: COL COS PAN.

Type: Costa Rica, Bogarín & Karremans 9234 (holotype, JBL-spirit; isotype, CR).

Voucher: Colombia, Moreno & Erazo 395 (CAUP).

Lepanthes satyrica Luer & Hirtz, Lindleyana 2(3): 141. 1987.

Illustration: Luer 1996b, f. 208 (p. 236); Luer & Escobar 1999, p. 139; Luer & Thoerle 2012a, f. 254; Baquero et al. 2019, f. 4d.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12430 (holotype, MO).

Lepanthes scalaris Luer, Phytologia 54(5): 369. 1983.

Illustration: Luer 1996b, f. 209 (p. 237); Luer & Thoerle 2012a, f. 255.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6255 (holotype, SEL).

Lepanthes schizix Luer, Phytologia 54: 370 (1983).

Illustration: Luer 1996b, f. 212 (p. 237). Fig. 9J.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 4411 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Galindo et al. 2020, p. 205.

Lepanthes schlechteriana J.M.H. Shaw, Orchid Review Suppl., 124(1314): 37. 2016. Lepanthes marginata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 96. 1920, nom. illeg. Non Lepanthes marginata Barb.Rodr. [1881].

Illustration: Mansfeld 1929, pl. 33 (n. 124).

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 33, No. 124)].

Lepanthes schnitteri Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 160. 1924.

Distribution: *COL.

Type: Colombia, Schnitter s.n. [holotype, B, destroyed; lectotype (selected here), AMES00100703, drawing of type].

Note: An obscure taxon.

Lepanthes scoliosa Luer & Hirtz, Harvard Pap. Bot. 17: 352. 2012.

Illustration: Luer & Thoerle 2012b, f. 24. Fig. 10L.

Distribution: COL ECU.

Type: Ecuador, Hirtz & Gonzalez 1031 (holotype, MO).

Voucher: Colombia, Zuluaga 4932 (CUVC).

Lepanthes scopuliferaLuer & R.Escobar, Orquideología 20(3): 310-311. 1997.

Illustration: Luer & Escobar 1997, f. 128 (p. 312); Luer & Thoerle 2012a, f. 256.

Distribution: *COL.

Type: Colombia, Luer et al. 17664 (holotype, MO).

Lepanthes sericinitens Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(10): 1048. 1984.

Illustration: Luer & Thoerle 2012a, f. 257.

Distribution: *COL.

Type: Colombia, Luer 10032 (holotype, MO; isotype, SEL).

Lepanthes series Luer & Hirtz, Lindleyana 2: 105 (1987).

Illustration: Luer 1996b, f. 216 (p. 238). Fig. 10H.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 10864 (holotype, MO).

Voucher: Colombia, Rincón-Gonzalez 1438 (TOLI).

Lepanthes setifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(8): 822. 1984.

Illustration: Luer 1996b, f. 217 (p. 239); Luer & Thoerle 2012a, f. 258.

Distribution: COL ECU GUA.

Type: Ecuador, Luer et al. 9863 (holotype, MO).

Voucher: Colombia, Escobar 3190 (JAUM, MO, SEL).

Lepanthes silverstonei Luer, Orquideología 22(1): 50-53. 2001.

Illustration: Luer 2001b, p. 51; Luer & Thoerle 2012a, f. 259.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 2864 (holotype, CUVC; isotype, MO).

Lepanthes sinuosaLuer & R.Escobar, Orquideología 19(2): 113-115. 1994.

Illustration: Luer & Escobar 1994, f. 105 (p. 80), f. 105 (p. 117); Luer & Thoerle 2012a, f. 260; Moreno et al. 2022b, f. 6d.

Distribution: *COL.

Type: Colombia, Luer et al. 16846 (holotype, MO).

Lepanthes skeleton Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(7): 722. 1984.

Illustration: Luer & Thoerle 2012a, f. 261.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 3231 (holotype, MO; isotypes, COL, JAUM, SEL).

Lepanthes smaragdinaLuer & R.Escobar, Orquideología 19(2): 115. 1994.

Illustration: Luer & Escobar 1994, f. 106 (p. 80), f. 106 (p. 117); Luer & Thoerle 2012a, f. 262; Baquero et al. 2021, f. 1b, f. 4b, f. 4d.

Distribution: *COL.

Type: Colombia, Escobar 5083 (holotype, JAUM; isotype, MO).

Lepanthes solicitor Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(1): 58. 1985.

Illustration: Luer & Thoerle 2012a, f. 263.

Distribution: *COL.

Type: Colombia, Luer et al. 10328 (holotype, MO).

Lepanthes speciosa Luer & Hirtz, Orchidee (Hamburg) 43: 67. 1992.

Illustration: Luer 1996b, f. 222 (p. 240). Fig. 9K.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9884 (holotype, MO).

Voucher: Colombia, Galindo & Espitia 1526 (CUVC).

Lepanthes spelynx Luer & R.Escobar, Amer. Orchid Soc. Bull. 54: 586. 1985.

Illustration: Luer & Thoerle 2012a, f. 264.

Distribution: *COL.

Type: Colombia, Luer et al. 10329 (holotype, JAUM; isotype, MO).

Lepanthes stelidantha Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 202. 1976.

Distribution: COL VEN.

Type: Venezuela, Dunsterville 67 (holotype, AMES).

Voucher: Colombia, Delgado-V. 69 (FMB).

Lepanthes stelidilabia Luer & R.Escobar, Orquideología 18(2): 146-148. 1991.

Illustration: Luer & Escobar 1991b, p. 144; Luer & Thoerle 2012a, f. 265.

Distribution: *COL.

Type: Colombia, Luer 14229 (holotype, MO).

Lepanthes stellaris Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 249-250. 2009.

Illustration: Luer 2009a, f. 11 (p. 255); Luer & Thoerle 2012a, f. 266; Pérez-Escobar et al. 2010a, f. 1, f. 2.

Distribution: COL ECU.

Type: Ecuador, Hirtz 8270 (holotype, MO).

Voucher: Colombia, Pérez 240 (VALLE).

Lepanthes stenoscleros Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 99. 1920.

Illustration: Mansfeld 1929, Pl. 34 (n. 128).

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 34, No. 128)].

Note: An obscure taxon not monographed by Luer.

Lepanthes steyermarkiiFoldats, Acta Bot. Venez. 3: 346. 1968.

Illustration: Luer & Thoerle 2012a, f. 267.

Distribution: COL VEN.

Type: Venezuela, Steyermark et al. 98381 (holotype, VEN).

Voucher: Colombia, Luer et al. 10297.

Lepanthes strumosa Luer & R.Escobar, Orquideología 16(2): 151. 1984.

Illustration: Luer & Escobar 1984a, p. 152; Luer & Thoerle 2012a, f. 268; Luer 1996b, f. 226 (p. 241).

Distribution: COL ECU.

Type: Colombia, Escobar & Valencia 2625 (holotype, SEL).

Lepanthes stupenda Luer, Phytologia 54(5): 372. 1983.

Illustration: Luer & Thoerle 2012a, f. 269.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6042 (holotype, SEL).

Voucher: Colombia, Escobar 5023 (MO).

Lepanthes subulata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(6): 609. 1984.

Illustration: Luer & Thoerle 2012a, f. 270.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2624 (holotype, SEL).

Lepanthes superposita Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 99. 1920.

Illustration: Luer & Thoerle 2012a, f. 271.

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype, tab. 34, n. 129 in Schlechter 1920 (Luer & Thoerle 2012a)].

Lepanthes surrogata Luer & Hirtz.

Illustration: Luer 1996b, f. 230 (p. 242).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11910 (holotype, MO).

Voucher: Colombia, Moreno 531 (CUVC).

Lepanthes sylvilagus E.Restrepo & S.Vieira-Uribe, Lankesteriana 22(1): 2. 2022.

Illustration: Restrepo et al. 2022, f. 1, f. 2, f. 3a-b. Fig. 9L.

Distribution: *COL.

Type: Colombia, Vieira-Uribe 53 (holotype, JAUM-spirit).

Lepanthes tachirensisFoldats, Acta Bot. Venez. 3: 348. 1968.

Illustration: Luer 1996b, f. 233 (p. 243); Luer & Thoerle 2012a, f. 272.

Distribution: COL ECU PER VEN.

Type: Venezuela, Steryermark & Dunsterville 98524 (holotype, VEN).

Voucher: Colombia, Luer 17682 (MO).

Lepanthes tamaensisFoldats, Acta Bot. Venez. 3: 351. 1968.

Illustration: Luer & Thoerle 2012a, f. 273.

Distribution: COL VEN.

Type: Venezuela, Steryermark & Dunsterville 98658 (holotype, VEN).

Voucher: Colombia, Luer 14268 (MO).

Lepanthes tanekes Luer & R.Escobar, Orquideología 18(2): 148. 1991.

Illustration: Luer & Escobar 1991b, p. 149; Luer & Thoerle 2012a, f. 274.

Distribution: *COL.

Type: Colombia, Hirtz et al. 4625 (holotype, SEL).

Lepanthes tatamaeS.Vieira-Uribe & J.S. Moreno, Orquideología 35(2): 184. 2018.

Illustration: Vieira-Uribe & Moreno 2018, f. 5, f. 6. Fig. 10A.

Distribution: *COL.

Type: Colombia, Vieira et al. 25 (holotype, JAUM).

Lepanthes telipogoniflora Schuit. & A.de Wilde, Orquideología 20: 27. 1996.

Illustration: Schuiteman & de Wilde 1996, p. 29, p. 31; Luer & Thoerle 2012a, f. 275.

Distribution: *COL.

Type: Colombia, de Wilde et al. 5010 (holotype, JAUM).

Lepanthes tetrachaeta Luer & L.Jost, Monogr. Syst. Bot. Missouri Bot. Gard. 79: 119. 2000.

Illustration: Luer 2011, f. 26a (p. 135). Fig. 10B.

Distribution: COL ECU.

Type: Ecuador, Jost & Souther 1806 (holotype, MO).

Voucher: Colombia, Moreno 527 (CAUP).

Lepanthes tetracolaLuer & R.Escobar, Orquideología 19(2): 116. 1994.

Illustration: Luer & Escobar 1994, f. 107 (p. 117); Luer & Thoerle 2012a, f. 276.

Distribution: *COL.

Type: Colombia, Luer 16875 (holotype, MO).

Lepanthes tibouchinicola Luer & R.Escobar, Amer. Orchid Soc. Bull. 52: 1269. 1983.

Illustration: Luer & Thoerle 2012a, f. 277.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2660 (holotype, SEL).

Lepanthes tomentosa Luer, Lindleyana 6(2): 76, 79. 1991.

Illustration: Ortiz et al. 2011, p. 23, p. 24; Luer & Thoerle 2012a, f. 278; Baquero et al. 2021, f. 1d.

Distribution: *COL.

Type: Colombia, Luer 13786 (holotype, MO).

Lepanthes tortilis Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 170. 1996.

Illustration: Luer 1996b, f. 242 (p. 245). Fig. 10C.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12445 (holotype, MO).

Voucher: Colombia, Zuluaga 5636 (CUVC).

Lepanthes tracheia Rchb.f., Flora 69: 557. 1886.

Illustration: Luer 1996b, f. 243 (p. 245); Luer & Thoerle 2012a, f. 279.

Distribution: COL ECU VEN.

Type: Colombia, Lehmann 3562 [lectotype, W-R54518 (Luer & Thoerle 2012); isolectotype, BM000082181].

Lepanthes transparens Luer, Phytologia 54(5): 374. 1983.

Illustration: Luer & Thoerle 2012a, f. 280.

Distribution: COL ECU.

Type: Ecuador, Luer et al 6260 (holotype, SEL).

Voucher: Colombia, Luer 8381 (SEL).

Lepanthes trichocaulisLuer & R.Escobar, Orquideología 17(3): 224-226. 1988.

Illustration: Luer & Escobar 1988, p. 227; Luer & Thoerle 2012a, f. 281.

Distribution: *COL.

Type: Colombia, Escobar 3245 (holotype, MO).

Lepanthes trifurcataLuer & R.Escobar, Orquideología 19(2): 119. 1994.

Illustration: Luer & Escobar 1994, f. 108 (p. 80), f. 108 (p. 117); Luer & Thoerle 2012a, f. 282.

Distribution: *COL.

Type: Colombia, Luer 16720 (holotype, MO).

Lepanthes trimerinx Luer, Phytologia 54(5): 374. 1983.

Illustration: Luer 1996b, f. 245 (p. 246); Luer & Thoerle 2012a, f. 283.

Distribution: COL ECU.

Type: Ecuador, Luer 1019 (holotype, SEL).

Voucher: Colombia, Ramos 954 (MO).

Lepanthes trinariaLuer & R.Escobar, Orquideología 20(3): 311. 1997.

Illustration: Luer & Escobar 1997, f. 129 (p. 312), p. 301 (bottom right); Luer & Thoerle 2012a, f. 284.

Distribution: *COL.

Type: Colombia, Luer 17530 (holotype, MO).

Lepanthes troglodytes Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(3): 320-321. 1985.

Illustration: Luer & Thoerle 2012a, f. 285.

Distribution: *COL.

Type: Colombia, Luer 10031 (holotype, JAUM; isotype, MO).

Lepanthes troxis Luer & R.Escobar, Orquideología 18(2): 150-152. 1991.

Illustration: Luer & Escobar 1991b, p. 135 (bottom), p. 151; Luer & Thoerle 2012a, f. 286.

Distribution: *COL.

Type: Colombia, Luer et al. 10366 (holotype, MO; isotype JAUM).

Lepanthes tsubotae Luer & R.Escobar, Orquideología 19(1): 39. 1993.

Illustration: Luer & Escobar 1993a, p. 20 (middle above), p. 29 (right); Luer & Thoerle 2012a, f. 287.

Distribution: *COL.

Type: Colombia, Luer 15261 (holotype, MO).

Lepanthes umbonata Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(10): 1050-1051. 1984.

Illustration: Luer & Thoerle 2012a, f. 288a, f. 288b.

Distribution: *COL.

Type: Colombia, Escobar 3206 (holotype, MO; isotype JAUM).

Lepanthes uncifera Luer & R.Escobar, Amer. Orchid Soc. Bull. 53: 580. 1984.

Illustration: Luer & Thoerle 2012a, f. 289.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2668 (holotype, SEL).

Lepanthes uribei Luer, Revista Soc. Boliv. Bot. 4(1): 11. 2003.

Illustration: Luer & Thoerle 2012a, f. 290.

Distribution: *COL.

Type: Colombia, Niessen & Uribe 095 (holotype, MO).

Lepanthes valenciae Luer & R.Escobar, Amer. Orchid Soc. Bull. 53: 825. 1984.

Illustration: Luer & Thoerle 2012a, f. 291.

Distribution: *COL.

Type: Colombia, Escobar 3185A (holotype, MO).

Lepanthes valerieae O.Pérez, Jaramillo & Bogarín, Lankesteriana 21(3): 356. 2021.

Illustration: Pérez-Escobar et al. 2021, f. 3, f. 4. Fig. 10E.

Distribution: *COL.

Type: Colombia, Pérez et al. 1981 (holotype, CUVC).

Lepanthes vareschii Garay, Canad. J. Bot. 34: 252. 1956.

Illustration: Garay 1956a, f. 11A.

Distribution: COL VEN.

Type: Venezuela, Vareschi 2306 (holotype, AMES3473; isotype, VEN).

Voucher: Colombia, Schneider 672 (COL).

Lepanthes vargasiiJ.S.Moreno, S.Vieira-Uribe & Karremans, Lankesteriana 17(2): 227. 2017.

Illustration: Moreno et al. 2017, f. 1, 2. Fig. 10F.

Distribution: *COL.

Type: Colombia, Moreno & Erazo 342 (holotype, CAUP).

Lepanthes vellicata Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 61(3): 177, f. 254. 1996.

Illustration: Luer 1996b, f. 254 (p. 248); Luer & Thoerle 2012a, f. 292.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12419 (holotype, MO).

Voucher: Colombia, Luer 16735 (MO).

Lepanthes vespertilio Rchb.f., Xenia Orchid. 1: 143. 1856.

Illustration: Luer 1996b, f. 256 (p. 248). ig. 10I.

Distribution: COL ECU.

Type: Ecuador, Jamieson 96 (holotype, W-R0054345; isotype, K-L).

Voucher: Colombia, Pennel 10497 (US).

Lepanthes venustaLuer & R.Escobar, Orquideología 19(2): 120. 1994.

Illustration: Luer & Escobar 1994, f. 109 (p. 118); Luer & Escobar 1999, p. 137; Luer & Thoerle 2012a, f. 293.

Distribution: *COL.

Type: Colombia, Luer et al. 16839 (holotype, MO).

Lepanthes viahoënsisLuer & R.Escobar, Orquideología 20(3): 311-313. 1997.

Illustration: Luer & Escobar 1997, f. 130 (p. 312), p. 301 (top left); Luer & Thoerle 2012a, f. 294; Moreno et al. 2020b, f. 4d, 5d.

Distribution: *COL.

Type: Colombia, Luer 17719 (holotype, MO).

Lepanthes vibrissa Luer & Hirtz, Lindleyana 2(3): 155. 1987.

Illustration: Luer 1996b, f. 258 (p. 249); Luer & Thoerle 2012a, f. 295.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11263 (holotype, MO).

Voucher: Colombia, Luer 17666 (MO).

Lepanthes vieiraeLuer & R.Escobar, Orquideología 17(3): 228-231. 1988.

Illustration: Luer & Escobar 1988, p. 231; Luer & Thoerle 2012a, f. 296.

Distribution: *COL.

Type: Colombia, Luer 10393 (holotype, MO).

Lepanthes volvox Luer & R.Escobar, Amer. Orchid Soc. Bull. 53: 481. 1984.

Illustration: Luer & Thoerle 2012a, f. 297.

Distribution: *COL.

Type: Colombia, Escobar 3194 (holotype, MO; isotype, JAUM).

Lepanthes wageneri Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Illustration: Luer 1996b, f. 260 (p. 249); Luer & Thoerle 2012a, f. 298; Vieira-Uribe & Moreno 2019, f. 4c.

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Wagener s.n. (holotype, W; isotype K).

Lepanthes wakemaniae S.Vieira-Uribe & J.S.Moreno, Lankesteriana 22(1): 54. 2022.

Illustration: Vieira-Uribe & Moreno 2022, f. 1, f. 2, f. 5a, f. 6a. Fig. 10G.

Distribution: *COL.

Type: Colombia, Vieira & Pérez 027 (holotype, JAUM; isotype, HUA).

Lepanthes xenos Luer & Hirtz, Orchidee (Hamburg) 38: 32. 1987.

Illustration: Luer 1996b, f. 261 (p. 250). Fig. 10D.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9582 (holotype, MO).

Voucher: A photo of a Colombian specimen is reproduced herein (Fig. 10D).

Lepanthes yubarta E.Calderón ex Luer & Thoerle, Monogr. Syst. Bot. Missouri Bot. Gard. 123: 217-218. 2012. Lepanthes yubarta E.Calderón, Orquideología 27: 200. 2010 [2011], nom. inval.

Illustration: Calderón-Sáenz 2010, f. 1, f. 2, p. 202; Luer & Thoerle 2012a, f. 299.

Distribution: *COL.

Type: Colombia, Calderón 602 (holotype, HPUJ027414).

Note: This taxon was originally published invalidly for not specifying a single specimen as holotype. It was unintentionally validated by Luer & Thoerle (2012a) who provided a direct reference and clearly indicated a holotype.

Lepanthes zapataeLuer & R.Escobar, Orquideología 19(2): 121. 1994.

Illustration: Luer & Escobar 1994, f. 110 (p. 79), f. 110 (p. 118); Luer & Thoerle 2012a, f. 300.

Distribution: *COL.

Type: Colombia, Luer 16724 (holotype, MO).

Lepanthopsis (Cogn.) Ames

Lepanthopsis abbreviata Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 8. 1991.

Illustration: Luer 1991a, pl. 1 (p. 9).

Distribution: COL ECU.

Type: Ecuador, Hirtz 1524 (holotype, MO).

Voucher: Colombia, Luer 2176 (SEL).

Lepanthopsis acetabulum Luer, Phytologia 55(3): 196-197. 1984.

Illustration: Luer 1991a, pl. 2 (p. 11).

Distribution: BOL COL ECU PER.

Type: Ecuador, Dalström 417 (holotype, SEL).

Voucher: Colombia, Luer 14270 (MO).

Lepanthopsis acuminata Ames, Bot. Mus. Leafl. 6(4): 70. 1938.

Illustration: Luer 1991a, pl. 3 (p. 13).

Distribution: COL ECU VEN.

Type: Colombia, Killip 11298 (holotype, AMES).

Lepanthopsis apoda (Garay & Dunst.) Luer, Selbyana 7(1): 100. 1982. Lepanthopsis apoda (Garay & Dunst.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 68. 1991, nom. superfl. Expedicula apoda (Garay & Dunst.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005. Pleurothallis apoda Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 3: 246. 1965. Trichosalpinx apoda (Garay & Dunst.) Luer, Phytologia 54(5): 394. 1983.

Illustration: Luer 1991a, pl. 33 (p. 69); Vásquez & Ibisch 2000, pl. 56; Ortiz & Medina 2012, f. 3, f. 4.

Distribution: BOL COL ECU VEN.

Type: Venezuela, Dunsterville 506 (holotype, AMES).

Voucher: Colombia, Luer 14333 (MO).

Lepanthopsis astrophora (Rchb.f. ex Kraenzl.) Garay, Caldasia 8: 520. 1962. Pleurothallis astrophora Rchb.f. ex Kraenzl., Xenia Orchid. 3: 85. 1892.

Illustration: Luer 1991a, pl. 6 (p. 19 ).

Distribution: COL VEN.

Type: Venezuela, Wendland s.n. (holotype, W).

Voucher: Colombia, Ospina 82 (JAUM).

Lepanthopsis dewildei Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 116. 1994. Expedicula dewildei (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005.

Illustration: Luer 1991a, pl. 38 (p. 117).

Distribution: *COL.

Type: Colombia, de Wilde 4436 (holotype, MO).

Lepanthopsis floripecten (Rchb.f.) Ames, Bot. Mus. Leafl. 1(9): 11. 1933. Humboldtia floripecten (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Lepanthes floripecten (Rchb.f.) Ames, Bot. Mus. Leafl. 1(9): 11. 1933. Pleurothallis floripecten Rchb.f., Bonplandia (Hannover) 2: 25. 1854.

Illustration: Cogniaux 1896, f. 2 (p. 122); Luer 1991a, pl. 14 (p. 35). Fig. 12A.

Distribution: BRA COL COS ECU ELS HON MEX NIC PAN VEN.

Type: Venezuela, Wagener s.n. [L564] (holotype, W-R-0053767; isotype, K-L-000079888).

Voucher: Colombia, Luer 13049 (MO).

Lepanthopsis hirtzii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 38, pl. 16. 1991.

Illustration: Luer 1991a, pl. 15 (p. 37).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 10953 (holotype, MO).

Voucher: Colombia, Luer 14217 (MO).

Lepanthopsis obliquipetala (Ames & C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 76. 1991. Physosiphon obliquipetalus Ames & C.Schweinf., Sched. Orch. 8: 12-14. 1925. Pleurothallis connata Luer, Selbyana 5(3-4): 388. 1981. Stelis obliquipetala (Ames & C.Schweinf.) L.O.Williams, Ceiba 5(1): 54. 1956. Trichosalpinx obliquipetala (Ames & C.Schweinf.) Luer, Phytologia 54(5): 396. 1983.

Illustration: Luer 1991a, pl. 37 (p. 77); Duque 1993, p. 311, p. 313.

Distribution: COL COS ECU.

Type: Costa Rica, Lankester 680 (holotype, AMES).

Voucher: Colombia, Duque 1637 (MO).

Lepanthopsis peniculus (Schltr.) Garay, Orchid J. 2(10): 468. 1953. Pleurothallis peniculus Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 110. 1920.

Illustration: Luer 1991a, pl. 23(p. 51); Mansfeld 1929, pl. 38 (n. 146). Fig. 12B.

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 38, No. 146)].

Lepanthopsis pristis Luer & R.Escobar, Orquideología 16(3): 27-28. 1986.

Illustration: Luer & Escobar 1986, p. 29; Luer 1991a, pl. 24 (p. 53).

Distribution: *COL.

Type: Colombia, Escobar et al. 3383 (holotype, MO).

Lepanthopsis prolifera Garay, Orchid J. 2: 468. 1954.

Illustration: Luer 1991a, pl. 25 (p. 54).

Distribution: *COL.

Type: Colombia, Schultes & Smith 12537 ( holotype, US; isotype, AMES).

Lepanthopsis undulata Kolan. & Szlach., Ann. Bot. Fenn. 57: 140-141. 2020.

Illustration: Kolanowska & Szlachetko 2020a, f. 1.

Distribution: *COL.

Type: Colombia, Schneider 200 (holotype, COL; isotype, COL).

Lepanthopsis vinacea C.Schweinf., Bot. Mus. Leafl. 18: 109. 1958.

Illustration: Luer 1991, pl. 32 (p. 67).

Distribution: COL ECU VEN.

Type: Venezuela, Wurdack 33866 (holotype, AMES).

Voucher: Colombia, Luer 14217 (MO).

Madisonia Luer

Madisonia kerrii (Braga) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004. Pleurothallis kerrii Braga, Bradea 3: 172. 1981.

Illustration: Luer 2006, f. 46; Smidt et al. 2021, f. 2, f. 3.

Distribution: BRA COL PER VEN.

Type: Brazil, Madison et al. PFE 366 (holotype, INPA).

Voucher: Colombia, Cardenas 24034 (COAH).

Madisonia spiculifera (Lindl.) Toscano & Smidt, Phytotaxa 505(1): 81. 2021. Anathallis spiculifera (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 259. 2009. Humboldtia spiculifera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis spiculifera Lindl., Fol. Orchid. ~Pleurothallis~ 43. 1859. Specklinia spiculifera (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001.

Illustration: Luer 2006, f. 35a, 35b.

Distribution: BOL BRA COL ECU FRE GUY JAM L.A. SUR VEN.

Type: Venezuela [incorrectly cited as Brazil], Spruce 3154 (holotype, K; isotypes, P, W).

Voucher: Colombia, Schultes & Cabrera 14559 (MO).

Masdevallia Ruiz & Pav.

Masdevallia aenigma Luer & R.Escobar, Selbyana 7: 69. 1982. Alaticaulia aenigma (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 4. 2006.

Illustration: Escobar 1983, p. 67; Luer 2000b, pl. 2, pl. 3.

Distribution: *COL.

Type: Colombia, Luer et al. 7571 (holotype, SEL; isotype, COL).

Masdevallia aguirrei Luer & R.Escobar, Harvard Pap. Bot. 9: 1. 1996. Alaticaulia aguirrei (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 4. 2006.

Illustration: Luer 2000b, pl. 4; Ortiz & Medina 2012, f. 7.

Distribution: *COL.

Type: Colombia, Luer 17988 (holotype, MO).

Masdevallia akemiana Königer & Sijm, Arcula 12: 298. 2003.

Illustration: Königer 2003, p. 299.

Distribution: *COL.

Type: Colombia, Königer WK-112 (holotype, M; isotype, JAUM).

Masdevallia alismifolia Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 422. 1921.

Illustration: Luer 2002b, pl. 394.

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, W-R0083477).

Masdevallia × alvaroi Luer & R.Escobar, Orquideología 13: 47. 1978. Spilotantha × alvaroi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Luer & Escobar 1978, p. 48, p. 49, p. 60 (top left); Luer 2003b, pl. 573.

Distribution: *COL.

Type: Colombia, Luer 1978 (holotype, SEL).

Masdevallia amanda Rchb.f., Bonplandia (Hannover) 2: 115. 1854. Spilotantha amanda (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Luer 2003b, pl. 574; Gutiérrez Morales & Oses 2017, f. 6.

Distribution: COL ECU VEN.

Type: Colombia, Warszewicz s.n. [lectotype (selected here), W-R-0047348, middle illustr.].

Masdevallia andreae P.Ortiz, Orquideología 26: 11. 2009.

Illustration: Ortiz 2009a, p. 8 (top), p. 11.

Distribution: *COL.

Type: Colombia, Ortiz 1306 (holotype, HPUJ, lost).

Masdevallia angulata Rchb.f., Otia Bot. Hamburg. 1: 15. 1878. Byrsella angulata (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 7. 2006.

Illustration: Luer 2000c, pl. 133.

Distribution: COL ECU.

Type: Ecuador, Lehmann s.n. (holotype, W-R38736; isotype, K).

Voucher: Colombia, Escobar 3213 (MO).

Masdevallia angulifera Rchb.f. ex Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 19. 1925.

Illustration: Luer 2002b, pl. 503, pl. 504. Fig. 12C.

Distribution: *COL.

Type: Colombia, Wallis s.n. (holotype, W-R0050954).

Masdevallia anisomorpha Garay, Orquideología 5: 79. 1970. Rodrigoa anisomorpha (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 14. 2006.

Illustration: Luer 2003b, pl. 621, pl. 622.

Distribution: *COL.

Type: Colombia, Escobar 446 (holotype, AMES).

Masdevallia apparitio Luer & R.Escobar, Orquideología 13(2): 145. 1979.

Illustration: Luer & Escobar 1979, p. 146, p. 147, p. 170 (bottom left); Luer 2002b, pl. 396.

Distribution: *COL.

Type: Colombia, Escobar 1490 (holotype, JAUM).

Masdevallia arangoi Luer & R.Escobar, Orquideología 13(2): 149. 1979. Acinopetala arangoi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer & Escobar 1979, p. 150, p. 151, p. 168 (top right); Luer 2001a, pl. 365.

Distribution: *COL.

Type: Colombia, Escobar 1391 (holotype, JAUM).

Masdevallia arminii Linden & Rchb.f., Bonplandia (Hannover) 2: 283. 1854.

Illustration: Luer 2002b, pl. 397, pl. 398.

Distribution: *COL.

Type: Colombia, Schlim 740 [lectotype (selected here), W-R-0050826/W-0254007; isolectotypes, G-00168822, G00168823, K-000077775, K-000880305, K-L, P-00493092].

Note: Both Schlim 740 (Ocaña) and Schlim 1182 (Pamplona) are cited in the protologue. The first is selected as lectotype.

Masdevallia assurgens Luer & R.Escobar, Orquideología 13(92): 153. 1979.

Illustration: Luer & Escobar 1979, p. 154, p. 155, p. 168 (bottom left); Luer 2001a, pl. 262, pl. 263.

Distribution: *COL.

Type: Colombia, Escobar 830 (holotype, JAUM; isotype, SEL).

Masdevallia bicolor Poepp. & Endl., Nov. Gen. Sp. Pl. 2: 6, pl. 108B. 1837. Alaticaulia bicolor (Poepp. & Endl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 4. 2006.

Illustration: Luer 2000b, pl. 8-11; Vásquez & Ibisch 2000, f. 26-28, pl. 60.

Distribution: BOL COL ECU PER VEN.

Type: Peru, Poeppig 1662 (holotype, W-0023683).

Voucher: Colombia, Warszewicz s.n. (W).

Masdevallia bonplandii Rchb.f., Bonplandia (Hannover) 3: 69. 1855. Byrsella bonplandii (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006. Masdevallia coriacea subsp. bonplandii (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 16: 23. 1986.

Illustration: Luer 2000c, pl. 137, pl. 138.

Distribution: COL ECU PER.

Type: Ecuador, Humboldt & Bonpland s.n. (holotype, P; isotype, G).

Voucher: Colombia, Luer 7879 (SEL).

Masdevallia brockmuelleri Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 87: 805-806, pl. 403. 2002.

Illustration: Luer 2002b, pl. 403.

Distribution: *COL.

Type: Colombia, Luer 10409 (holotype, MO).

Masdevallia buccinator Rchb.f. & Warsz., Bonplandia (Hannover) 2: 115. 1854. Alaticaulia buccinator (Rchb.f. & Warsz.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 4. 2006.

Illustration: Escobar 1983, p. 84, p. 85; Luer 2000b, pl. 15.

Distribution: *COL.

Type: Colombia, Schlim 1181 (holotype, W-R47217; isotypes, BR-0000006570781, P-00493089).

Masdevallia bucculenta Luer, Lindleyana 10: 115. 1995. Buccella bucculenta (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 7. 2006.

Illustration: Luer 2003b, pl 634. Fig. 12D.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9862 (holotype, MO).

Voucher: Colombia, Galindo et al. 1476 (CUVC).

Masdevallia cacodes Luer & R.Escobar, Selbyana 7: 70. 1982. Byrsella cacodes (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 140.

Distribution: *COL.

Type: Colombia, Luer 1841 (holotype, SEL; isotype, JAUM).

Masdevallia caesia Roezl, Orchidophile 1: 599. 1881. Byrsella caesia (Roezl) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 141.

Distribution: *COL.

Type: Colombia, Roezl s.n. Colombia, Parque Nacional Natural Farallones de Cali, Restrepo 016 [neotype (selected here), FAUC-spirit; Fig. 11].

Note: No type specimen was apparently prepared.

Masdevallia campyloglossa Rchb.f., Gard. Chron., n.s. 10: 588. 1878. Byrsella campyloglossa (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 142-145.

Distribution: COL ECU PER.

Type: Colombia, Veitch 47 (holotype, W-R-2344/W-0254009; type ill., W-R23267/W-0254010).

Masdevallia carrilloana P.Ortiz, Orquideología 26: 9. 2009.

Illustration: Ortiz 2009a, p. 7 (top), p. 9.

Distribution: *COL.

Type: Colombia, Ortiz 1304 (holotype, HPUJ).

Note: Described as M. carrillana P.Ortiz.

Masdevallia caudata Lindl., Gen. Sp. Orchid. Pl. 193. 1833.

Illustration: Luer 2002b, pl. 404, pl. 405; Gil & Oses 2020a, f. 44.

Distribution: COL VEN.

Type: Colombia, Goudot s.n. (holotype, K; tracing, W).

Masdevallia caudivolvula Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 128. 1922. Streptoura caudivolvula (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 16. 2006.

Illustration: Luer 2003b, pl. 648.

Distribution: *COL.

Type: Colombia, Kalbreyer 1731 (holotype, B, destroyed). Colombia, Schmidtchen s.n. [lectotype (selected here), W-R-0010785, illustr.].

Note: Luer (2003b) selected Schmidtchen s.n. at W-R as lectotype. In the protologue, Kraenzlin indeed cites having seen Schmidtchen’s material at Reichenbach’s herbarium. However, he only refers to an illustration (W-R-10785), not a specimen (WR-11179), and in fact annotates the latter as type. Luer did not explicitly state which he selected, but he annotated the specimen as “neotype”. We specifically select the illustration as lectotype instead.

Masdevallia cerastes Luer & R.Escobar, Orquideología 13(1): 51-55. 1978. Megema cerastes (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer & Escobar 1978, p. 52, p. 53, p. 57 (bottom); Luer 2003b, pl. 609.

Distribution: *COL.

Type: Colombia, Luer 2785 (holotype, JAUM; isotype, SEL).

Masdevallia chimboensis Kraenzl., Bull. Misc. Inform. Kew 1925: 106. 1925. Luzama chimboensis (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 10. 2006.

Illustration: Luer 2000c, pl. 224.

Distribution: *COL.

Type: Colombia, Lehmann 6748 (holotype, K-000077875).

Masdevallia civilis Rchb.f. & Warsz., Bonplandia (Hannover) 2: 115. 1854. Byrsella civilis (Rchb.f. & Warsz.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 146.

Distribution: COL PER.

Type: Peru, Warszewicz 21 [lectotype (selected here), W-R-0038730, illustr. top right].

Voucher: Colombia, Ospina 578 (JAUM).

Note: Luer (2000c) argues that the types of M. civilis and M. rufolutea Lindl., nom. nud. are different elements in a mixed collection by Warszewicz. The sheet at W-R is indeed a mix of elements, but what Luer labelled as type of M. civilis is a collection by Roezl. The color illustration by Reichenbach, which Luer excludes from his concept of M. civilis, agrees with the protologue and is here selected as lectotype.

Masdevallia clandestina Luer & R.Escobar, Selbyana 7: 71. 1982.

Illustration: Escobar 1983, p. 95; Luer 2001a, pl. 267.

Distribution: COL VEN.

Type: Colombia, Luer et al. 7975 (holotype, SEL).

Masdevallia coccinea Linden ex Lindl., Orchid. Linden. 5, n. 26. 1846.

Illustration: Williams & Williams 1894, p. 495; Escobar 1983, p. 97; Luer 2003b, pl. 545-547; Karremans & Moreno 2019, f. 31.

Distribution: *COL.

Type: Colombia, Linden 1262 (holotype, K-L000880325; isotypes, BR-0000006570132, W-R-47365).

Masdevallia copiosa Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 129. 1922.

Distribution: *COL.

Type: Colombia, Kalbreyer 736 (holotype, B, destroyed?).

Note: An obscure name not monographed by Luer.

Masdevallia coriacea Lindl., Ann. Mag. Nat. Hist. 15: 257. 1845. Byrsella coriacea (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Escobar 1983, p. 74; Luer 2000c, pl. 149, pl. 150; Gil & Oses 2020b, f. 45

Distribution: COL ECU PER.

Type: Colombia, Hartweg s.n. (holotype, K-L; isotype, W).

Masdevallia corniculata Rchb.f., Gard. Chron., n.s. 9: 72. 1878. Megema corniculata (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2003b, pl. 610.

Distribution: COL ECU.

Type: Colombia, Backhouse s.n. (holotype, W-R50959).

Masdevallia crescenticola F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 428. 1921. Acinopetala crescenticola (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer 2001a, pl. 371.

Distribution: COL ECU.

Type: Ecuador, Lehmann 230 (holotype, W-R0083518; isotype, G-00168810).

Voucher: Colombia, Escobar 1459 (JAUM).

Masdevallia cucullata Lindl., Orchid. Linden. 4. 1846. Megema cucullata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2003b, pl. 611.

Distribution: COL ECU.

Type: Colombia, Linden 869 (holotype, K-L; isotypes, BR-0000006570163, GENT-10168608, W-R-50966).

Masdevallia densiflora Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 77. 1920. Spilotantha densiflora (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Mansfeld 1929, pl. 26 (n. 96).

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 26, No. 96)].

Note: An obscure name not monographed by Luer.

Masdevallia discolorLuer & R.Escobar, Orquideología 15(2-3): 129. 1982.

Illustration: Luer & Escobar 1982, p. 130; Luer 2002b, pl. 413, pl. 414.

Distribution: *COL.

Type: Colombia, Escobar 1915 (holotype, JAUM; isotype, SEL).

Masdevallia dryada Luer & R.Escobar, Lindleyana 4(3): 108, 111. 1989. Petalodon dryadus (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2000c, pl. 188.

Distribution: *COL.

Type: Colombia, Luer et al. 14161 (holotype, MO; isotypes, JAUM, K).

Masdevallia elephanticeps Rchb.f. & Warsz., Bonplandia (Hannover) 2: 116. 1854. Byrsella elephanticeps (Rchb.f. & Warsz.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 151.

Distribution: *COL.

Type: Colombia, Warszewicz s.n. [lectotype (selected here), W-R-0038728, illustr.].

Masdevallia encephala Luer & R.Escobar, Orquideología 13(1): 56. 1978.

Illustration: Luer & Escobar 1978, p. 57 (top), p. 68, p. 69; Luer 2001a, pl. 280. Fig. 12E; Gutiérrez Morales & Karremans 2017, f. 7.

Distribution: *COL.

Type: Colombia, Escobar 1682 (holotype, JAUM).

Masdevallia ensata Rchb.f., Linnaea 22: 818. 1850. Masdevallia xiphium Rchb.f. ex Kraenzl., Repert. Spec Nov. Regni Veg. Beih. 34: 78. 1925.

Illustration: Luer 2001 b, pl. 281.

Distribution: COL VEN.

Type: Venezuela, Funk & Schlim 1543 (holotype, W-R-0039206; isotypes, BM-000082155, BR-0000006570491, BR-0000006571269, G-00168802, G-00168801, G-00168803, K-L!, LE-00006499, P-00493062, W-R-38794; type illustr., AMES-00101245).

Voucher: Colombia, Ortiz-Valdivieso 930 (HPUJ).

Masdevallia estradae Rchb.f., Gard. Chron. 2: 435. 1874.

Illustration: Luer 2002b, pl. 416; Williams & Williams 1894, p. 488.

Distribution: *COL.

Type: Colombia, Wallis s.n. (holotype, W-R-50934).

Masdevallia exilipes Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 79. 1920.

Illustration: Mansfeld 1929, pl. 26 (n. 98).

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 26, No. 98)].

Note: An obscure name not monographed by Luer.

Masdevallia expansa Rchb.f., Otia Bot. Hamburg. 16. 1878.

Illustration: Luer 2002b, pl. 419.

Distribution: *COL.

Type: Colombia, Schlim 1182 [lectotype, W-R50818 (Luer 2002b); isolectotype, G].

Masdevallia falcago Rchb.f., Otia Bot. Hamburg. 16. 1878.

Illustration: Luer 2001a, pl. 283.

Distribution: *COL.

Type: Colombia, Purdie s.n. (holotype, W-R0051153; isotypes, K-000573975, K-000573976).

Masdevallia fasciata Rchb.f., Gard. Chron., n.s. 15(1): 202. 1881. Rodrigoa fasciata (Rchb.f.) Braas, Orchidee (Hamburg) 30(6): 218. 1979.

Illustration: Luer 2003b, pl. 623.

Distribution: *COL.

Type: Colombia, Schmidtchen s.n. [lectotype (selected here), W-R-0014091; type illustr., W-R0051150].

Masdevallia filaria Luer & R.Escobar, Orquideología 13(1): 70. 1978. Masdevallia ventricularia subsp. filaria (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 16: 36. 1986.

Illustration: Luer & Escobar 1978, p. 64 (top, bottom right), p. 72, p. 73; Luer 2002b, pl. 511.

Distribution: COL ECU.

Type: Colombia, Escobar 456E (holotype, JAUM).

Masdevallia floribunda Lindl., Edwards’s Bot. Reg. 29: misc. 72. 1843. Acinopetala floribunda (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006. Masdevallia floribunda subsp. floribunda.

Illustration: Luer 2001a, pl. 373-376; Bogarín et al. 2017, f. 5.

Distribution: BEL COL COS GUA HON MEX.

Type: Mexico, Rogers s.n. (holotype, K-L000463103; tracing of type, W-R-19726).

Voucher: Colombia, unknown collector (LE).

Note: Mostly known from northern Central America, however a cultivated specimen of Colombian origin is cited by Luer (2001b).

Masdevallia foetens Luer & R.Escobar, Orquideología 13(2): 157. 1979. Byrsella foetens (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer & Escobar 1979, p. 160, p. 161, p. 169 (middle); Luer 2000c, pl. 152.

Distribution: *COL.

Type: Colombia, Escobar 1072 (holotype, JAUM).

Masdevallia fragrans Woolward, Genus Masdevallia (3): , t. 18. 1896. Byrsella fragrans (Woolward) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Escobar 1983, p. 73; Luer 2000c, pl. 155.

Distribution: *COL.

Type: Colombia, Woolward s.n. (holotype, BM000084327).

Masdevallia gargantua Rchb.f., Gard. Chron., n.s. 6: 516. 1876. Byrsella gargantua (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 156, pl. 157.

Distribution: *COL.

Type: Colombia, Wallis sub Veitch s.n. (holotype, W-R-0038742).

Masdevallia geminiflora P.Ortiz, Orquideología 13: 75. 1978. Acinopetala geminiflora (P.Ortiz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Ortiz 1981b, p. 226, p. 227; Luer 2001a, pl. 377.

Distribution: COL ECU.

Type: Colombia, Ortiz 607 (holotype, COL).

Masdevallia gerhardii Luer & Sijm, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 142. 2010.

Illustration: Luer 2010, f. 9 (p. 151).

Distribution: *COL.

Type: Colombia, Sijm 290814 (holotype, MO).

Masdevallia gilbertoi Luer & R.Escobar, Orquideología 13(1): 75. 1978.

Illustration: Luer & Escobar 1978, p. 59 (top), p. 76, p. 77; Luer 2001a, pl. 287.

Distribution: *COL.

Type: Colombia, Escobar 440 (holotype, JAUM).

Masdevallia hercules Luer & Andreetta, Lindleyana 3(4): 198. 1988 [1989]. Megema hercules (Luer & Andreetta) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2003b, pl. 613.

Distribution: COL ECU.

Type: Ecuador, Luer 13256 (holotype, MO).

Voucher: Colombia, Schultes & Villarreal 5219 (AMES).

Masdevallia herradurae F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 455. 1898. Acinopetala herradurae (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer 2001a, pl. 379.

Distribution: *COL.

Type: Colombia, Lehmann 4139 (holotype, K; isotypes, AMES, G, NY, US).

Masdevallia heteroptera Rchb.f., Gard. Chron., n.s. 3(1): 590. 1875. Rodrigoa heteroptera (Rchb.f.) Braas, Orchidee (Hamburg) 30(6): 218. 1979.

Illustration: Luer 2003b, pl. 624.

Distribution: *COL.

Type: Colombia, Norman s.n. [lectotype, W-R51149, flower bottom right (Luer 2003b); type illustr., W-R-0051149, top left]. Colombia, Patin s.n. (syntype, W-R-0051149, center and bottom left).

Masdevallia hians Lindl. & Rchb.f., Bonplandia (Hannover) 2: 283. 1854.

Illustration: Luer 2001a, pl. 294.

Distribution: *COL.

Type: Colombia, Schlim 1163 (holotype, W-R50110).

Masdevallia hieroglyphica Rchb.f., Gard. Chron. 2: 230. 1882.

Illustration: Luer 2001a, pl. 295.

Distribution: *COL.

Type: Colombia, Arnold 82 [lectotype (selected here), W-R-0050819, bottom left flowers].

Masdevallia hoeijeri Luer & Hirtz, Lindleyana 1(3): 184. 1986. Diodonopsis hoeijeri (Luer & Hirtz) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001. Pteroon hoeijeri (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 12. 2006.

Illustration: Luer 2000c, pl. 230.

Distribution: COL ECU.

Type: Ecuador, Luer 11905 (holotype, MO).

Voucher: Colombia, Escobar 4064 (MO).

Masdevallia hortensis Luer & R.Escobar, Orquideología 16(2): 154. 1984. Rodrigoa hortensis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 14. 2006.

Illustration: Luer & Escobar 1984b, p. 155, p. 156; Luer 2003b, pl. 625. Fig. 12F; Vieira-Uribe & Karremans 2017c, f. 8.

Distribution: *COL.

Type: Colombia, Escobar et al. 2707 (holotype, SEL; isotypes, COL, JAUM).

Masdevallia hubeinii Luer & Würstle, Orchideën 48: 50. 1986.

Illustration: Luer 2002b, pl. 421.

Distribution: *COL.

Type: Colombia, Würstle 1324 (holotype, MO).

Masdevallia hylodes Luer & R.Escobar, Orquideología 13(2): 162. 1979. Byrsella hylodes (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer & Escobar 1979, p. 164, p. 165, p. 170 (top); Luer 2000c, pl. 158.

Distribution: *COL.

Type: Colombia, Escobar et al. 1899 (holotype, JAUM; isotype, SEL).

Masdevallia ignea Rchb.f., Gard. Chron. 1871: 1482. 1871.

Illustration: Escobar 1983, p. 71; Luer 2003b, pl. 552-554.

Distribution: *COL.

Type: Colombia, Veitch s.n. [lectotype (selected here), W-R-0047372]. Colombia, Day s.n. (syntype, W-R-0047375).

Masdevallia impostor Luer & R.Escobar, Orquideología 13(2): 170. 1979. Alaticaulia impostor (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Luer & Escobar 1979, p. 168 (top left), p. 172, p. 173; Luer 2000b, pl. 45. Fig. 12G.

Distribution: COL ECU VEN.

Type: Colombia, Segura s.n. (holotype, JAUM; isotype, SEL).

Masdevallia indecora Luer & R.Escobar, Novon 1(4): 170. 1991. Luzama indecora (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 10. 2006.

Illustration: Luer 2000c, pl. 231.

Distribution: *COL.

Type: Colombia, Luer 15265 (holotype, JAUM; isotype, MO).

Masdevallia jaderiS.Vieira-Uribe & Bogarín, Orquideología 33(1): 16. 2016.

Illustration: Vieira-Uribe & Bogarín 2016, f. 1, f. 2; Domínguez Vargas & Bogarín 2018, f. 19.

Distribution: *COL.

Type: Colombia, Vieira 21 (holotype, JAUM).

Masdevallia klabochiorum Rchb.f., Gard. Chron. 1: 270. 1876. Masdevallia xanthina subsp. klabochiorum (Rchb.f.) Luer, Lindleyana 3(1): 64. 1988.

Illustration: Escobar 1983, p. 93; Luer 2002b, pl. 427, pl. 428; Gutiérrez Morales & Sierra-Ariza 2020, f. 46.

Distribution: COL ECU PER.

Type: Colombia, Klaboch s.n. (holotype, W-R50830).

Note: Published as M. klabochorum.

Masdevallia lamia Luer & Hirtz, Lindleyana 3(1): 40. 1988. Buccella lamia (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 7. 2006.

Illustration: Luer 2003b, pl. 636, pl. 637.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12383 (holotype, MO).

Voucher: A photo of a Colombian specimen is reproduced herein (Fig. 12D).

Masdevallia laevis Lindl., Ann. Mag. Nat. Hist. 15: 257. 1845.

Illustration: Escobar 1983, p. 80, p. 100; Luer 2001a, pl. 301-305.

Distribution: COL ECU.

Type: Colombia, Hartweg s.n. (holotype, K-L; isotype, W-R-0039199).

Masdevallia leonor-baeziana Os.Perd., Edwin Trujillo & Karremans, Lankesteriana 23(2): 140. 2023.

Illustration: Perdomo et al. 2023, f. 2, f. 3F.

Distribution: *COL.

Type: Colombia, Perdomo et al. OP465 (holotype, CUVC; isotype, HUAZ).

Masdevallia leontoglossa Rchb.f., Bonplandia (Hannover) 3: 69. 1855. Byrsella leontoglossa (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 160.

Distribution: *COL.

Type: Colombia, Wagener s.n. (holotype, W-0038745; type illustr., W-R-0038743).

Masdevallia leptoura Luer, Phytologia 54(5): 381. 1983. Masdevallia pachyura subsp. leptoura (Luer) Luer, Lindleyana 3(1): 44. 1988. Spilotantha leptoura (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Luer 2003b, pl. 589.

Distribution: COL ECU PER.

Type: Ecuador, Dalström 7264 (holotype, SEL).

Voucher: Colombia, Luer 17982 (MO).

Masdevallia × ligiae Luer & R.Escobar, Orquideología 15(2): 131. 1982.

Illustration: Luer & Escobar 1982, p. 132; Luer 2002b, pl. 514.

Distribution: *COL.

Type: Colombia, Luer 6727 (holotype, JAUM; isotypes, SEL).

Note: Described initially as Masdevallia ligiae, since then regarded as a hybrid (Luer 2002b).

Masdevallia livingstoneana Roezl ex Rchb.f., Gard. Chron., n.s. 2(37): 322. 1874. Acinopetala livingstoneana (Roezl ex Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 118. 2007. Acinopetala livingstoneana (Roezl ex Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006, nom. inval.

Illustration: Luer 2001a, pl. 381.

Distribution: COL COS PAN.

Type: Panama, Roezl 74 [lectotype (selected here), W-R-0019729/W-0028829, type illustr.].

Voucher: Colombia, Duke 9911 (NY).

Note: Roezl s.n. at W was selected as lectotype by Kolanowska (2014). However, there are several Roezl materials under M. livingstoneana in Reichenbach’s herbarium, all except the illustration labeled ‘Roezl 74’ were prepared after publication and are therefore strictly speaking not original material.

Masdevallia ludibunda Rchb.f., Gard. Chron. 1: 179. 1882. Masdevallia estradae var. ludibunda (Rchb.f.) Woolward, Masdevallia 13. 1892.

Illustration: Luer 2002b, pl. 430.

Distribution: *COL.

Type: Colombia, Lawrence s.n. [lectotype (selected here), W-R-0050820, illustr. bottom right].

Note: Sheet 50820 in Reichenbach’s herbarium contains 3 illustrations, two dissimilar leaves and a note. Only the illustration on the bottom right and the note can be confidently associated with the protologue.

Masdevallia ludibundella Luer & R.Escobar, Lindleyana 10(2): 120, 123. 1995.

Illustration: Luer 2002b, pl. 431.

Distribution: *COL.

Type: Colombia, Luer 16881 (holotype, MO).

Masdevallia macrogenia (Arango) Luer & R.Escobar, Orquideología 13(1): 79. 1978.

Masdevallia collina var. macrogenia O.J. Arango, Orquideología 11(1): 13. 1976. Petalodon macrogenius (O.J. Arango) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer & Escobar 1978, p. 62 (bottom right), p. 80, p. 81; Luer 2000c, pl. 189.

Distribution: *COL.

Type: Colombia, Arango 160 (holotype, JAUM).

Masdevallia macroglossa Rchb.f., Otia Bot. Hamburg. 1: 15. 1878. Byrsella macroglossa (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 162, pl. 163.

Distribution: COL VEN.

Type: Venezuela, Fendler 1373 [lectotype (selected here), W-R-0019734, tracing.; isolectotype, K-L].

Voucher: Colombia, Ortiz-Valdivieso 954 (HPUJ).

Masdevallia macrura Rchb.f., Gard. Chron., n.s. 1: 240. 1874. Megema macrura (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2000c, pl. 164.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W-R0050960).

Voucher: Colombia, Luer 10244 (MO).

Masdevallia mandarina (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 87: 857. 2002. Masdevallia xanthina subsp. mandarina Luer & R.Escobar, Lindleyana 3(1): 64. 1988.

Illustration: Luer 2002b, pl. 434.

Distribution: COL ECU.

Type: Colombia, Luer et al. 11383 (holotype, MO; isotype, JAUM).

Masdevallia marthae Luer & R.Escobar, Orquideología 13(1): 84. 1978.

Illustration: Luer & Escobar 1978, p. 64 (bottom left), p. 86, p. 87; Luer 2002b, pl. 516.

Distribution: *COL.

Type: Colombia, Escobar 452 (holotype, JAUM).

Masdevallia mastodon Rchb.f., Bonplandia (Hannover) 3: 69. 1855.

Illustration: Escobar 1983, p. 77; Luer 2001a, pl. 315.

Distribution: *COL.

Type: Colombia, Wagener s.n. (holotype, W-R-47216).

Masdevallia medusa Luer & R.Escobar, Selbyana 7: 72. 1982. Alaticaulia medusa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Escobar 1983, p. 99; Luer 2000b, pl. 59.

Distribution: *COL.

Type: Colombia, Luer et al. 8025 (holotype, SEL; isotype, JAUM).

Masdevallia mejiana Garay, Orquideología 5: 17. 1970. Reichantha mejiana (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 13. 2006.

Illustration: Luer 2000c, pl. 209.

Distribution: *COL.

Type: Colombia, Escobar 426 (holotype, AMES).

Masdevallia melanoxantha Linden & Rchb.f., Bonplandia (Hannover) 2: 283. 1854. Alaticaulia melanoxantha (Linden & Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Luer 2000b, pl. 60.

Distribution: COL VEN.

Type: Colombia, Schlim 1166 (holotype, W-R0047240).

Masdevallia meleagris Lindl., Ann. Mag. Nat. Hist. 15: 257. 1845. Rodrigoa meleagris (Lindl. ex Rchb.f.) Braas, Orchidee (Hamburg) 30(6): 218. 1979.

Illustration: Luer 2003b, pl. 626.

Distribution: *COL.

Type: Colombia, Hartweg s.n. (holotype, K-L).

Masdevallia minuta Lindl., Ann. Mag. Nat. Hist. 12: 396. 1843. Acinopetala minuta (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer 2001a, pl. 382, pl. 383; Vásquez & Ibisch 2000, pl. 79.

Distribution: BOL BRA COL ECU FRE GUY PER SUR VEN.

Type: Surinam, Hostmann 151 (holotype, K-L; isotypes, BM-000082164, G, LE, W-R, W-R-0050770; type illustr., W-R-0019708).

Voucher: Colombia, Dawe 243 (K).

Masdevallia mirandae Gal.-Tar., Haelt. & Zuluaga, Phytotaxa 502(3): 278, 281. 2021.

Illustration: Galindo-Tarazona et al. 2021, f. 1-2.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona 1478 (holotype, CUVC; isotype, COL).

Masdevallia misasii Braas, Orchidee (Hamburg) 33(4): 148. 1982. Byrsella misasii (Braas) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006. Masdevallia reflexa Misas, Orquideología 12(23): 149. 1978, nom. illeg. Non Masdevallia reflexa Schltr. [1923].

Illustration: Misas Urreta 1977, p. 151, p. 153; Luer 2000c, pl. 167; Oses & Gutiérrez Morales 2018, f. 20.

Distribution: *COL.

Type: Colombia, Laserna s.n. (holotype, JAUM).

Masdevallia molossus Rchb.f., Linnaea 41: 10. 1877. Buccella molossus (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 7. 2006.

Illustration: Luer 2003b, pl. 639.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W-R51154).

Masdevallia mooreana Rchb.f., Gard. Chron., n.s. 21: 408. 1884.

Byrsella mooreana (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 168, pl. 169.

Distribution: *COL.

Type: Unknown country, Moore s.n. (holotype, WR-0030394/W-0075796; isotypes, BM, K; type illustr., W-R-0038789/0075797A).

Voucher: Colombia, Escobar 480 (JAUM).

Masdevallia mutica Luer & R.Escobar, Orquideología 13(1): 88. 1978. Fissia mutica (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 9. 2006.

Illustration: Luer & Escobar 1978, p. 60 (top right), p. 90, p. 91; Luer 2003b, pl. 615A, pl. 615.

Distribution: *COL.

Type: Colombia, Escobar 1483 (holotype, JAUM).

Masdevallia × mystica Luer, Lindleyana 4(3): 119. 1989.

Illustration: Luer 2002b, pl. 520.

Distribution: *COL.

Type: Colombia, Luer 13793 (holotype, MO).

Masdevallia nidifica Rchb.f., Otia Bot. Hamburg. 1: 18. 1878. Buccella nidifica (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 7. 2006.

Illustration: Luer 2003b, pl. 640.

Distribution: COL COS ECU NIC PAN.

Type: Ecuador, Lehmann 24 [lectotype, W-R0051151/W-0028827 (Luer 2003b), top right specimen (‘Marz 1877’)].

Voucher: Colombia, Ortiz-Valdivieso 4267 (HPUJ).

Masdevallia niesseniae Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 110, pl. 35. 1998.

Illustration: Luer 2003b, pl. 554.

Distribution: *COL.

Type: Colombia, Luer 18789 (holotype, MO).

Masdevallia nivea (Luer & R.Escobar) Luer & R.Escobar, Thesaurus Masdevalliarum 13: t. 9. 1989. Masdevallia purpurella subsp. nivea Luer & R.Escobar, Lindleyana 4: 55. 1988.

Illustration: Luer 2002b, pl. 436.

Distribution: *COL.

Type: Colombia, Luer 12960 (holotype, MO).

Masdevallia norae Luer, Lindleyana 3: 44. 1988. Alaticaulia norae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Luer 2000b, pl. 64.

Distribution: COL VEN.

Type: Venezuela, Luer 8082 (holotype, SEL).

Voucher: Colombia, Plowman 2430 (AMES).

Masdevallia odontocera Luer & R.Escobar, Selbyana 7: 73. 1982.

Illustration: Escobar 1983, p. 91; Luer 2001a, pl. 322.

Distribution: *COL.

Type: Colombia, Luer et al. 7933 (holotype, SEL; isotypes, AAU, COL, JAUM).

Masdevallia orchipayanensium P.Ortiz, Orquideología 23(2): 73. 2004.

Illustration: Ortiz 2004, p. 75, p. 76.

Distribution: *COL.

Type: Colombia, Ortiz 1210 (holotype, HPUJ).

Masdevallia os-draconis Luer & R.Escobar, Orquideología 13(1): 92. 1978.

Illustration: Luer & Escobar 1978, p. 59 (bottom), p. 94, p. 95; Luer 2002b, pl. 523.

Distribution: *COL.

Type: Colombia, Escobar 1766 (holotype, JAUM).

Masdevallia oscarii Luer & R.Escobar, Orquideología 13(2): 175. 1979. Byrsella oscarii (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer & Escobar 1979, p. 168 (bottom right), p. 176, p. 177; Luer 2000c, pl. 171.

Distribution: *COL.

Type: Colombia, Escobar 849 (holotype, JAUM).

Masdevallia pachyantha Rchb.f., Gard. Chron., n.s. 21: 174. 1884. Byrsella pachyantha (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 172.

Distribution: *COL.

Type: Colombia, Cross s.n. [lectotype (selected here), W-R-38738; isolectotype, BM-000082150].

Masdevallia pachysepala (Rchb.f.) Luer, Lindleyana 9(4): 245. 1994. Byrsella pachysepala (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006, nom. inval. Masdevallia elephanticeps fo. pachysepala (Rchb.f.) O. Gruss & M. Wolff, Orchid Atlas 213. 2007. Masdevallia elephanticeps var. pachysepala Rchb.f., Bonplandia (Hannover) 2: 283. 1854.

Illustration: Luer 2000c, pl. 173.

Distribution: COL VEN.

Type: Colombia, Schlim 410 (holotype, W-R0038729; isotypes, BR-0000006571603, G-00168809, G-00168808, G-00168807, K-L000880310, P-00493017).

Masdevallia pardina Rchb.f., Otia Bot. Hamburg. 1: 15. 1878. Byrsella pardina (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 174.

Distribution: COL ECU.

Type: Colombia, Chesterton s.n. (holotype, W-R0039198).

Masdevallia parsonsii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 241, pl. 701. 2009.

Illustration: Luer 2009a, pl. 701.

Distribution: *COL.

Type: Colombia, Luer 21302 (holotype, MO).

Masdevallia parvula Schltr., Repert. Spec. Nov. Regni Veg. Beih. 8: 49. 1921. Rodrigoa parvula (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 14. 2006.

Illustration: Luer 2003b, pl. 629, pl. 630.

Distribution: BOL COL ECU PER.

Type: Ecuador, Sodiro s.n. (holotype, B, destroyed). Ecuador, Jost 1678 [neotype, MO (Luer 2003b)].

Voucher: Colombia, Escobar 8344 (JAUM, MO)

Masdevallia pastinata Luer, Harvard Pap. Bot. 2(1): 60, f. 4. 1997. Alaticaulia pastinata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Luer 2000b, pl. 69.

Distribution: *COL.

Type: Colombia, Luer 18425 (holotype, MO).

Masdevallia peristeria Rchb.f., Gard. Chron., n.s. 1: 500. 1874. Byrsella peristeria (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006. Masdevallia peristeria subsp. peristeria.

Illustration: Luer 2000c, pl. 175, pl. 176.

Distribution: COL ECU.

Type: Colombia, Veitch 288 (holotype, W-R38734; type illustr., W-R-38733, top right).

Masdevallia pescadoensis Luer & R.Escobar, Orquideología 13(2): 179. 1979. Acinopetala pescadoensis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer & Escobar 1979, p. 167 (bottom left), p. 180, p. 181; Luer 2001a, pl. 385.

Distribution: *COL.

Type: Colombia, Escobar 1092 (holotype, JAUM; isotypes, SEL).

Masdevallia picturata Rchb.f., Otia Bot. Hamburg. 1: 16. 1878. Fissia picturata (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 9. 2006.

Illustration: Vásquez & Ibisch 2000, f. 42, pl. 87; Luer 2003b, pl. 616-618; Ortiz & Medina 2012, f. 8; Moreno 2019b, f. 32.

Distribution: BOL COL COS ECU GUY PAN PER VEN.

Type: Venezuela, Wagener s.n. [lectotype (selected here), W-R-0050825/W-0028826, type illustr.; isolectotype, W-R-347, Xenia Orchid. I t. 75(1)].

Voucher: Colombia, Ortiz-Valdivieso 64 (HPUJ).

Masdevallia pileata Luer & Würstle, Orchideën 48: 50. 1986.

Illustration: Luer 2002b, pl. 441.

Distribution: *COL.

Type: Colombia, Luer 8132 (holotype, SEL).

Masdevallia planadensis Luer & R.Escobar, Lindleyana 3(1): 50. 1988. Rodrigoa planadensis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 14. 2006.

Illustration: Luer 2003b, pl. 631.

Distribution: COL ECU.

Type: Colombia, Luer 12488 (holotype, MO).

Masdevallia platyglossa Rchb.f., Gard. Chron., n.s. 18: 552. 1882. Byrsella platyglossa (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 178.

Distribution: COL ECU.

Type: Colombia, Lawrence s.n. (holotype, W-R0038737).

Masdevallia posadae Luer & R.Escobar, Selbyana 7: 74. 1982. Alaticaulia posadae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 5. 2006.

Illustration: Luer 2000b, pl. 74, pl. 75; Doucette et al. 2016b, f. 4, f. 11c.

Distribution: COL ECU PER.

Type: Colombia, Luer 2936 (holotype, SEL; isotype, JAUM).

Masdevallia pteroglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 81. 1920. Masdevallia wageneriana var. pteroglossa (Schltr.) Braas, Orchidee (Hamburg) 33: 148. 1982.

Illustration: Mansfeld 1929, pl. 27 (n. 101); Braas 1977, p. 28, p. 29, p. 49; Luer 2002b, pl. 489.

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 27, No. 101)]. Colombia, Luer 10405 [neotype, MO (Luer 2002b), here rejected and selected as epitype].

Note: We follow Pupulin et al. (2022) in recognizing Schlechter’s floral analyses reproduced by Mansfeld as original material and therefore reject Luer’s neotype in favor of a lectotype.

Masdevallia pterygiophora Luer & R.Escobar, Lindleyana 3(1): 50. 1988. Diodonopsis pterygiophora (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 253. 2001. Pteroon pterygiophorum (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 12. 2006.

Illustration: Luer 2000c, pl. 241.

Distribution: *COL.

Type: Colombia, Luer 2288 (holotype, SEL).

Masdevallia pumila Poepp. & Endl., Nov. Gen. Sp. Pl. 2: 6, pl. 108C. 1837.

Illustration: Luer 2001a, pl. 330; Vásquez & Ibisch 2000, pl. 88.

Distribution: BOL COL ECU PER.

Type: Peru, Poeppig s.n. (holotype, W-R-0023685; isotypes, K-L, W-R-0050753; type illustr., W-R19707, W-R-0023684).

Voucher: Colombia, Ortiz-Valdivieso 442 (HPUJ).

Masdevallia purpurellaLuer & R.Escobar, Orquideología 15(2-3): 136. 1982.

Illustration: Luer & Escobar 1982, p. 137, 138; Luer 2002b, pl. 444.

Distribution: *COL.

Type: Colombia, Escobar 1469 (holotype, JAUM; isotype, SEL).

Masdevallia racemosa Lindl., Ann. Mag. Nat. Hist. 15: 256. 1845. Spectaculum racemosum (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 14. 2006.

Illustration: Luer 2003b, pl. 564; Moreno & Díaz-Morales 2018, f. 21; Galindo-Tarazona et al. 2021, f. 3.

Distribution: *COL.

Type: Colombia, Hartweg 1432 (holotype, K-L; isotypes, BM-000082142, G-00168770, K-000077755, K-000573980, LD-1404037, P-00493011, W-R0047336, W-R; type tracing, W-R-0019727).

Masdevallia renzii Luer, Lindleyana 10(4): 232. 1995.

Illustration: Luer 2002b, pl. 445.

Distribution: *COL.

Type: Colombia, Renz 8461 (holotype, BAS).

Masdevallia rhinophora Luer & R.Escobar, Lindleyana 10(2): 126, f. 7. 1995.

Illustration: Luer 2002b, pl. 447.

Distribution: *COL.

Type: Colombia, Luer 16879 (holotype, MO).

Masdevallia rosea Lindl., Ann. Mag. Nat. Hist. 15: 257. 1845. Masdevallia rosea subsp. rosea.

Illustration: Luer 2003b, pl. 555, pl. 556; Williams & Williams 1894, p. 502.

Distribution: COL ECU.

Type: Ecuador, Hartweg s.n. (holotype, K-L; isotype, W-R-0047338).

Voucher: Colombia, Lehmann 203 (AMES, BM, G, K, LE, NY).

Masdevallia saltatrix Rchb.f., Linnaea 41: 10. 1877.

Illustration: Luer 2002b, pl. 527.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W-R0050953).

Masdevallia sanctae-fidei Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 414. 1921. Alaticaulia sanctae-fidei (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 81, pl. 82.

Distribution: COL VEN.

Type: Colombia, Man 1015 [lectotype (selected here), W-R-0014963; isolectotypes, HBG-501774, W-R-0013474].

Note: There are three sheets annotated by Kraenzlin himself as “Typus”. These were communicated to him by Sander, and all presumably originally collected by Man. However, a single sheet actually cites Man, it is selected as lectotype as per the protologue.

Masdevallia sanctae-rosae Kraenzl., Bull. Misc. Inform. Kew 102. 1925. Byrsella sanctae-rosae (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006. Pleurothallis sanctae-rosae (Kraenzl.) Garay, Orquideología 3: 17. 1968.

Illustration: Luer 2000c, pl. 181.

Distribution: *COL.

Type: Colombia, Lehmann 7269 (holotype, K; isotypes, AMES, LE, NY).

Masdevallia sceptrum Rchb.f., Bonplandia (Hannover) 2: 283. 1854. Alaticaulia sceptrum (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006. Masdevallia schlimii var. sceptrum (Rchb.f.) Woolward, Masdevallia 8. 1890.

Illustration: Luer 2000b, pl. 84, pl. 85.

Distribution: COL VEN.

Type: Colombia, Schlim 1200 (holotype, W-R0047250; isotype, BR-0000006570484).

Masdevallia schizantha Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 134. 1922.

Illustration: Luer 2002b, pl. 451.

Distribution: *COL.

Type: Colombia, Kalbreyer 1095 (holotype, B, destroyed). Colombia, Luer et al. 10327 [neotype, MO (Luer 2002b)].

Masdevallia schizopetala Kraenzl., Bull. Misc. Inform. Kew 1925(3): 101-102. 1925. Acinopetala schizopetala (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer 2000c, pl. 246, pl. 247.

Distribution: BOL COL COS PAN.

Type: Colombia, Lehmann 3836 (holotype, K-000077758; isotype, HBG-501779).

Masdevallia schlimii Linden ex Lindl., Orchid. Linden.: 5. 1846. Alaticaulia schlimii (Linden ex Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 128.

Distribution: COL VEN.

Type: Venezuela, Linden 666 (holotype, K-L; isotypes, BR-0000006570811, W-R-12766!, W-R-0047251, W-R-0047243, left infl.; type illustr. W-R-0047255).

Voucher: Colombia, Luer 2328 (SEL).

Masdevallia schmidt-mummii Luer & R.Escobar, Orquideología 13(1): 96. 1978.

Illustration: Luer & Escobar 1978, p. 59 (middle), p. 98, p. 99; Luer 2002b, pl. 452.

Distribution: *COL.

Type: Colombia, Escobar 1822 (holotype, JAUM; isotype, SEL).

Masdevallia scobina Luer & R.Escobar, Selbyana 7: 75. 1982. Alaticaulia scobina (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 88.

Distribution: *COL.

Type: Colombia, Luer 1904 (holotype, SEL; isotype, JAUM).

Masdevallia segurae Luer & R.Escobar, Orquideología 13(1): 100. 1978. Rodrigoa segurae (Luer & R.Escobar) Braas, Orchidee (Hamburg) 30(6): 219. 1979.

Illustration: Luer & Escobar 1978, p. 62 (bottom left), p. 102, p. 103; Luer 2003b, pl. 632.

Distribution: *COL.

Type: Colombia, Escobar 1601 (holotype, JAUM).

Masdevallia senghasiana Luer, Lindleyana 9(4): 248. 1994.

Illustration: Ortiz 2000, 119 (p. 143).

Distribution: *COL.

Type: Colombia, Senghas 524 (holotype, HEID; isotype, MO).

Note: Also cited as a hybrid.

Masdevallia sernae Luer & R.Escobar, Orquideología 13(2): 183. 1979. Alaticaulia sernae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer & Escobar 1979, p. 169 (top), p. 184, p. 185; Luer 2000b, pl. 90.

Distribution: *COL.

Type: Colombia, Escobar 1820 (holotype, JAUM).

Masdevallia siphonantha Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 87: 1033-1034, pl. 528. 2002.

Illustration: Luer 2002b, pl. 528.

Distribution: *COL.

Type: Colombia, Luer 14906 (holotype, W).

Masdevallia stenorhynchos Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 130. 1922. Alaticaulia stenorhynchos (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 93; Sierra-Ariza & Rincón-González 2022, f. 55.

Distribution: *COL.

Type: Colombia, Kalbreyer 1528 (holotype, B, destroyed). Colombia, Luer 1862 (neotype, SEL).

Masdevallia × strumella Luer, Schlechteriana 1(2): 9, 13. 1988.

Illustration: Luer 2001a, pl. 339.

Distribution: *COL.

Type: Colombia, Luer et al. 10180 (holotype, MO).

Masdevallia strumifera Rchb.f., Flora 69: 560. 1886.

Illustration: Luer 2001a, pl. 340, pl. 341.

Distribution: COL ECU VEN.

Type: Colombia, Lehmann 2720 (holotype, W).

Masdevallia strumosa P.Ortiz & E.Calderón, Orquideología 22: 120. 2002.

Illustration: Ortiz & Calderón-Sáenz 2002, p. 122-123.

Distribution: *COL.

Type: Colombia, Calderón 110 (holotype, FMB; isotypes, COL, TULV, JAUM).

Masdevallia sumapazensis P.Ortiz, Orquideología 14(3): 223. 1981. Byrsella sumapazensis (P.Ortiz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Ortiz 1981b, p. 223, 225; Luer 2000c, pl. 184.

Distribution: *COL.

Type: Colombia, Ortiz 314 (holotype, COL).

Masdevallia teaguei Luer, Selbyana 2(4): 381. 1978. Jostia teaguei (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 79: 2-3. 2000.

Illustration: Ortiz 2009a, p. 7 (bottom); Luer 2000a, pl. 1; Luer 2003b, pl. 647.

Distribution: COL ECU.

Type: Ecuador, Luer 2035 (holotype, SEL).

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz (2009, p. 7).

Masdevallia thienii Dodson, Selbyana 2(1): 54, f. 15B. 1977. Alaticaulia thienii (Dodson) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 118. 2007. Reichantha thienii (Dodson) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 13. 2006.

Illustration: Luer 2000b, pl. 131.

Distribution: COL COS ECU PAN.

Type: Ecuador, Dodson & Thien 1133 (holotype, SEL).

Voucher: Colombia, Ortiz-Valdivieso 923 (HPUJ).

Masdevallia torta Rchb.f., Gard. Chron., n.s. 19: 110. 1883. Byrsella torta (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 185.

Distribution: *COL.

Type: Colombia, Bull s.n. (holotype; W-R-001414; type illustr., 38746).

Masdevallia triangularis Lindl., Orchid. Linden. 5. 1846.

Illustration: Luer 2002b, pl. 454-456.

Distribution: COL ECU VEN.

Type: Venezuela, Linden 1471 (holotype, K-L; isotypes, BM-0000923767, BR-0000006570507, G-00168761, W-R-50950; tracing of type, W-R0019743).

Voucher: Colombia, Schlim 1704 (K, LE).

Masdevallia tridens Rchb.f., Otia Bot. Hamburg. 1: 13. 1879. Spilotantha tridens (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Luer 2003b, pl. 604.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. [lectotype, K-L000880308 (Luer 2003b); isolectotype, W-R47364; tracing of type, W-R-19741, left].

Masdevallia trigonopetala Kraenzl., Bull. Misc. Inform. Kew 1925: 105. 1925.

Illustration: Luer 2001a, pl. 346.

Distribution: COL ECU.

Type: Colombia, Lehmann 345 (holotype, K-000077804; tracing of type, W-R-0008145).

Masdevallia trochilus Linden & André, Gard. Chron. 711. 1873. Alaticaulia trochilus (Linden & André) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 98; Luer 2003b, pl. 98 (p. 196a).

Distribution: COL ECU PER.

Type: Colombia, Wallis s.n. (holotype, W).

Masdevallia tsubotae Luer, Lindleyana 9(2): 114. 1994. Alaticaulia tsubotae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 100.

Distribution: *COL.

Type: Colombia, Luer 15515 (holotype, MO).

Masdevallia tubulosa Lindl., Orchid. Linden. 4. 1846.

Illustration: Luer 2001a, pl. 348-351.

Distribution: COL ECU PER VEN.

Type: Venezuela, Linden 685 (holotype, K-L; tracing of type, W-R-0050956, top).

Voucher: Colombia, Luer 7797 (AMES, SEL).

Masdevallia uncifera Rchb.f., Otia Bot. Hamburg. 1: 16. 1878.

Illustration: Luer 2001a, pl. 352.

Distribution: COL ECU.

Type: Colombia, Lehmann s.n. [lectotype (selected here), W-R-19674, illustr.].

Note: Luer (2001a) and other authors have indicated the type as being a specimen by Jameson kept at K, a tracing of which is in Reichenbach’s herbarium at W. However, the protologue suggests the type is a Lehmann collection, for which there is an illustration at W-R.

Masdevallia unguentum A.Doucette, Phytotaxa 48: 34. 2012.

Illustration: Doucette 2012, f. 1, f. 2.

Distribution: *COL.

Type: Colombia, Doucette 4000 (holotype, BH; isotype, PMA).

Masdevallia urceolaris Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 132. 1922.

Illustration: Luer 2001a, pl. 354.

Distribution: *COL.

Type: Colombia, Kalbreyer 545 [holotype, B, destroyed; lectotype, W-R-0052945 (Luer 2001)].

Masdevallia ustulata Luer, Phytologia 54(5): 384. 1983.

Illustration: Luer 2001a, pl. 355, pl. 356.

Distribution: COL ECU PER.

Type: Ecuador, Luer et al. 1660 (holotype, SEL).

Voucher: Colombia, Luer 14131 (MO)

Masdevallia valenciae Luer & R.Escobar, Orquideología 16(3): 36-39. 1986.

Illustration: Luer & Escobar 1986, p. 37; Luer 2002b, pl. 458.

Distribution: *COL.

Type: Colombia, Luer 11309 (holotype, JAUM; isotype, MO).

Masdevallia vargasii C.Schweinf., Amer. Orchid Soc. Bull. 19: 34, pl.. 1950. Alaticaulia vargasii (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer & Escobar 1982, p. 135; Luer 2000b, pl. 100-104; Vásquez & Ibisch 2000, pl. 98.

Distribution: BOL BRA COL ECU PER.

Type: Peru, Vargas 5528 (holotype, herb. Vargas; isotypes, AMES, USM).

Voucher: Colombia, Escobar 1984 (JAUM).

Masdevallia velella Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 87: 367a-368a, pl. 535. 2002. Byrsella velella (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2002b, pl. 535.

Distribution: *COL.

Type: Colombia, Rao 85 (holotype, MO).

Masdevallia velifera Rchb.f., Gard. Chron., n.s. 2: 98. 1874. Byrsella velifera (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 8. 2006.

Illustration: Luer 2000c, pl. 186; Williams & Williams 1894, p. 507.

Distribution: *COL.

Type: Colombia, Patin s.n. (holotype, W-R0038740; isotype, K-000077838).

Masdevallia ventricularia Rchb.f., Otia Bot. Hamburg. 1: 14. 1878.

Illustration: Luer 2002b, pl. 531.

Distribution: COL ECU.

Type: Ecuador, Lehmann 23 (holotype, W-R-50951).

Voucher: Colombia, Ortiz-Valdivieso 634 (HPUJ).

Masdevallia vieirana Luer & R.Escobar, Lindleyana 3(4): 211. 1989. Zahleria vieirana (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 17. 2006.

Illustration: Luer 2000c, pl. 252.

Distribution: *COL.

Type: Colombia, Luer 13055 (holotype, MO).

Masdevallia villegasii Königer, Arcula 9: 253. 1999.

Illustration: Ortiz 2000, 142 (p. 166).

Distribution: *COL.

Type: Colombia, Königer WK-96 (holotype, M; isotype, K, JAUM, Herb. Königer).

Masdevallia virgo-rosea N.Peláez, Buitr.-Delg. & Gary Mey., Orquideología 26(1): 46. 2009.

Illustration: Peláez et al. 2009, f. 8 (p. 40), f. 10.

Distribution: *COL.

Type: Colombia, Peláez NP0289 (holotype, JAUM-44182).

Note: Peláez NP0290 cited as isotype but is a paratype.

Masdevallia vittatula Luer & R.Escobar, Harvard Pap. Bot. 9: 9. 1996. Spilotantha vittatula (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 15. 2006.

Illustration: Luer 2003b, pl. 605A, pl. 605.

Distribution: COL ECU.

Type: Colombia, Luer 17983 (holotype, MO).

Masdevallia wuellneri P.Ortiz, Orquideología 24: 4. 2005.

Illustration: Ortiz 2005, pl. 2 (p. 11), ph. 2 (p. 15).

Distribution: *COL.

Type: Colombia, Ortiz 1182 (holotype, HPUJ).

Masdevallia wendlandiana Rchb.f., Gard. Chron., ser. 3 1: 174. 1887. Acinopetala wendlandiana (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 3. 2006.

Illustration: Luer 2001a, pl. 390; Cogniaux 1896, t. 78 (II); Vásquez & Ibisch 2000, f. 44.

Distribution: BOL BRA COL ECU PER VEN.

Type: Colombia, Wendland s.n. (holotype, W-R0050767).

Masdevallia wetzelii Luer & Sijm, Harvard Pap. Bot. 17(2): 358. 2012.

Illustration: Luer & Thoerle 2012b, f. 32.

Distribution: *COL.

Type: Colombia, Sijm 20111211 (holotype, MO).

Masdevallia wuerstlei Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 154. 1999. Alaticaulia wuerstlei (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 1999a, f. 12a; Luer 2000b, pl. 132.

Distribution: *COL.

Type: Colombia, Luer 8129 (holotype, SEL; isotype, MO).

Masdevallia xanthina Rchb.f., Gard. Chron. 1: 681. 1880. Masdevallia estradae var. xanthina (Rchb.f.) H.J.Veitch, Man. Orchid. Pl. 5: 42. 1889.

Illustration: Luer 2002b, pl. 463, pl. 464; Gutiérrez Morales & Sierra-Ariza 2020, f. 46.

Distribution: COL ECU.

Type: Colombia, Veitch 165 (holotype, W-R-50973).

Masdevallia xylina Rchb.f., Bot. Zeitung (Berlin) 31: 390. 1873. Alaticaulia xylina (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 6. 2006.

Illustration: Luer 2000b, pl. 112.

Distribution: *COL.

Type: Colombia, Roezl 717 [lectotype (selected here), W-R-0015007/W-0019499].

Note: A collection by Roezl at W-R is annotated by Reichenbach with the name M. xylina, but there is no direct indication from the protologue or on the sheet that suggests this specimen is the holotype. It is, however, the only material at W-R that bears the name and being consistent with the description is selected as lectotype.

Masdevallia zahlbruckneri Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 413. 1921. Zahleria zahlbruckneri (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 17. 2006, nom. inval. Zahleria zahlbruckneri (Kraenzl.) Luer, Fl. Mesoamer. 7(2): 545. 2023.

Illustration: Luer 2000b, pl. 253-256.

Distribution: BOL COL COS ECU PAN.

Type: Costa Rica, Endres 463 (holotype, W-R0015007/W-0019499).

Voucher: Colombia, Sarrazola et al. 1607 (HUA).

Masdevallia zapatae Luer & R.Escobar, Novon 1(4): 175. 1991. Petalodon zapatae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 11. 2006.

Illustration: Luer 2000b, pl. 190.

Distribution: *COL.

Type: Colombia, Escobar 4054 (holotype, JAUM; isotype, MO).

Myoxanthus Poepp. & Endl.

Myoxanthus affinis (Lindl.) Luer, Selbyana 7(1): 35. 1982. Humboldtia affinis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis affinis Lindl., Companion Bot. Mag. 2: 354. 1836.

Illustration: Luer 1991b, f. 390; Luer 1992, pl. 1; Ortiz & Uribe 2007 (CD Edition); Rojas-Alvarado & Karremans 2017, f. 1-4; Luer 1975f, f. 48; Karremans & Vieira-Uribe 2020, p. 189, p. 190; Rojas-Alvarado et al. 2021, f. 2e, f. 3a, f. 12a-c.

Distribution: BOL COL COS ECU PER.

Type: Peru, Mathews 1899 (holotype, K).

Voucher: Colombia, Luer 13413 (MO).

Myoxanthus ceratothallis (Rchb.f.) Luer, Selbyana 7(1): 36. 1982. Humboldtia ceratothallis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis ceratothallis Rchb.f., Bonplandia (Hannover) 2: 25. 1854.

Illustration: Luer 1992, pl. 4; Ortiz & Uribe 2007 (CD Edition); Karremans & Vieira-Uribe 2020, p. 198, p. 208; Rojas-Alvarado et al. 2021, f. 12e, f. 13.

Distribution: COL ECU PER VEN.

Type: Venezuela, Wagener sub Schiller s.n. [L611] (holotype, W-R-0024745/W-0117997; isotype, K-L).

Voucher: Colombia, Pennell 7581 (AMES, K).

Myoxanthus cimex (Luer & R.Escobar) Luer, Selbyana 7(1): 36. 1982. Pleurothallis cimex Luer & R.Escobar, Orquideología 14: 140. 1981.

Illustration: Luer & Escobar 1981a, p. 141; Luer 1992, pl. 6; Ortiz & Uribe 2007 (CD Edition); Karremans & Vieira-Uribe 2020, p. 196; Rojas-Alvarado et al. 2021, f. 12h-i.

Distribution: *COL.

Type: Colombia, Luer 2312 (holotype, JAUM; isotype, SEL).

Myoxanthus epibator Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 44: 30-32. 1992.

Illustration: Luer 1992, pl. 10.

Distribution: *COL.

Type: Colombia, Luer 14312 (holotype, MO).

Myoxanthus exasperatus (Lindl.) Luer, Selbyana 7(1): 36. 1982. Humboldtia exasperata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis exasperata Lindl., Fol. Orchid. ~Pleurothallis~ 15. 1859.

Illustration: Luer 1992, pl. 37; Karremans & Vieira-Uribe 2020, p. 203; Rojas-Alvarado et al. 2021, f. 2d, f. 2h, f. 3i, f. 14e.

Distribution: BRA COL COS ELS GUA PAN PER VEN.

Type: Venezuela, Fendler 1490 (holotype, K; isotypes, K, AMES, GH).

Voucher: Colombia, Renz 3613 (RENZ).

Myoxanthus frutex (Schltr.) Luer, Selbyana 7(1): 36. 1982. Pleurothallis frutex Schltr., Repert. Spec. Nov. Regni Veg. 10: 454. 1912.

Illustration: Luer 1992, pl. 14; Karremans & Vieira-Uribe 2020, p. 204; Rojas-Alvarado et al. 2021, f. 14h.

Distribution: BOL COL PER.

Type: Bolivia, Buchtien 2613 [holotype, B, destroyed; letotype, K-000583610; isolectotypes, AMES, L, US (Rojas-Alvarado et al. 2021)].

Voucher: Colombia, Schneider 571 (COL).

Myoxanthus hirsuticaulis (Ames & C.Schweinf.) Luer, Selbyana 7(1): 43. 1982. Pleurothallis hirsuticaulis Ames & C.Schweinf., Sched. Orch. 10: 29-30. 1930.

Illustration: Luer 1992, pl. 18; Karremans & Vieira-Uribe 2020, p. 197, p. 205, p. 206; Rojas-Alvarado et al. 2021, f. 2a, f. 3f, f. 4c-d, f. 5d, f. 6d, f. 7a, f. 11c-d, f. 15a.

Distribution: COL COS PAN PER.

Type: Costa Rica, Standley & Valerio 47052 (holotype, AMES).

Voucher: Colombia, Misas 226d (HPUJ).

Myoxanthus hystrix (Rchb.f.) Luer, Selbyana 7(1): 43. 1982. Dubois-reymondia hystrix (Rchb.f.) Brieger, Trab. Congr. Nac. Bot. 26: 43. 1977. Humboldtia hystrix (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis hystrix Rchb.f., Bonplandia (Hannover) 2: 26. 1854.

Illustration: Luer 1991b, f. 391; Luer 1992, pl. 19; Karremans & Vieira-Uribe 2020, p. 195; Rojas-Alvarado et al. 2021, f. 15b.

Distribution: COL VEN.

Type: Venezuela, Wagener s.n. [L612] (holotype, W-R-0025537/W-0117999; isotype, K-L).

Voucher: Colombia, Lehmann 3613, 3619 (G).

Myoxanthus melittanthus (Schltr.) Luer, Selbyana 7(1): 47. 1982. Pleurothallis melittantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 49. 1924.

Illustration: Luer 1991b, f. 392; Luer 1992, pl. 22; Karremans & Vieira-Uribe 2020, p. 195; Rojas-Alvarado et al. 2021, f. 15d; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Colombia, Hopp 136 (holotype, B, destroyed). Colombia, Luer et al. 3023 [neotype, SEL-021582; isoneotype, SEL-021538 (Rojas-Alvarado et al. 2021); type illustration Luer 1982, f. 8; Luer 1992: pl. 22].

Myoxanthus merae (Luer) Luer, Selbyana 7(1): 47. 1982. Pleurothallis merae Luer, Selbyana 3: 144, 1976.

Illustration: Luer 1992, pl. 23; Karremans & Vieira-Uribe 2020, p. 202; Rojas-Alvarado et al. 2021, f. 15e.

Distribution: COL ECU PER.

Type: Ecuador, Luer et al. 682 (holotype, SEL).

Voucher: Colombia, Arias 1020 (HUAZ).

Myoxanthus monophyllus Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 50. 1836. Pleurothallis myoxanthus Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 141. 1921. Pleurothallis poeppigii Lindl., Companion Bot. Mag. 2: 354. 1836, nom. superfl.

Illustration: Luer 1975f, f. 67; Luer 1992, pl. 24; Karremans & Vieira-Uribe 2020, p. 186; Rojas-Alvarado et al. 2021, f. 15f-h.

Distribution: COL ECU PER.

Type: Peru, Poeppig P.1741 (holotype, W; isotype, K).

Voucher: Colombia, André 2553 (K).

Myoxanthus montanus P.Ortiz, Orquideología 20: 320. 1997.

Illustration: Ortiz 1997, p. 317 (top right), p. 322 (top right); Rojas-Alvarado et al. 2021, f. 15i.

Distribution: *COL.

Type: Colombia, Ortiz 937 (holotype, HPUJ).

Myoxanthus ortizianus Kolan. & Szlach., Biodiv. Res. Conserv. 36(1): 8. 2014.

Illustration: Karremans & Vieira-Uribe 2020, p. 202; Rojas-Alvarado et al. 2021, f. 15l.

Distribution: *COL.

Type: Colombia, Medina 466 (holotype, HPUJ; photos, MEDEL; drawing, UGDA).

Myoxanthus parahybunensis (Barb.Rodr.) Luer, Selbyana 7(1): 47. 1982. Anathallis parahybunensis Barb.Rodr., Gen. Sp. Orchid. 2: 76. 1882.

Illustration: Cogniaux 1896, f. 2 (p. 111); Luer 1975f, f. 65; Luer 1992, pl. 12; Karremans & Vieira-Uribe 2020, p. 187, p. 188.

Distribution: BRA COL COS ECU PER VEN.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype, tab. 150, in Iconogr. Orchid. Bresil, vol. 3, at the Library of Rio de Janeiro Botanical Garden, cited as tab. 789 [(unpublished) in Barbosa Rodrigues (1882: 76); copied and reproduced in black and white in Cogniaux [1896: tab. 111, fig. II, 1896]; reproduced in colour in Sprunger et al. (1996, vol. 1: 208) (Rojas-Alvarado & Karremans 2020].

Voucher: Colombia, Ospina 81 (JAUM).

Myoxanthus parvilabius (C.Schweinf.) Luer, Selbyana 7(1): 49. 1982. Pleurothallis parvilabia C.Schweinf., Bot. Mus. Leafl. 20: 14. 1962.

Illustration: Luer 1992, pl. 28; Karremans & Vieira-Uribe 2020, p. 199; Rojas-Alvarado et al. 2021, f. 17d. Fig. 12H.

Distribution: COL ECU GUY SUR VEN.

Type: Guyana, Forestry Dept. G340 R.N. 7316 (holotype, NY).

Voucher: Colombia, Gutierrez & Moreno 64 (JBB).

Myoxanthus reymondii (H.Karst.) Luer, Selbyana 7(1): 49. 1982. Dubois-reymondia palpigera H.Karst., Bot. Zeitung (Berlin) 6: 397-398. 1848, nom. superfl. Dubois-reymondia reymondii (H.Karst.) Brieger, Trab. Congr. Nac. Bot. 26: 43. 1977. Duboisia reymondii H.Karst., Allg. Gartenzeitung 15(50): 394-395. 1847. Humboldtia reymondii (H.Karst.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis palpigera Karsten, Bot. Zeitung (Berlin) 6: 397-398. 1848, nom. illeg. Pleurothallis palpigera (H.Karst.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 6: 63. 1919, nom. illeg. Pleurothallis reymondii (H.Karst.) Rchb.f., Ann. Bot. Syst. 3: 520. 1852.

Illustration: Luer 1991b, f. 393; Luer 1992, pl. 32; Karremans & Vieira-Uribe 2020, p. 196; Rojas-Alvarado et al. 2021, f. 17k-l. Fig. 12I.

Distribution: COL ECU VEN.

Type: Venezuela, Wagener s.n. (holotype, W).

Voucher: Colombia, Luer 9056 (SEL).

Myoxanthus trachychlamys (Schltr.) Luer, Selbyana 7(1): 51. 1982. Pleurothallis trachychlamys Schltr., Repert. Spec. Nov. Regni Veg. Beih. 17: 23. 1922.

Illustration: Luer 1992, pl. 38; Misas Urreta 2005, p. 436. Fig. 12J; Karremans & Vieira-Uribe 2020, p. 185; Rojas-Alvarado et al. 2021, f. 1b, f. 2b, f. 2j, f. 3j, f. 4f, f. 5f, f. 6f, f. 10a, f. 18i.

Distribution: BRA COL COS ECU NIC PAN PER VEN.

Type: Panama, Powell 215 [holotype, B, destroyed; lectotype (Christenson 1991), AMES-23974; isolectotypes, AMES, MO; illustrat., AMES].

Voucher: Colombia, Schultes 8584 (IAN, AMES).

Myoxanthus uxorius (Luer) Luer, Selbyana 7(1): 51. 1982. Pleurothallis uxoria Luer, Selbyana 5(2): 186-187. 1979.

Illustration: Luer 1992, pl. 39; Karremans & Vieira-Uribe 2020, p. 198; Rojas-Alvarado et al. 2021, f. 18j.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 2533 (holotype, SEL).

Voucher: Colombia, Teague s.n. (MO).

Myoxanthus xiphion Luer, Selbyana 7(1): 51. 1982.

Illustration: Luer 1992, pl. 40; Karremans & Vieira-Uribe 2020, p. 199; Perdomo et al. 2020, f. 1, f. 2; Rojas-Alvarado et al. 2021, f. 18l.

Distribution: BOL COL ECU PER.

Type: Ecuador, Luer et al. 3276 (holotype, SEL).

Voucher: Colombia, Perdomo 180 (CUVC).

Octomeria R.Br.

Octomeria colombiana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 121. 1920.

Illustration: Luer 2010, f. 5a, 5b.

Distribution: BOL BRA COL ECU.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 43, No. 163)]. Colombia, Luer 17707 [neotype, MO (Luer 2010), here rejected and selected as epitype].

Note: We follow Pupulin et al. (2022) in recognizing Schlechter’s floral analyses reproduced by Mansfeld as original material and therefore reject Luer’s neotype in favor of a lectotype.

Octomeria costaricensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 111. 1923.

Illustration: Luer 2010, f. 10a, f. 10b.

Distribution: COL COS PAN.

Type: Costa Rica, Brade 1217 [holotype, B, destroyed; lectotype, HB (Luer 2010); illustration of type AMES-31270].

Voucher: Colombia, Cardenas 2149 (MO).

Octomeria crassilabiaPabst, Rodriguésia 31. 1956.

Illustration: Pabst 1956, pl. 2b.

Distribution: BRA COL.

Type: Brazil, Dusén 16077 (holotype, S).

Voucher: Colombia, Clavijo-Romero 1007 (COAH).

Octomeria exigua C.Schweinf., Bot. Mus. Leafl. 3: 86-87. 1935.

Illustration: Luer 2010, f. 15.

Distribution: COL GUY SUR VEN.

Type: Guyana, Thurn 115 B (holotype, K).

Voucher: Colombia, Gonzales & Cardenas 2693 (COL).

Octomeria filifolia C.Schweinf., Bot. Mus. Leafl. 19: 206. 1961.

Illustration: Luer 2010, f. 16.

Distribution: COL GUY VEN.

Type: Venezuela, Steyermark 75181 (holotype, AMES).

Voucher: Colombia, Duke 12254(6) (AMES).

Octomeria gemmula Carnevali & I.Ramírez, Ernstia 39: 13. 1986.

Illustration: Luer 2010, f. 17.

Distribution: COL VEN.

Type: Venezuela, Carnevali et al. 1715 (holotype, VEN).

Voucher: Colombia, Schultes & Cabrera 17516 (AMES).

Octomeria graminifolia (L.) R.Br., Hort. Kew. (ed. 2) 5: 211. 1813. Dendrobium graminifolium (L.) Willd., Sp. Pl. 4: 135. 1805. Epidendrum graminifolium L., Sp. Pl. ed. 2: 1353. 1763.

Illustration: Luer 2010, f. 19.

Distribution: BOL BRA COL COS ECU GUA MEX NIC PAN PER VEN.

Type: Martinique, Plumier s.n. (holotype, P); lectotype, Plumier in Burman, Pl. Amer.: 171, tab. 176, fig. 1 (1758) Convallaria? Caule articulato? (Garay & Sweet 1974).

Voucher: Colombia, Hirtz 4128 (MO).

Octomeria grandiflora Lindl., Edwards’s Bot. Reg. 28: Misc. 64-65. 1842.

Illustration: Luer 2010, f. 20a, 20b.

Distribution: BOL BRA COL ECU VEN.

Type: Brazil, Loddiges s.n. [lectotype, K; (Luer 2010)].

Voucher Colombia, Garcia-Barriga 14095 (NY).

Octomeria guentheriana Kraenzl., Repert. Spec. Nov. Regni Veg. 25: 19. 1928.

Illustration: Luer 2010, f. 21.

Distribution: BOL COL.

Type: Bolivia, Buchtien 562 (holotype, B, destroyed; lectotype, AMES; isolectotypes, HGB, US (Luer 2010)].

Voucher: Colombia, Idrobo & Schultes 857 (US).

Octomeria harantiana I.Bock, Orchidee (Hamburg) 35(2): 49. 1984.

Illustration: Luer 2010, f. 22.

Distribution: COL ECU PAN.

Type: Colombia, Bock, Bock & E. Harant 10/t. 10 (holotype, Herb. Irene Bock; isotypes, K, MO).

Octomeria heleneana Carnevali & Delascio, Ernstia 45: 12. 1987.

Illustration: Luer 2010, f. 23.

Distribution: COL VEN.

Type: Venezuela, Delacio 12354 (holotype, VEN).

Voucher: Colombia, Cardenas 49571 (COAH).

Octomeria longifolia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 58. 1924.

Illustration: Luer 2010, f. 28.

Distribution: BRA COL ECU GUY VEN.

Type: Colombia, Hopp 78, 86 (holotype, B, destroyed) Colombia, Schultes 19785 [neotype, AMES; (Luer 2010); isoneotype COL].

Octomeria mocoana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 59. 1924.

Illustration: Luer 2010, f. 32.

Distribution: BOL COL PER.

Type: Colombia, Hopp 76 (holotype, B, destroyed). Colombia, Palacios 1956 [neotype, W (Luer 2010)].

Octomeria napoleon Luer ex Gil & Karremans, Phytotaxa 406(5): 263. 2019. Octomeria napoleon Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 101-102. 2010, nom. inval.

Illustration: Luer 2010, f. 36.

Distribution: *COL.

Type: Colombia, Schultes & Cabrera 14519B (holotype, AMES-105032; isotypes, AMES-105033, AMES-barcode00287435).

Octomeria odontoglossoidesLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 102. 2010.

Illustration: Luer 2010, f. 37.

Distribution: *COL.

Type: Colombia, Luer 6729 (holotype, SEL).

Octomeria parvula C.Schweinf., Bot. Mus. Leafl. 3: 90. 1935.

Illustration: Luer 2010, f. 40.

Distribution: BRA COL GUY VEN.

Type: Guyana, Sandwith 234a (holotype, K).

Voucher: Colombia, Hoyos & Hernandez 941 (JAUM, MO)

Octomeria scirpoidea (Poepp. & Endl.) Rchb.f., Bot. Zeitung (Berlin) 10: 856. 1852. Aspegrenia scirpoidea Poepp. & Endl., Nov. Gen. Sp. Pl. 2: 12, t. 116. 1836.

Illustration: Cogniaux 1896, f. 2 (p. 133), Luer 2010, f. 48; Misas Urreta 2005, p. 448.

Distribution: BOL BRA COL COS ECU FRE GUY PER.

Type: Peru, Poeppig 1581 [lectotype (selected here), W-R; isolectotype, W-R-0054470!].

Voucher: Colombia, García-Barriga 14109 (AMES, US).

Note: There are two specimens with the number Poeppig 1581 at W-R. Both have the same ticket. One, which is numbered, has a ticket written by Reichenbach. It is probably the material he analyzed when studying this species. The other has (as of Aug 2017) no herbarium or barcode number, but it has an original ticket by Poeppig.

Octomeria seegeriana Kraenzl., Xenia Orchid. 3: 101, t. 257. 1892. Octomeria grandiflora var. seegeriana (Kraenzl.) Cogn., Fl. Bras. 3(4): 606. 1896. Octomeria robusta Barb.Rodr., Gen. Sp. Orchid. 2: 97. 1881, nom. illeg. Octomeria grandiflora var. robusta Cogn., Fl. Bras. 3(4): 606. 1896. Non Octomeria robusta Rchb.f. & Warm. [1881].

Illustration: Cogniaux 1896, f. 1 (p. 123), Luer 2010, f. 49.

Distribution: BOL BRA COL ECU.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype (Luer 2010) tab. 185, vol. 3, Barb. Rodr., Icon. Orchid. Brésil, orig. Illustr. At the Library, Rio de Janeiro Bot. Gard., cited as tab. 560 (unpub.) Barb.Rodr., 1881; Cogn. In Mart., Fl. Bras. 3(4), tab. 123, fig. I, 1896; reprod. in Sprunger et al., 1996, 1: 243, fig. B].

Voucher: Colomba, Warner 42 (AMES, MO).

Note: Octomeria seegeriana was proposed as a new name for the illegitimate O. robusta Barb.Rodr. Therefore, contrary to what Cogniaux (1896) and Luer (2010) suggests, they are homotypic. This also means that O. grandiflora var. robusta Cogn. and O. grandiflora var. seegirana are also homotypic.

Octomeria taracuana Schltr., Beih. Bot. Centralbl., Abt. 2 42(11): 93. 1925.

Illustration: Luer 2010, f. 54.

Distribution: BRA COL VEN.

Type: Brazil, Hübner 177 (holotype, B, destroyed). Brazil, Romero 3219A [neotype, TFAV (Luer 2010)].

Voucher: Colomba, Cárdenas-López 24493 (COAH).

Octomeria tridentata Lindl., Edwards’s Bot. Reg. 25: Misc. 35. 1839.

Illustration: Cogniaux 1896, f. 3 (p. 126), Luer 2010, f. 55a; Misas Urreta 2005, p. 450.

Distribution: BRA COL ECU FRE GUY NIC PER SUR VEN.

Type: Guyana, Loddiges s.n. (holotype, K).

Voucher: Colombia, Giraldo 360 (HUA, MO).

Octomeria yauaperyensis Barb.Rodr., Vellosia (ed. 2) 1: 122, t. 837. 1891.

Illustration: Cogniaux 1896, f. 3 (p. 132), Luer 2010, f.58.

Distribution: BRA COL COS ECU GUY HON PER VEN.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype (Luer 2010) fig. A, tab. 200, vol. 3, Barb.Rodr., Icon. Orchid. Brésil, orig. Illustr. At the Library, Rio de Janeiro Bot. Gard., reproduced as tab. 837 (unpub.) Barb.Rodr., 1891; reprod. in Sprunger et al., 1996, 1: 258].

Voucher: Colombia, Castaño-A. 1280 (COAH).

Octomeria zygoglossaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 117, f. 59. 2010.

Illustration: Luer 2010, f. 59.

Distribution: *COL.

Type: Colombia, Schultes & Cabrera 14521 (holotype, AMES; isotype, COL).

Ophidion Luer

Ophidion alphonsianum (Doucette ex L.E.Matthews) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila alphonsiana Doucette ex L.E.Matthews, OrchideenJ. 6(3): 8. 2018.

Illustration: Matthews 2018, p. 8 A-C, p. 9 (top); Vierling 2019, p. 146 (right), p. 147; Karremans & Vieira-Uribe 2020, p. 220.

Distribution: *COL.

Type: Colombia, Doucette 252 (holotype, WIS).

Ophidion barbilabium (Vierling) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila barbilabia Vierling, OrchideenJ. 26(4): 141. 2019.

Illustration: Vierling 2019, p. 141, p. 142; Karremans & Vieira-Uribe 2020, p. 221.

Distribution: *COL.

Type: Unknown country, Vierling s.n. (holotype, M).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 221.

Ophidion carrilloi P.Ortiz, Orquideología 24: 6. 2005. Phloeophila carrilloi (P.Ortiz) Pfahl, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 7 Jan 2014. 2014.

Illustration: Ortiz 2005, pl. 4 (13), ph. 4 (p. 15); Karremans & Vieira-Uribe 2020, p. 222.

Distribution: *COL.

Type: Colombia, Ortiz 1217 (holotype, HPUJ).

Ophidion compactum (Vierling) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila compacta Vierling, OrchideenJ. 26(4): 143. 2019.

Illustration: Vierling 2019, p. 143, p. 144; Karremans & Vieira-Uribe 2020, p. 215.

Distribution: *COL.

Type: Unknown, Vierling s.n. (holotype, M).

Voucher: Colombia, Gutierrez 2018 (JBB).

Ophidion cunabulum (Luer & R.Escobar) Luer & R.Escobar, Selbyana 7(1): 79. 1982. Cryptophoranthus cunabulus Luer & R.Escobar, Orquideología 14: 114. 1981. Phloeophila cunabulum (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16: 254. 2001.

Illustration: Luer & Escobar 1981a, p. 115; Luer 2004, f. 1 (p. 209); Luer 1991c, f. 449; Misas Urreta 2005, p. 464; Karremans & Vieira-Uribe 2020, p. 218. Fig. 12K.

Distribution: *COL.

Type: Colombia, Luer 2165 (holotype, SEL).

Ophidion dasyglossum (Luer & R.Escobar) Luer, Selbyana 7(1): 80. 1982. Cryptophoranthus dasyglossus Luer & R.Escobar, Orquideología 14: 116. 1981. Phloeophila dasyglossa (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 254. 2001.

Illustration: Luer & Escobar 1981a, p. 117; Luer 2004, f. 3 (p. 209); Luer 1991c, f. 451; Ortiz & Uribe 2007 (CD Edition); Karremans & VieiraUribe 2020, p. 222.

Distribution: *COL.

Type: Colombia, Escobar 1627 (holotype, JAUM).

Ophidion doucetteanum (Doucette ex L.E.Matthews) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila doucetteana Doucette ex L.E.Matthews , OrchideenJ. 6(3): 9. 2018.

Illustration: Matthews 2018, p. 9 (below), p. 10 F-J; Karremans & Vieira-Uribe 2020, p. 217.

Distribution: COL ECU?

Type: Ecuador?, Doucette 345 (holotype, WIS).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 217.

Ophidion erectilabrum Reina-Rodr., O.Pérez & Bogarín, Syst. Biodivers. 21(1, 2160504): 9.

Illustration: Reina-Rodríguez et al. 2023, f. 2, f. 3.

Distribution: *COL.

Type: Colombia, Reina-Rodríguez et al. 3044 (holotype, CUVC).

Ophidion nekidapono J.S.Moreno & Karremans, Pleurothallids Neotropical Jewels I: 220. 2020.

Illustration: Karremans & Vieira-Uribe 2020, p. 219.

Distribution: *COL.

Type: Colombia, Erazo & Moreno 702 (holotype, CAUP).

Ophidion nigerrimum (Vierling) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila nigerrima Vierling, OrchideenJ. 26(4): 149. 2019.

Illustration: Vierling 2019, p. 149 (right), p. 150; Karremans & Vieira-Uribe 2020, p. 216.

Distribution: COL.

Type: Unknown, Vierling s.n. (holotype, M).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 216.

Ophidion parvum (Vierling) Karremans & J.S.Moreno, Pleurothallids Neotropical Jewels I: 222. 2020. Phloeophila parva Vierling, OrchideenJ. 26(4): 153. 2019.

Illustration: Vierling 2019, p. 152, p. 153; Karremans & Vieira-Uribe 2020, p. 213, p. 214. Fig. 12L.

Distribution: *COL.

Type: Unknown, Vierling s.n. (holotype, M).

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 213, p. 214.

Ophidion pleurothallopsis (Kraenzl.) Luer, Selbyana 7(1): 80. 1982. Cryptophoranthus pleurothallopsis Kraenzl., Bull. Misc. Inform. Kew 1925(3): 115-116. 1925. Phloeophila pleurothallopsis (Kraenzl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 254. 2001.

Illustration: Luer 1991c, f. 450; Luer 2004, f. 4 (p. 209); Karremans & Vieira-Uribe 2020, p. 209, p. 210

Distribution: BOL COL ECU PAN VEN.

Type: Colombia, Lehmann 7272 (holotype, K).

Pabstiella Brieger & Senghas

Pabstiella aryter (Luer) F.Barros, Bradea 8(43): 296. 2002. Anthereon aryter (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 252. 2001. Pleurothallis aryter Luer, Selbyana 3(1-2): 54-56. 1976. Ronaldella aryter (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 195-197, f. 152a-b. 2006. Specklinia aryter (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004.

Illustration: Luer 2006, f. 152a, f. 152b, f. 152c.

Distribution: COL COS PER.

Type: Peru, Fuchs, Jr. s.n. (holotype, SEL).

Voucher: Colombia, Luer 13095 (MO).

Pabstiella syringodes (Luer) F.Barros, Bradea 8: 296. 2002. Anthereon syringodes (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 252. 2001. Gyalanthos syringodes (Luer) Szlach. & Marg., Polish Bot. J. 46: 117. 2001. Pleurothallis syringodes Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 176, f. 32a. 1999. Specklinia syringodes (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Luer 1999a, f. 32a.

Distribution: *COL.

Type: Colombia, Luer 19186 (holotype, MO).

Pabstiella tripterantha (Rchb.f.) F.Barros, Bradea 8(43): 296. 2002. Anthereon tripteranthus (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 252. 2001. Humboldtia tripterantha (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pabstiella tripterantha (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 140-141. 2006, nom. superfl. Pleurothallis tripterantha Rchb.f., Bonplandia (Hannover) 2: 24. 1854. Specklinia tripterantha (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Cogniaux 1896, f. 2; Luer 1975a, f. 20 (p. 97); Luer 2000a, f. 41, f. 42, f. 43 (p. 100); Luer 2006, f. 97, f. 98.

Distribution: BOL BRA COL COS ECU PAN PER VEN.

Type: Venezuela, Wagener 120 (holotype, W-R0053610).

Voucher: Colombia, Lehmann 134 (W).

Pabstiella yauaperyensis (Barb.Rodr.) F.Barros, Bradea 8: 296. 2002. Lepanthes yauaperyensis Barb. Rodr., Vellosia (ed. 2) 117. 1891. Specklinia yauaperyensis (Barb.Rodr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 2006, f. 99.

Distribution: BOL BRA COL PER VEN.

Type: Brazil, Barbosa Rodrigues s.n. [holotype, lost; lectotype (selected here), tab. 126, fig. B, in Iconogr. Orchid. Bresil, vol. 2, at the Library of Rio de Janeiro Botanical Garden; reproduced in color in Sprunger et al. (1996, vol. 1: 182)].

Voucher: Colombia, Navarro 1244 (COAH).

Pendusalpinx Karremans & Mel.Fernández

Pendusalpinx berlineri (Luer) Karremans & Mel. Fernández, Phytotaxa 340(2): 132. 2018. Pleurothallis berlineri Luer, Selbyana 3(1-2): 60. 1976. Trichosalpinx berlineri (Luer) Luer, Phytologia 54(5): 394. 1983.

Illustration: Luer 1997, f. 1; Ortiz & Uribe 2007 (CD Edition).

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Berliner s.n. (holotype, SEL; isotype, MO).

Voucher: Colombia, Portillo et al. 7767 (JAUM).

Pendusalpinx dependens (Luer) Karremans & Mel. Fernández, Phytotaxa 340(2): 132. 2018. Pleurothallis dependens Luer, Selbyana 3(1-2): 94, f. 150. 1976. Trichosalpinx dependens (Luer) Luer, Phytologia 54(5): 394. 1983. Fig. 13A.

Illustration: Luer 1997, f. 7, f. 8.

Distribution: BOL COL ECU FRE PER VEN.

Type: Ecuador, Brenner 22 (holotype, SEL).

Voucher: Colombia, Lehmann 7055 (AMES, K).

Pendusalpinx patula (Luer) Karremans & Mel. Fernández, Phytotaxa 340(2): 132. 2018. Trichosalpinx patula Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 31. 1997, nom. inval. Trichosalpinx patula Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 82. 1998.

Illustration: Luer 1997, f. 17; Ortiz & Uribe 2007 (CD Edition). Fig. 13B.

Distribution: BOL COL ECU VEN.

Type: Ecuador, Luer et al. 13395 (holotype, MO).

Voucher: Colombia, Schneider 353 (AMES, UGDA).

Platystele Schltr.

Platystele acicularis Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 14. 1990.

Illustration: Luer 1990, pl. 1.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 12693 (holotype, MO).

Voucher: Colombia, Sierra 5 (UPTC).

Platystele aculeata Luer, Phytologia 49(3): 197-198). 1981.

Illustration: Luer 1990, pl. 2.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6303 (holotype, SEL).

Voucher: Colombia, Luer et al. 17665 (MO).

Platystele acutilingua Kapuler & Hascall, Amer. Orchid Soc. Bull. 35: 544. 1966.

Illustration: Luer 1990, pl. 3.

Distribution: COL ECU.

Type: Colombia, Kapuler et al. 268 (holotype, COL).

Platystele alucitae Luer, Phytologia 46(6): 355. 1980.

Illustration: Ortiz 1981a, p. 209, p. 211; Luer 1990, pl. 5; Baquero & Zuchan 2017, f. 4; Baquero & Verkovitch 2019, f. 6a.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3943 (holotype, SEL).

Voucher: Colombia, Ortiz 969 (COL).

Platystele argentosa Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 22. 1990.

Illustration: Luer 1990, pl. 6; Ortiz 1977, p. 143 (top); Misas Urreta 2005, p. 490, p. 491.

Distribution: *COL.

Type: Colombia, Cogollo & Ramírez 3137 (holotype, JAUM; isotype, MO).

Platystele aurea Garay, Orquideología 8(3): 182. 1973.

Illustration: Luer 1986, pl. 37; Karremans et al. 2016, f. 8a.

Distribution: COL ECU PAN.

Type: Colombia, Escobar 886 (holotype, AMES).

Platystele baqueroiL. Jost & G. Iturralde, Lankesteriana 17(1): 55. 2017.

Illustration: Jost & Iturralde, f. 1-4. Fig. 13C.

Distribution: COL ECU.

Type: Ecuador, Baquero 3100 (holotype, QCNE).

Voucher: Colombia, Galindo et al. 1513 (CUVC).

Platystele beatricis P.Ortiz, Orquideología 22: 126. 2002.

Illustration: Ortiz 2002, p. 129.

Distribution: *COL.

Type: Colombia, Ortiz 1095 (holotype, COL).

Platystele calantha P.Ortiz, Orquideología 12: 138. 1977.

Illustration: Ortiz 1977, f. 2 (p. 139), p. 143 (bottom); Luer 1990, pl. 8.

Distribution: COL ECU.

Type: Colombia, Escobar 1414 (holotype, COL).

Platystele caudatisepala (C.Schweinf.) Garay, Orquideología 9(2): 119. 1974. Pleurothallis caudatisepala C.Schweinf., Bot. Mus. Leafl. 4(7): 113. 1937.

Illustration: Luer 1990, pl. 10; Bogarín & Karremans 2010, f. 4a; Karremans et al. 2016, f. 8b; Baquero & Verkovitch 2019, f. 6b.

Distribution: COL COS ECU GUA MEX PAN.

Type: Costa Rica, Brenes 1445 (holotype, AMES).

Voucher: Colombia, Escobar 4001 (MO).

Platystele colombiana S.Vieira-Uribe & Karremans, Spec. Orchid. 1(1). 2017.

Illustration: Vieira-Uribe & Karremans 2017b, pl. 12.

Distribution: *COL.

Type: Colombia, Vieira & Domínguez 0023 (holotype, JAUM).

Platystele consobrinaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 32. 1990.

Illustration: Luer 1990, pl. 12.

Distribution: *COL.

Type: Colombia, Luer 14765 (holotype, MO).

Platystele dasyglossa P.Ortiz, Orquideología 14(1): 18. 1979.

Illustration: Ortiz 1979a, p. 16 (bottom left), p. 19; Luer 1990, pl. 14.

Distribution: *COL.

Type: Colombia, Escobar 1857 (holotype, COL).

Platystele densiflora P.Ortiz, Orquideología 12(2-3): 140. 1977.

Illustration: Ortiz 1977, f. 3 (p. 141), p. 148; Luer 1991d, f. 482; Luer 1990, pl. 15.

Distribution: *COL.

Type: Colombia, Ortiz 909 (holotype, COL).

Platystele dewildei Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 121. 1994.

Illustration: Luer 1994, pl. 83.

Distribution: *COL.

Type: Colombia, De Wilde 3908 (holotype, MO).

Platystele examen-culicum Luer, Selbyana 3(1-2): 28. 1976.

Illustration: Luer 1990, pl. 20. Fig. 13C.

Distribution: COL ECU PER.

Type: Ecuador, Luer et al 574 (holotype, SEL).

Voucher: Colombia, Luer 2313 (SEL).

Platystele filamentosaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 46-47. 1990.

Illustration: Luer 1990, pl. 21.

Distribution: *COL.

Type: Colombia, Dodson et al. 18051 (holotype, MO).

Platystele gyroglossa Luer, Selbyana 3(3-4): 228. 1977.

Illustration: Ortiz 1981a, p. 217, p. 218; Luer 1990, pl. 23.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 208 (holotype, SEL).

Voucher: Colombia, Ortiz 971 (COL).

Platystele jesupiorum Luer, Phytologia 49(3): 199. 1981.

Illustration: Luer 1990, pl. 25. Fig. 13E.

Distribution: COL ECU.

Type: Ecuador, Luer 2235 (holotype, SEL).

Voucher: Colombia, Gutierrez & Moreno 73 (JBB).

Platystele johnstonii (Ames) Garay, Caldasia 10: 233. 1968. Pleurothallis johnstonii Ames, Orchidaceae 2: 271. 1908.

Illustration: Luer 1990, pl. 26.

Distribution: COL ECU VEN.

Type: Venezuela, Johnston 238 (holotype, AMES).

Voucher: Colombia, Hanbury-Tracy 553 (K).

Platystele lancilabris (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 102. 1923. Pleurothallis lancilabris (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. 12(317-321): 205. 1913. Stelis lancilabris Rchb.f., Beitr. Orchid.-K. C. Amer. 94, t. 8, 3-5. 1866.

Illustration: Luer 1990, pl. 28.

Distribution: COL COS.

Type: Costa Rica, Wendland 904 (holotype, W).

Voucher: Colombia, Cuatrecasas 12542 (COL).

Platystele lehmannii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 124. 1994.

Illustration: Luer 1994, pl. 85.

Distribution: *COL.

Type: Colombia, Lehmann 7090 (holotype, K).

Platystele londonoana Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 59, pl. 29. 1990.

Illustration: Luer 1990, pl. 29.

Distribution: *COL.

Type: Colombia, Luer 14301 (holotype, MO).

Platystele medinae Thoerle, Novon 26(2): 180. 2018.

Illustration: Thoerle 2018, f. 1-2.

Distribution: *COL.

Type: Colombia, Medina Trejo 920a (holotype, JAUM).

Platystele megaloglossa Luer & R.Escobar, Orquideología 16(3): 39-42. 1986.

Illustration: Luer & Escobar 1986, p. 41; Luer 1990, pl. 31.

Distribution: *COL.

Type: Colombia, Luer et al. 11374 (holotype, JAUM; isotype, MO).

Platystele microglossa P.Ortiz, Orquideología 14: 212. 1981.

Illustration: Ortiz 1981a, p. 213; Luer 1990, pl. 32.

Distribution: *COL.

Type: Colombia, Escobar 1748 (holotype, COL).

Platystele microscopica Luer, Phytologia 45: 356. 1980.

Illustration: Luer 1990, pl. 33.

Distribution: *COL.

Type: Colombia, Luer et al. 4608 (holotype, SEL).

Platystele misasiana P.Ortiz, Orquideología 14(3): 214. 1981.

Illustration: Ortiz 1981a, p. 214, p. 215; Luer 1990, pl. 36.

Distribution: *COL.

Type: Colombia, Ortiz 960 (holotype, COL).

Platystele misera (Lindl.) Garay, Bot. Mus. Leafl. 21(9): 251. 1967. Humboldtia misera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis misera Lindl., Fol. Orchid. ~Pleurothallis~ 36. n. 215. 1859.

Illustration: Luer 1990, pl. 37.

Distribution: COL PER.

Type: Peru, Mathews 3197 (holotype, K-L; isotypes, BM-0000074494, G).

Voucher: Colombia, Zarucchi et al. 7009 (HUA, MO).

Platystele myoxura Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 75. 1990.

Illustration: Luer 1990, pl. 39; Ortiz & Medina 2012, f. 10.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11711 (holotype, MO)

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz & Medina (2012).

Platystele orchestris P.Ortiz, Orquideología 14(1): 20. 1979.

Illustration: Ortiz 1979a, p. 16 (top left), p. 21; Luer 1990, pl. 40; Misas Urreta 2005, p. 494, p. 495.

Distribution: *COL.

Type: Colombia, Misas 10 (holotype, COL).

Platystele orectoglossa P.Ortiz, Orquideología 14: 22. 1979.

Illustration: Ortiz 1979a, p. 16 (top right), p. 23; Luer 1990, pl. 41.

Distribution: COL ECU.

Type: Colombia, Escobar 1053 (holotype, COL).

Platystele ortiziana Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 80-82. 1990.

Illustration: Luer 1991d, f. 483, Luer 1990, pl. 43.

Distribution: COL PAN.

Type: Colombia, Luer 13092 (holotype, MO).

Platystele ovalifolia (H.Focke) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 2: 268-269. 1961. Pleurothallis ovalifolia (H.Focke) Rchb.f., Ann. Bot. Syst. 6(2): 188. 1861. Stelis ovalifolia H.Focke, Tijdschr. WisNatuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 2: 202. 1849.

Illustration: Stenzel 2007, f. 15; Luer 1990, pl. 44.

Distribution: BRA COL CUB FRE GUY L.A. PAN SUR VEN.

Type: Guyana, Thurn 36 (holotype, K).

Voucher: Colombia, Ospina 80 (JAUM).

Platystele oxyglossa (Schltr.) Garay, Orquideología 9(2): 120. 1974. Pleurothallis lancilabris var. oxyglossa (Schltr.) C.Schweinf., Bot. Mus. Leafl. 6(10): 200. 1938. Pleurothallis oxyglossa Schltr., Repert. Spec. Nov. Regni Veg. 10(254-256): 354. 1912.

Illustration: Luer 1991d, f. 486, Luer 1990, pl. 46; Bogarín & Karremans 2010, f. 4b, c; Karremans et al. 2016, f. 4c.

Distribution: BOL BRA COL COS ECU ELS GUA MEX NIC PAN PER VEN.

Type: Guatemala, von Tuerckheim no. II 2422a (holotype, B, destroyed).

Voucher: Colombia, Luer 14175 (MO).

Platystele papillosa Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 127. 1994.

Illustration: Luer 1994, pl. 87.

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K).

Platystele pisifera (Lindl.) Luer, Selbyana 3(3,4): 232. 1977. Humboldtia pisifera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pisifera Lindl., Fol. Orchid. ~Pleurothallis~ 36. 1859.

Illustration: Luer & Escobar 1983, p. 15; Luer 1990, pl. 50.

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Luer et al. 7898 (SEL).

Platystele posadarum Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 96. 1990.

Illustration: Luer 1990, pl. 51.

Distribution: *COL.

Type: Colombia, Luer 7451 (holotype, SEL).

Platystele psix Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 44: 117. 1992.

Illustration: Luer 1992, pl. 80.

Distribution: COL ECU.

Type: Ecuador, Hirtz 4830 (holotype, MO).

Voucher: Colombia, Rincón et al. 1262 (TOLIspirit).

Platystele pubescens Luer, Phytologia 55(3): 200-201. 1984.

Illustration: Luer 1990, pl. 53; Baquero & Verkovitch 2019, f. 6d.

Distribution: COL ECU.

Type: Ecuador, Hirtz 1044 (holotype, SEL).

Voucher: Colombia, Luer 19900 (MO).

Platystele pyriformisLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 99. 1990.

Illustration: Luer 1990, pl. 54.

Distribution: *COL.

Type: Colombia, Luer 676 (holotype, SEL).

Platystele rhinocera Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 108-110. 1990.

Illustration: Luer 1990, pl. 60.

Distribution: *COL.

Type: Colombia, Luer 12493 (holotype, MO).

Platystele riograndensis Reina-Rodr. & Karremans, Phytotaxa 375(3): 236. 2018.

Illustration: Reina-Rodríguez & Karremans 2018, f. 1-3.

Distribution: *COL.

Type: Colombia, Reina-Rodríguez et al. 1603 (holotype, CUVC-spirit).

Note: Originally published as P. riograndense.

Platystele risaraldae Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 130. 1994.

Illustration: Luer 1994, pl. 88.

Distribution: *COL.

Type: Colombia, De Wilde 326 (holotype, MO).

Platystele schmidtchenii Schltr., Repert. Spec. Nov. Regni Veg. 27: 166. 1924.

Illustration: Luer 1990, pl. 61.

Distribution: BOL COL ECU.

Type: Colombia, Schmidtchen s.n. (holotype, B, destroyed).

Platystele schneideri P.Ortiz, Orquideología 14(1): 24. 1979.

Illustration: Ortiz 1979a, p. 16 (bottom right), p. 25; Luer 1990, pl. 62; Luer 1991d, f. 485.

Distribution: *COL.

Type: Colombia, Ortiz 48 (holotype, COL).

Platystele stenostachya (Rchb.f.) Garay, Caldasia 8(40): 520-521. 1962. Humboldtia stenostachya (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis stenostachya Rchb.f., Linnaea 18: 399. 1844 [1845].

Illustration: Luer 1990, pl. 65; Misas Urreta 2005, p. 504, p. 505.

Distribution: BEL COL COS ECU ELS GUA HON MEX NIC PAN SUR VEN.

Type: Mexico, Leibold 20 [L547] [lectotype (selected here), W-R-0021894; isolectotype, HAL-0076897]. Mexico, Leibold 32 (syntype, K-L-000364141).

Voucher: Colombia, Cuatrecasas 15642 (AMES).

Platystele stonyxLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 38: 122. 1990.

Illustration: Luer 1990, pl. 67.

Distribution: *COL.

Type: Colombia, Luer et al. 12519 (holotype, MO).

Platystele sylvestrei Karremans & Bogarín, Lankesteriana 13(3): 268-271. 2014.

Illustration: Fernández et al. 2014, f. 6, f. 7.

Distribution: COL COS PAN.

Type: Costa Rica, Bogarín et al. 8240 (holotype, JBL-spirit).

Voucher: Colombia, Vieira 040 (JAUM).

Platystele tica Karremans & Bogarín, Lankesteriana 13(3): 271-274. 2014.

Illustration: Fernández et al. 2014, f. 8, f. 9, f. 14g- h; Rincón-González et al. 2020a, f. 3.

Distribution: COL COS.

Type: Costa Rica, Karremans et al. 5315 (holotype, JBL-spirit; isotype, JBL-spirit).

Voucher: Colombia, Vieira 047 (JAUM).

Platystele umbellata P.Ortiz, Orquideología 12(2-3): 136. 1977.

Illustration: Ortiz 1977, f. 1 (p. 137), p. 143 (middle); Luer 1990, pl. 70; Misas Urreta 2005, p. 506, p. 507; Karremans et al. 2016, f. 6b.

Distribution: *COL.

Type: Colombia, Ortiz 916 (holotype, COL).

Pleurothallis R.Br.

Pleurothallis abortiva Luer, Phytologia 47(2): 72-73. 1980. Tigivesta abortiva (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 121. 2007. Vestigium abortivum (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 309. 2005.

Illustration: Wilson et al. 2018b, f. 2a; Luer 1998b, f.1.

Distribution: *COL.

Type: Colombia, Luer et al. 4547 (holotype, SEL).

Pleurothallis acutilabia Luer, Orquideología 21(2): 116-118. 1999. Acronia acutilabia (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 83. 2005. Zosterophyllanthos acutilabius (Luer) Szlach. & Kulak, Richardiana 6: 186. 2006.

Illustration: Luer 1999b, p. 117; Luer 2005, f. 2 (p. 208).

Distribution: *COL.

Type: Colombia, Ramos et al. 1225 (holotype, CUVC; isotype, MO).

Pleurothallis adonis Luer, Selbyana 3(1-2): 40. 1976. Acronia adonis (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 85. 2005. Zosterophyllanthos adonis (Luer) Szlach. & Kulak, Richardiana 6: 186. 2006.

Illustration: Luer 2005, f. 5a, f. 5b, f. 5c (p. 209).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 454 (holotype, SEL).

Voucher: Colombia, Lehmann 3897 (K).

Pleurothallis aggeris Luer, Lindleyana 11(3): 143. 1996. Rhynchopera aggeris (Luer) Szlach. & Marg., Polish Bot. J. 46: 117. 2001.

Illustration: Luer 1998a, f. 1 (p. 92); Luer 1996c, f. 3.

Distribution: COL ECU PER.

Type: Ecuador, Luer et al. 1632 (holotype, SEL).

Voucher: Colombia, Ospina 64 (JAUM).

Pleurothallis aguirrei Luer & R.Escobar, Orquideología 22(3): 255-257, 280-281. 2003. Acronia aguirrei (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 85. 2005. Zosterophyllanthos aguirrei (Luer & R.Escobar) Szlach. & Kulak, Richardiana 6: 132. 2006.

Illustration: Luer 2003c, p. 257; Luer 2005, f. 6 (p. 210).

Distribution: *COL.

Type: Colombia, Escobar 8310 (holotype, MO).

Pleurothallis altimonile Luer & R.Escobar, Orquideología 14(2): 120. 1981. Acronia altimonile (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 86. 2005. Zosterophyllanthos altimonile (Luer & R.Escobar) Szlach. & Marg., Polish Bot. J. 46: 118. 2001.

Illustration: Luer & Escobar 1981a, p. 121; Luer 2005, f. 8 (p. 210).

Distribution: *COL.

Type: Colombia, Luer 2339 (holotype, JAUM; isotype, SEL).

Pleurothallis alvaroi Luer & R.Escobar, Orquideología 14(2): 122. 1981. Acronia alvaroi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1981a, p. 123; Luer 1998b, f. 3.

Distribution: *COL.

Type: Colombia, Luer 2088 (holotype, SEL; isotype, JAUM).

Pleurothallis ambyx Luer & R.Escobar, Orquideología 14(2): 126. 1981. Acronia ambyx (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 87. 2005. Zosterophyllanthos ambyx (Luer & R.Escobar) Szlach. & Marg., Polish Bot. J. 46: 118. 2001.

Illustration: Luer & Escobar 1981a, p. 127; Luer 2005, f. 9 (p. 210).

Distribution: COL ECU.

Type: Colombia, Luer 2045 (holotype, SEL).

Pleurothallis amphigya Luer & R.Escobar, Orquideología 14(2): 128. 1981. Acronia amphigya (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1981a, p. 129; Luer 1998b, f. 26, f. 27.

Distribution: *COL.

Type: Colombia, Luer 2046 (holotype, SEL).

Pleurothallis amplectens Luer, Phytologia 47(2): 73. 1980.

Illustration: Luer 1999a, f. 48.

Distribution: COL ECU.

Type: Ecuador, Luer & Luer 5458 (holotype, SEL).

Voucher: Colombia, André 2972 (K).

Pleurothallis anceps Luer, Selbyana 5(2): 159. 1979. Ancipitia anceps (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1989, pl. 1.

Distribution: COL ECU.

Type: Ecuador, Madison et al. s.n. (holotype, SEL).

Voucher: Colombia, Luer 2056 (SEL).

Pleurothallis andreae Mark Wilson, B.T.Larsen & J.Portilla, Lankesteriana 17(2): 177. 2017.

Illustration: Luer 1989 f, 6; Wilson et al. 2017c, f. 3, 4, 6-8, 10, 11, 15-20.

Distribution: *COL.

Type: Colombia, Wilson & Niessen PL0962 (holotype, COCO).

Pleurothallis antennifera Lindl., Fol. Orchid. ~Pleurothallis~ 30, n. 178. 1859. Humboldtia antennifera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 2; Luer 1975f, f. 49.

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Jameson 84 (holotype, K-L; isotypes, K-L, W-R-53840).

Voucher: Colombia, Triana 124 (K, W).

Pleurothallis anthrax Luer & R.Escobar, Orquideología 14(2): 130. 1981. Ancipitia anthrax (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer & Escobar 1981a, p. 131; Luer 1989, pl. 2.

Distribution: *COL.

Type: Colombia, Luer et al. 3724 (holotype, SEL).

Pleurothallis applanata Luer & Dalström, Lindleyana 11(3): 147. 1996. Acronia applanata (Luer & Dalström) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 89. 2005. Zosterophyllanthos applanatus (Luer & Dalström) Szlach. & Kulak, Richardiana 6: 186. 2006.

Illustration: Luer 2005, f. 13 (p. 211); Luer 1996c, f. 8.

Distribution: COL ECU PER.

Type: Ecuador, Dalström & Höijer 1060 (holotype, MO).

Voucher: Colombia, Otero 261 (MO).

Pleurothallis archidonae Lindl., Fol. Orchid. ~Pleurothallis~ 15, no. 74. 1859. Humboldtia archidonae (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 49.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K-L000820778).

Voucher: Colombia, Schneider 3252 (COL).

Pleurothallis ascera Luer & R.Escobar, Orquideología 20: 36. 1996. Acronia ascera (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 90. 2005.

Illustration: Luer & Escobar 1996, p. 37, p. 58 (top, right); Luer 2005, f. 15 (p. 212).

Distribution: *COL.

Type: Colombia, Escobar 5052 (holotype, JAUM; isotype, MO).

Pleurothallis baccata (Luer) J.M.H.Shaw, Orchid Rev. 122(1308): 76. 2014. Acronia baccata Luer, Selbyana 30(1): 2. 2009.

Illustration: Luer 2009b, f. 3.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 8871 (holotype, CUVC; isotype, MO).

Pleurothallis baezensis (Luer & Hirtz) J.M.H.Shaw, Orchid Rev. 122: 76. 2014. Acronia baezensis Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard 103: 91. 2005.

Illustration: Luer 2005, f. 18 (p. 213).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3188 (holotype, CUVC; isotype, MO).

Voucher: Colombia, Valencia & Rincón 187 (TOLI).

Pleurothallis baudoensisLuer & R.Escobar, Orquideología 21(1): 73-75, pl. 1. 1998. Acronia baudoensis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 92. 2005.

Illustration: Luer & Escobar 1998, p. 74; Wilson et al. 2018b, f. 1b; Luer 2005, f. 20 (p. 213); Misas Urreta 2005, p. 523.

Distribution: *COL.

Type: Colombia, de Wilde 3915 (holotype, MO).

Pleurothallis belocardia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 161. 1924. Acronia belocardia (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 93. 2005. Zosterophyllanthos belocardia (Schltr.) Szlach. & Kulak, Richardiana 6: 186. 2006.

Distribution: *COL.

Type: Colombia, Hopf s.n. (holotype, B, destroyed).

Note: An obscure name, no type material seems to be extant.

Pleurothallis bicochlearis Luer, Orquideología 21(2): 118-120, f. 2. 1999.

Illustration: Luer 1999b, p. 119; Luer 1999a, f. 4.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4282 (holotype, CUVC; isotype, MO).

Pleurothallis bicornis Lindl., Fol. Orchid. ~Pleurothallis~ 7, no. 29. 1859. Humboldtia bicornis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 51; Wilson et al. 2017e, f. 2, 3, 5, 8.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K-L001097657).

Voucher: Colombia, Luer et al. 10266 (MO).

Pleurothallis bicruris Lindl., Fol. Orchid. ~Pleurothallis~ 8. 1859. Humboldtia bicruris (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 5.

Distribution: COL ECU.

Type: Ecuador, Jameson 72 (holotype, K-L; isotype, W-R-21853).

Voucher: Colombia, Triana 603 (P).

Pleurothallis bivalvis Lindl., Orchid. Linden. 2. 1846. Acronia bivalvis (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 93-96, f. 22. 2005. Humboldtia bivalvis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 2003c, p. 266; Luer 2005, f. 22a, f. 22b, f. 22c, f. 22d (p. 214), f. 22e, f. 22f, f. 22g, f. 22h (p. 215).

Distribution: BOL COL COS ECU FRE GUA GUY L.A. MEX PAN PER VEN.

Type: Venezuela, Linden 1480 [holotype, K-L000820780; superfl. selected as lectotype by Kolanowska (2014)].

Voucher: Colombia, Roezl s.n. (W).

Pleurothallis bovilinguaLuer & R.Escobar, Orquideología 21(1): 75-77, pl. 2, photo 2. 1998. Acronia bovilingua (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 97. 2005.

Illustration: Luer & Escobar 1998, p. 75, p. 76; Luer 2005, f. 25 (p. 216); Jiménez et al. 2018, f. 4b, f. 5d.

Distribution: *COL.

Type: Colombia, Escobar 986 (holotype, AMES).

Pleurothallis brachiata Luer, Selbyana 1(3): 230. 1975.

Illustration: Luer 1999a, f. 6; Luer 1975f, f. 50; Misas Urreta 2005, p. 525.

Distribution: COL ECU.

Type: Ecuador, Luer & Kent 337 (holotype, SEL).

Voucher: Colombia, John 20860 (AMES, COL).

Pleurothallis bulbosa (Luer & R.Escobar) Pfahl, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 3a: 1. 2012. Acronia bulbosa Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 98, f. 28. 2005.

Illustration: Luer 2005, f. 28 (p. 217).

Distribution: *COL.

Type: Colombia, Luer et al. 9043 (holotype, SEL).

Pleurothallis calamifolia Luer & R.Escobar, Orquideología 20: 40. 1996. Acronia calamifolia (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 99. 2005.

Illustration: Luer & Escobar 1996, p. 41; Wilson et al. 2018b, f. 1c; Luer 2005, f. 29 (p. 217).

Distribution: *COL.

Type: Colombia, Luer et al. 10325 (holotype, MO).

Pleurothallis calceolaris Rchb.f., Linnaea 41: 15. 1877. Acronia calceolaris (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 99. 2005.

Illustration: Luer & Escobar 1984b, p. 161; Luer 2005, f. 30 (p. 217); Wilson et al. 2018b, f. 1d.

Distribution: *COL.

Type: Colombia, Roezl s.n. (holotype, W-R0025681).

Pleurothallis calogramma Luer, Orquideología 22(3): 256-259, 281-282. 2003. Acronia calogramma (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 100. 2005.

Illustration: Luer 2003c, p. 258; Luer 2005, f. 32 (p. 218).

Distribution: *COL.

Type: Colombia, Luer 2115 (holotype, SEL).

Pleurothallis calolalaxLuer & R.Escobar, Orquideología 21(1): 78-80. 1998. Acronia calolalax (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 101. 2005.

Illustration: Luer & Escobar 1998, p. 79, f. 3 (p. 97); Luer 2005, f. 33 (p. 218).

Distribution: *COL.

Type: Colombia, Luer et al. 17693 (holotype, JAUM; isotype, MO).

Pleurothallis calvariola Luer & R.Escobar, Orquideología 14(2): 136. 1981.

Illustration: Luer & Escobar 1981a, p. 137; Luer 1999a, f. 54; Luer 1996c, f. 55.

Distribution: COL ECU.

Type: Colombia, Luer et al. 3056 (holotype, SEL).

Pleurothallis canaligera Rchb.f., Linnaea 41: 15, f. 51. 1877. Acronia canaligera (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 101. 2005. Zosterophyllanthos canaliger (Rchb.f.) Szlach. & Kulak, Richardiana 6(4): 187. 2006.

Illustration: Luer 2005, f. 34 (p. 218); Luer 1975f, f. 51.

Distribution: COL ECU PER.

Type: Colombia, Roezl s.n. (holotype, W-R0025679).

Pleurothallis canidentisLuer & R.Escobar, Orquideología 21: 80-82. 1998. Acronia canidentis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 102. 2005.

Illustration: Luer & Escobar 1998, p. 81, f. 4 (p. 97); Luer 2005, f. 35 (p. 219).

Distribution: *COL.

Type: Colombia, Escobar 5228 (holotype, JAUM; isotype, MO).

Pleurothallis capitonis Luer & R.Escobar, Orquideología 20: 44. 1996.

Illustration: Luer & Escobar 1996, p. 43; Luer 1999a, f. 55.

Distribution: COL ECU.

Type: Colombia, Luer et al. 10110 (holotype, MO).

Pleurothallis caprina Luer & R.Escobar, Orquideología 14(2): 138. 1981. Ancipitia caprina (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer & Escobar 1981a, p. 139; Luer 1989, pl. 4; Luer 1991e, f. 491.

Distribution: *COL.

Type: Colombia, Escobar 226 (holotype, JAUM).

Pleurothallis cardiophylax Rchb.f., Flora 71: 154. 1888. Acronia cardiophylax (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 103. 2005. Zosterophyllanthos cardiophylax (Rchb.f.) Szlach. & Kulak, Richardiana 6: 187. 2006.

Illustration: Luer 2005, f. 37 (p. 219).

Distribution: COL ECU PER.

Type: Colombia, Low s.n. (holotype, W-R0025682).

Pleurothallis cardiostola Rchb.f., Bonplandia (Hannover) 2: 26. 1854. Acronia cardiostola (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 104. 2005. Humboldtia cardiostola (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Zosterophyllanthos cardiostola (Rchb.f.) Szlach. & Kulak, Richardiana 6: 187. 2006.

Illustration: Luer 2005, f. 38a (p. 219), f. 38b (p. 220); ; Wilson et al. 2016, f. 1b, f. 2b; Jiménez et al. 2021a, f.3b, f. 4, f. 5; Schuiteman 2018, f. 6 (p. 57); Doucette et al. 2016b, f. 5, f. 11g.

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Wagener 49 (holotype, W).

Voucher: Colombia, Lehmann s.n. (K).

Pleurothallis cardiothallis Rchb.f., Bot. Zeitung (Berlin) 15(10): 158-159. 1857. Acronia cardiothallis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 106, f. 39. 2005. Humboldtia cardiothallis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Zosterophyllanthos cardiothallis (Rchb.f.) Szlach. & Kulak, Richardiana 6(4): 187. 2006.

Illustration: Luer 2005, f. 39 (p. 220), Pupulin et al. 2017a, fig. 1-5, 22, 23; Pupulin et al. 2021, f. 2(10), f. 5, f. 21e-f, f. 26; Sierra-Ariza et al. 2022, f. 5.

Distribution: COL COS ECU ELS GUA MEX NIC PAN.

Type: Unknown, Schiller s.n. (holotype, W; illustration, AMES).

Voucher: Colombia, Luer & Escobar 6592 (SEL).

Pleurothallis carnosa Braid, Kew Bull. 1924: 200. 1924. Acronia carnosa (Braid) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 107. 2005. Zosterophyllanthos carnosus (Braid) Szlach. & Kulak, Richardiana 6: 187. 2006.

Illustration: Luer 2005, f. 41 (p. 220).

Distribution: *COL.

Type: Unknown (holotype, K).

Voucher: Colombia, Albert et al. s.n. (HUA, MO).

Pleurothallis cassidataLuer & R.Escobar, Orquideología 21: 82-84, t. 5. 1998. Acronia cassidata (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 108. 2005.

Illustration: Luer & Escobar 1998, p. 83, f. 5 (p. 97); Luer 2005, f. 43 (p. 221).

Distribution: *COL.

Type: Colombia, Escobar 8289 (holotype, JAUM; isotype, MO).

Pleurothallis caucensis Mark Wilson, Lankesteriana 17(2): 122, 124. 2017.

Illustration: Wilson et al. 2017e, f. 1, 6, 10-13.

Distribution: *COL.

Type: Colombia, Wilson et al. PL0993 (holotype, JAUM).

Pleurothallis cauda-hirundinis Luer & J. Portilla, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 107. 2002. Acronia cauda-hirundinis (Luer & J. Portilla) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer 2002a, f. 17 (p. 119); Buitrago-Delgado et al. 2005, f. 1 (p. 51), f. 2 (p. 52), ph. 1 (p. 52).

Distribution: COL ECU.

Type: Ecuador, Luer 19675 (holotype, MO).

Voucher: Colombia, Valencia-D. 474 (COL).

Pleurothallis celsia Gal.-Tar. & J.S.Moreno, ined.

Illustration: Moreno et al. 2023, f. 1-3.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona 1463 (holotype, CUVC).

Pleurothallis cernua Luer, Selbyana 1(3): 234. 1975.

Illustration: Luer 1975f, f. 52; Luer 1998b, f.52.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 426 (holotype, SEL).

Voucher: Colombia, Silverstone-Sopkin et al. 1678 (holotype, CUVC; isotype, MO).

Pleurothallis chicalensis M.M. Jiménez & Baquero, Lankesteriana 18(2): 104. 2018.

Illustration: Jiménez et al. 2018, f. 1, f. 2a, f. 2b, f. 5a.

Distribution: COL ECU.

Type: Ecuador, LB 3033 (holotype, QCNE).

Voucher: No Colombian voucher available but see Jiménez et al. (2018).

Pleurothallis chuscalica Luer, Orquideología 21(2): 120-122. 1999.

Illustration: Luer 1999b, p. 121; Luer 1999a, f. 56.

Distribution: *COL.

Type: Colombia, Lehmann 7134 (holotype, K).

Pleurothallis circinata Luer, Orquideología 20: 206. 1996.

Illustration: Luer 1996f, 207; Luer 1998b, f. 54.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 3951 (holotype, CUVC; isotype, MO).

Pleurothallis colossus Kraenzl. ex Kerch., Livre Orchid. 376. 1894.

Illustration: Luer 1999a, f. 57.

Distribution: COL COS ECU PAN.

Type: Colombia, Lehmann 6929 [lectotype, LE (Christenson 1995); isolectotypes, AMES (superfl. selected as lectotype by Kolanowska 2014), K; photo of type, MO].

Pleurothallis constrictaLuer & R.Escobar, Orquideología 21: 84-86, t. 6. 1998. Acronia constricta (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 113. 2005.

Illustration: Luer & Escobar 1998, p. 85; Luer 2005, f. 52 (p. 223).

Distribution: *COL.

Type: Colombia, Escobar 5297 (holotype, JAUM; isotype, MO).

Pleurothallis cordata (Ruiz & Pav.) Lindl., Gen. Sp. Orchid. Pl. 5. 1830. Acronia cordata (Ruiz & Pav.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 114. 2005. Humboldtia cordata Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 234. 1798. Stelis cordata (Ruiz & Pav.) Willd., Sp. Pl. 4: 140. 1805.

Illustration: Luer 2005, f. 54a, f. 54b, f. 54c (p. 224), f. 54d, f. 55a, f. 55b, f.56 (p. 225); Pupulin 2013, f. 36b.

Distribution: BOL COL ECU PER.

Type: Peru, Ruiz & Pavón s.n. (holotype, M; isotype, BM-000082395).

Voucher: Colombia, Cogollo 2687 (JAUM, MO).

Pleurothallis cordifolia Rchb.f. & Wagener, Bonplandia (Hannover) 2: 26. 1854. Acronia cordifolia (Rchb.f. & Wagener) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 117, f. 57. 2005. Humboldtia cordifolia (Rchb.f. & Wagener) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Zosterophyllanthos cordifolius Szlach. & Kulak, Richardiana 6: 188. 2006.

Illustration: Luer 2005, f. 57 (p. 226).

Distribution: COL ECU PAN VEN.

Type: Venezuela, Wagener 42 [lectotype (selected here), W-R-0025499, type illustr. top left].

Voucher: Colombia, Hutchison 3133 (F, K, MO, M, MICH, NY, P, UC, US).

Note: The type sheet at W-R is mistakenly annotated as “Holotype” and “Wagener 147”. It consists of an illustration, a flower sketch and an empty envelope. The drawings, Wagener 42, agree with the information provided by the protologue. The top left illustration is selected as lectotype.

Pleurothallis coriacardia Rchb.f., Bonplandia (Hannover) 2: 26. 1854. Acronia coriacardia (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 117. 2005. Humboldtia coriacardia (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Zosterophyllanthos coriacardia (Rchb.f.) Szlach. & Kulak, Richardiana 6: 188. 2006.

Illustration: Luer 2005, f. 58 (p. 226).

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Hartweg, s.n. (holotype, K).

Pleurothallis cosmetronLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 44: 120, pl. 7. 1992. Colombiana cosmetron (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 249. 2006.

Illustration: Luer 1992, pl. 7 (p. 121); Wilson et al. 2017d, f. 8a.

Distribution: *COL.

Type: Colombia, Pennell 4352 (holotype, NY).

Pleurothallis cotyligera Luer & R.Escobar, Orquideología 20: 46. 1996.

Illustration: Luer & Escobar 1996, p. 48; Luer 1999a, f. 59.

Distribution: *COL.

Type: Colombia, Luer 16765 (holotype, MO).

Pleurothallis crocodiliceps Rchb.f., Bonplandia (Hannover) 3: 72. 1855. Ancipitia crocodiliceps (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Humboldtia crocodiliceps (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1975a, f. 6 (p. 69); Wilson et al. 2017c, f. 1, 2.

Distribution: COL ECU.

Type: Colombia, Wagener s.n. [lectotype (selected here), W-R-0024689/W-0073521, stem and flower].

Note: There is no clear indication as to the holotype of this controversial taxon. A specimen at W-R which has been regarded as holotype is here selected as lectotype given it conforms with the protologue. The sheet consists of a rough sketch and a single ramicaul, leaf and flower. The lip has been severed and is unfortunately glued back to front.

Pleurothallis cunabularis Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 42. 1998.

Illustration: Luer 1998b, f. 56.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4909 (holotype, CUVC; isotype, MO).

Pleurothallis cyanea Luer & R.Escobar, Orquideología 16(2): 157. 1984. Acronia cyanea (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 123. 2005.

Illustration: Luer & Escobar 1984b, p. 158, p. 159; Luer 2005, f. 67 (p. 228).

Distribution: *COL.

Type: Colombia, Luer 6730 (holotype, SEL).

Pleurothallis cymbisepala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 45. 1924.

Illustration: Luer 1999a, f. 7.

Distribution: COL ECU.

Type: Colombia, Hopp 175 (holotype, B, destroyed). Colombia, Luer et al. 3078 (neotype, K; isoneotype, SEL).

Pleurothallis deflexa Luer, Selbyana 1(2): 178. 1975. Acronia deflexa (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 123. 2005. Zosterophyllanthos deflexus (Luer) Szlach. & Kulak, Richardiana 6: 189. 2006.

Illustration: Luer 2005, f. 68 (p. 228); Luer 1975b, f. 25.

Distribution: COL ECU VEN.

Type: Ecuador, Luer & Kent 225 (holotype, SEL).

Voucher: Colombia, Luer 16870 (MO).

Pleurothallis depressa Luer & R.Escobar, Orquideología 20: 50. 1996. Acronia depressa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 125. 2005.

Illustration: Luer & Escobar 1996, p. 53, p. 58 (bottom right); Wilson et al. 2018b, f. 1g; Luer 2005, f. 70 (p. 229).

Distribution: *COL.

Type: Colombia, Luer 17569 (holotype, MO).

Pleurothallis dewildeiLuer & R.Escobar, Orquideología 21: 86-88. 1998. Acronia dewildei (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 125. 2005.

Illustration: Luer & Escobar 1998, p. 87; Luer 2005, f. 71 (p. 229); Jiménez et al. 2018, f. 2c, f. 2d, f.4a, f. 5b.

Distribution: *COL.

Type: Colombia, Luer 16854 (holotype, MO).

Pleurothallis diabolica Luer & R.Escobar, Orquideología 14(2): 142. 1981. Acronia diabolica (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 125. 2005. Zosterophyllanthos diabolicus (Luer & R.Escobar) Szlach. & Marg., Polish Bot. J. 46: 118. 2001.

Illustration: Luer & Escobar 1981a, p. 143; Luer 2005, f. 72 (p. 229); Misas Urreta 2005, p. 533.

Distribution: COL ECU.

Type: Colombia, Luer 2161 (holotype, SEL).

Pleurothallis discoidea Lindl., Edwards’s Bot. Reg. 21: sub t. 1797. 1835. Acronia discoidea (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 128-129. 2005. Humboldtia discoidea (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1996f, 230; Luer 2005, f. 76a (p. 230), f. 76b (p. 231).

Distribution: BOL BRA COL ECU HON L.A. PER SUR VEN.

Type: Trinidad (holotype, K).

Voucher: Colombia, Luer 13093 (MO).

Pleurothallis divaricans Schltr., Repert. Spec. Nov. Regni Veg. 10(257-259): 387. 1912.

Illustration: Luer 1999a, f. 40; Luer 1975b, f. 26; Luer 2011, f. 12.

Distribution: BOL COL ECU PAN PER.

Type: Peru, Köhler s.n. [holotype, B, destroyed; lectotype (selected here), AMES-00074216, photo of type; isolectotype, F, photo of type]. Peru, Schuncke 1330 [neotype, AMES, designated by Luer (1999), here rejected].

Voucher: Colombia, Hartman 62 (CUVC).

Note: The holotype of this taxon was lost at Berlin, and no other original material is extant except for a photograph of the type kept at AMES and a duplicate of this photograph at F. Luer (1999) selected a neotype based on a different collection, but we follow Pupulin et al. (2022) in considering the photographs of the holotype a faithful reproduction of original type material. The neotype is here rejected in favor of the lectotypes.

Pleurothallis dunstervilleiFoldats, Acta Bot. Venez. 3: 371. 1968. Ancipitia dunstervillei (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1989, pl. 7.

Distribution: COL ECU PER VEN.

Type: Venezuela, Steyermark & Dunsterville 100843 (holotype, VEN).

Voucher: Colombia, Luer 2744 (SEL).

Pleurothallis duplex Luer & R.Escobar, Orquideología 16(1): 22. 1983. Ancipitia duplex (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer & Escobar 1983, p. 23; Luer 1989, pl. 8.

Distribution: *COL.

Type: Colombia, Luer et al. 6684 (holotype, SEL).

Pleurothallis eduardoi Mark Wilson, Orquideología 34(1): 41. 2017.

Illustration: Wilson et al. 2017d, f. 4-6.

Distribution: *COL.

Type: Colombia, Wilson & Mejía PL1003 (holotype, JAUM).

Note: Type number mistakenly omitted in the original protologue.

Pleurothallis erythrium Luer, Selbyana 2(4): 386. 1978. Acronia erythrium (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 132. 2005. Zosterophyllanthos erythrium (Luer) Szlach. & Kulak, Richardiana 6: 189. 2006.

Illustration: Luer 2005, f. 82 (p. 232).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 1626 (holotype, SEL).

Voucher: Colombia, Escobar 8327 (MO).

Pleurothallis escobarii (Luer) J.M.H.Shaw, Orchid Rev. 122(1308): 76. 2014. Acronia escobarii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 132, f. 83. 2005.

Illustration: Luer 2005, f. 83 (p. 232).

Distribution: *COL.

Type: Colombia, Escobar 8304 (holotype, MO).

Pleurothallis excavata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 287. 1923. Acronia excavata (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 133, f. 84. 2005. Zosterophyllanthos excavatus (Schltr.) Szlach. & Kulak, Richardiana 6(4): 189. 2006.

Illustration: Luer 2005, f. 84 (p. 233); Luer 1975c, f. 36; Misas Urreta 2005, p. 536, p. 537.

Distribution: COL COS PAN.

Type: Costa Rica, Acosta s.n. [holotype, B, destroyed; lectotype, AMES-31280 (Luer 2005)].

Voucher: Colombia, Misas Urreta 108 (HPUJ).

Note: AMES-31280 was superfluously selected as neotype by Luer (2023).

Pleurothallis excelsa Garay, Caldasia 10(47): 232-233. 1968. Elongatia excelsa (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Stelis excelsa (Garay) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001.

Illustration: Luer 1994, pl. 2 (p. 75).

Distribution: *COL.

Type: Colombia, Mejía s.n. (holotype, AMES; clonotypes, K, MO, SEL).

Pleurothallis flavomarginata A.Doucette, H.Medina & J.Portilla, Phytotaxa 295(2): 194. 2017.

Illustration: Doucette et al. 2017, f. 1, 2.

Distribution: COL ECU.

Type: Ecuador, Portilla s.n. (holotype, QCNE).

Voucher: Colombia, Zuluaga 4913 (CUVC).

Pleurothallis fonnegrae (Luer & Endara) J.M.H.Shaw, Orchid Rev. 122(1308): 76. 2014. Acronia fonnegrae Luer & Endara, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 135, f. 87. 2005.

Illustration: Luer 2005, f. 87 (p. 233).

Distribution: *COL.

Type: Colombia, Fonnegra 3496 (holotype, HUA, isotypes, MO, SEL).

Pleurothallis forceps-cancri Luer & R.Escobar, Orquideología 14(2): 144. 1981. Acronia forceps-cancri (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1981a, p. 145; Luer 1998b, f. 32.

Distribution: *COL.

Type: Colombia, Luer 612A (holotype, SEL).

Pleurothallis fossulataLuer & R.Escobar, Orquideología 21(1): 91-83. 1998. Acronia fossulata (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 135. 2005.

Illustration: Luer & Escobar 1998, p. 92, f. 9 (p. 98); Luer 2005, f. 88 (p. 234).

Distribution: COL ECU.

Type: Colombia, Escobar 5203 (holotype, JAUM; isotype, MO).

Pleurothallis francianaSierra-Ariza, Lankesteriana 23(1): 36. 2023.

Illustration: Sierra-Ariza 2023, f. 1, f. 2, f. 5a, f. 6d, f. 7c.

Distribution: *COL.

Type: Colombia, Sierra-Ariza & Bohorquez 430 (holotype, TOLI).

Pleurothallis fugax Luer & R.Escobar, Orquideología 14(2): 146. 1981. Acronia fugax (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1981a, p. 147; Luer 1998b, f. 33.

Distribution: *COL.

Type: Colombia, Luer et al. 2352 (holotype, JAUM; isotype, SEL).

Pleurothallis furcifera Luer, Selbyana 3: 108. 1976. Ancipitia furcifera (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1989, pl. 10.

Distribution: COL PER.

Type: Peru, Luer 598 (holotype, SEL).

Voucher: Colombia, Gutierrez 220 (JBB).

Pleurothallis fustifera Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 173. 1999. Lindleyalis fustifera (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004.

Illustration: Luer 1999a, f. 26a; Luer 2000a, f. 1 (p. 113); Rincón-González et al. 2020a, f. 4

Distribution: COL PER.

Type: Peru, Tokach P-48 (holotype, MO).

Voucher: Colombia, Rincón et al. 1244 (TOLI-spirit).

Pleurothallis ganymedes Luer & R.Escobar, Orquideología 14(2): 148. 1981. Acronia ganymedes (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 136. 2005. Zosterophyllanthos ganymedes (Luer & R.Escobar) Szlach. & Marg., Polish Bot. J. 46: 119. 2001.

Illustration: Luer & Escobar 1981a, p. 149; Luer 2005, f. 90 (p. 234).

Distribution: *COL.

Type: Colombia, Luer 4086 (holotype SEL; isotype JAUM).

Pleurothallis garayana (Ospina) Luer, Selbyana 1: 90. 1975. Colombiana garayana Ospina, Orquideología 8: 232. 1973.

Illustration: Luer 1989, pl. 2 (p. 59); Wilson et al. 2018a, f. 1b; Karremans & Vieira-Uribe 2020, p. 9.

Distribution: *COL.

Type: Colombia, Ospina 752 (holotype, Herb. Ospina).

Pleurothallis genychila Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 74. 1921.

Illustration: Luer 1999a, f. 11.

Distribution: COL ECU PER.

Type: Peru, Weberbauer 2135 (holotype, B, destroyed). Ecuador, Luer et al. 3429 [neotype, SEL (Luer 1999a)].

Voucher: Colombia, Bristol 811 (AMES).

Pleurothallis giraldoi Luer, Orquideología 21(2): 125-127, f. 5. 1999. Acronia giraldoi (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 138. 2005.

Illustration: Luer 1999b, p. 126; Luer 2005, f. 94 (p. 235); Misas Urreta 2005, p. 541.

Distribution: *COL.

Type: Colombia, Giraldo & Olver 97 (holotype, TULV).

Pleurothallis glabra Luer & R.Escobar, Orquideología 20: 56. 1996. Acronia glabra (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 139. 2005.

Illustration: Luer & Escobar 1996, p. 59; Luer 2005, f. 95 (p. 235).

Distribution: *COL.

Type: Colombia, Luer et al. 14187 (holotype, MO).

Pleurothallis globosa Luer & R.Escobar, Orquideología 20: 60. 1996. Acronia globosa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 140. 2005.

Illustration: Luer & Escobar 1996, p. 57 (top left), p. 61; Luer 2005, f. 96 (p. 236).

Distribution: *COL.

Type: Colombia, Escobar 5012 (holotype, JAUM; isotype, MO).

Pleurothallis glochis Luer & R.Escobar, Orquideología 14(2): 150. 1981. Acronia glochis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1981a, p. 151; Luer 1998b, f. 34.

Distribution: COL ECU.

Type: Colombia, Luer 3735 (holotype, SEL).

Pleurothallis glossopogon Rchb.f., Bonplandia (Hannover) 3: 71. 1855. Humboldtia glossopogon (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Lindleyalis glossopogon (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004.

Illustration: Luer 2000a, f. 2, f. 3 (p. 113).

Distribution: COL VEN.

Type: Colombia, Wagener s.n. [L339] (holotype, W-R-0053212; isotype, K-L)

Pleurothallis gomezii Luer & R.Escobar, Orquideología 20: 62. 1996. Acronia gomezii (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer & Escobar 1996, p. 58 (bottom left), p. 63; Luer 1998b, f.7.

Distribution: *COL.

Type: Colombia, Luer et al. 16789 (holotype, MO).

Pleurothallis gonzaloi J.S. Moreno, Rinc.-González & Gal.-Tar, Harvard Pap. Bot. 27(2): 187-220. 2022.

Illustration: Wilson et al. 2022, f. 20-22

Distribution: *COL.

Type: Colombia, Rincón & Moreno 1354 (holotype, TOLI).

Pleurothallis gracilicolumna Mark Wilson, Orquideología 35(1): 35. 2018.

Illustration: Wilson 2018, f. 9D, 10D, 12-14.

Distribution: *COL.

Type: Colombia, Mejía 002 (holotype, JAUM).

Pleurothallis gracilipedunculataFoldats, Acta Bot. Venez. 3: 374. 1968. Acronia gracilipedunculata (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 57. 2005.

Illustration: Luer 1998b, f.35.

Distribution: COL VEN.

Type: Venezuela, Dunsterville & Dunsterville 1031 (holotype, VEN).

Voucher: Colombia, Luer 7581 (SEL).

Pleurothallis grandiflora Lindl., Edwards’s Bot. Reg. 21: sub. t. 1797. 1835. Acronia grandiflora (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 140. 2005. Zosterophyllanthos grandiflorus (Lindl.) Szlach. & Marg., Polish Bot. J. 46(2): 119. 2001. Humboldtia grandiflora (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 2005, f. 97 (p. 236).

Distribution: COL ECU.

Type: Ecuador, Hall s.n. (holotype, K-L, specimen on the right).

Voucher: Colombia, Escobar 5026 (MO).

Pleurothallis gratiosa Rchb.f., Bonplandia (Hannover) 2: 25. 1854. Ancipitia gratiosa (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004. Humboldtia gratiosa (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1989, pl. 11.

Distribution: COL VEN.

Type: Venezuela, Wagener 116 [lectotype (selected here), W-R-0025590, type illustr.].

Voucher: Colombia, Acosta-Arteaga 581 (COL).

Note: The holotype seems to be lost. But the sheet marked as type at W-R bears several elements, including a very good full illustration which agrees with the protologue.

Pleurothallis gustavoi Mark Wilson, Orquideología 34(1): 36. 2017.

Illustration: Wilson et al. 2017d, f. 1-3.

Distribution: *COL.

Type: Colombia, Wilson & Aguirre PL1002 (holotype, JAUM).

Pleurothallis habenula Luer & R.Escobar, Orquideología 14(2): 152. 1981. Acronia habenula (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1981a, p. 153; Luer 1998b, f. 36; Ortiz & Medina 2012, f. 12.

Distribution: COL ECU.

Type: Colombia, Luer et al. 3119 (holotype, SEL).

Pleurothallis harpago Luer, Selbyana 5(2): 167. 1979. Ancipitia harpago (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1989, pl. 12.

Distribution: COL PAN.

Type: Panama, Luer 2798 (holotype, SEL).

Voucher: Colombia, Hartman 12423B (MO).

Note: A specimen collected exactly on the Panama/Colombia border was cited in Kolanowska (2014).

Pleurothallis hartwegii Lindl., Edwards’s Bot. Reg. 28: Misc. 80. 1842. Humboldtia hartwegii (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 63.

Distribution: COL ECU.

Type: Ecuador, Hartweg s.n. (holotype, K-L000820779; isotype, BM-000074384).

Voucher: Colombia, Hartweg 1406 (K).

Pleurothallis helleri A.D.Hawkes, Phytologia 14(1): 12-14. 1966.

Illustration: Luer 1998b, f. 58.

Distribution: COL COS ECU NIC.

Type: Nicaragua, Heller s.n. (holotype, F; isotype, AMES).

Voucher: Colombia, Misas Urreta 265 (COL).

Pleurothallis hemisphaerica Luer & R.Escobar, Orquideología 14(2): 154. 1981.

Illustration: Luer & Escobar 1981a, p. 155; Luer 1999a, f. 64.

Distribution: *COL.

Type: Colombia, Luer et al. 2349 (holotype, SEL; isotype, JAUM).

Pleurothallis hippocrepica Luer & R.Escobar, Orquideología 16(2): 165. 1984.

Illustration: Luer & Escobar 1984b, p. 166; Luer 1999a, f. 12.

Distribution: COL ECU.

Type: Colombia, Escobar et al. 2704 (holotype, SEL).

Pleurothallis hispidula (Luer & R.Escobar) J.M.H.Shaw, Orchid Rev. 122(1308): 77. 2014. Acronia hispidula Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 142, f. 100. 2005.

Illustration: Luer 2005, f. 100 (p. 237).

Distribution: *COL.

Type: Colombia, Luer et al. 8923 (holotype, SEL).

Pleurothallis hitchcockii Ames, Orchidaceae 7: 117. 1922. Talpinaria hitchcockii (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 2011, f. 60.

Distribution: COL ECU GUY VEN.

Type: Guyana, Hitchcock 17394 (holotype, AMES; isotype, HB).

Voucher: Colombia, Callejas & Marulanda 6030 (HUA, MO).

Pleurothallis holtonii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 52: 76, 78, pl. 4. 1994. Elongatia holtonii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004.

Illustration: Luer 1994, pl. 4 (p. 79).

Distribution: *COL.

Type: Colombia, Holton s.n. (holotype, NY).

Pleurothallis homalantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 24, 106. 1923. Acronia homalantha (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 143, f. 102. 2005. Zosterophyllanthos homalanthus (Schltr.) Szlach. & Marg., Polish Bot. J. 46(2): 119. 2001. Zosterophyllanthos homalanthus (Schltr.) Szlach. & Kulak, Richardiana 6(4): 189. 2006, nom. superfl.

Illustration: Luer 2005, f. 102 (p. 237).

Distribution: COL COS HON NIC PAN.

Type: Costa Rica, Brade 1160 [holotype, B, destroyed; lectotype, HB (Luer 2005), AMES-31260, illustration of type].

Voucher: Colombia, Contreras s.n. (UPTC).

Note: The HB specimen was superfluously selected as lectotype by Luer (2023).

Pleurothallis imbaburae Luer & Hirtz, Lindleyana 11(3): 163. 1996.

Illustration: Luer 1999a, f. 14; Luer 1996c, f. 28.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 15167 (holotype, MO).

Voucher: Colombia, Silverstone-Sopkin et al. 1606, 1627 (CUVC, MO).

Pleurothallis imber-florum Luer & R.Escobar, Orquideología 16(2): 167. 1984.

Illustration: Luer & Escobar 1984b, p. 168; Luer 1999a, f. 66.

Distribution: *COL.

Type: Colombia, Escobar 2451 (holotype, SEL).

Pleurothallis incongrua Luer, Orquideología 21(2): 127-130. 1999. Mixis incongrua (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004.

Illustration: Luer 1999b, p. 128; Luer 2007, f. 1 (p. 104).

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 2686 (holotype, CUVC; isotype, MO).

Pleurothallis indecora Rodr.-Mart. & Karremans, Lankesteriana 17(2): 251. 2017.

Illustration: Karremans et al. 2017, f. 1, f. 5.

Distribution: *COL.

Type: Colombia, Rodríguez-Martínez & RincónUseche 084 (holotype, COL).

Pleurothallis index Luer, Phytologia 47(2): 76. 1980. Acronia index (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 145, f. 105. 2005. Zosterophyllanthos index (Luer) Szlach. & Kulak, Richardiana 6: 189. 2006.

Illustration: Luer 2005, f. 105 (p. 238).

Distribution: COL ECU.

Type: Colombia, Luer et al. 4582 (holotype, SEL).

Pleurothallis inflata Rolfe, Bull. Misc. Inform. Kew 1894: 154. 1894.

Illustration: Luer 1999a, f. 67.

Distribution: COL VEN.

Type: Venezuela, Sander s.n. (holotype, K).

Voucher: Colombia, Wagener s.n. (W).

Pleurothallis iris (Luer & Endara) J.M.H.Shaw, Orchid Rev. 122(1308): 77. 2014. Acronia iris Luer & Endara, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 145. 2005.

Illustration: Luer 2005, f. 106 (p. 238).

Distribution: *COL.

Type: Colombia, Luer et al. 8932 (holotype, MO).

Pleurothallis jaculiferaLuer & R.Escobar, Orquideología 21: 93. 1998.

Illustration: Luer & Escobar 1998, p. 94, f. 10 (p. 97); Luer 1999a, f. 68.

Distribution: *COL.

Type: Colombia, Luer et al. 16843 (holotype, JAUM; isotype, MO).

Pleurothallis jaramilloi Luer, Orquideología 22(1): 56-58. 2001. Acronia jaramilloi (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 146. 2005. Zosterophyllanthos jaramilloi (Luer) Szlach. & Kulak, Richardiana 6: 189. 2006.

Illustration: Luer 2001b, p. 57; Luer 2005, f. 108 (p. 239).

Distribution: *COL.

Type: Colombia, Jaramillo 20254 (holotype, COL; isotype, AMES).

Pleurothallis juvenilis Rodr.-Mart. & Karremans, Lankesteriana 17(2): 253. 2017.

Illustration: Karremans et al. 2017, f. 2, f. 5.

Distribution: *COL.

Type: Colombia, Rodríguez-Martínez 275 (holotype, VALLE).

Pleurothallis killipii Garay, Canad. J. Bot. 34: 254. 1956. Acronia killipii (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 147, f. 110. 2005. Zosterophyllanthos killipii (Garay) Szlach. & Marg., Polish Bot. J. 46: 119. 2001.

Illustration: Luer 2005, f. 110 (p. 239).

Distribution: *COL.

Type: Colombia, Killip 34061 (holotype, US; isotype, AMES).

Pleurothallis lacera Luer, Selbyana 1(3): 248. 1975. Acronia lacera (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 148. 2005. Zosterophyllanthos lacer (Luer) Szlach. & Marg., Polish Bot. J. 46: 119. 2001.

Illustration: Luer 2005, f. 111 (p. 239); Luer 1975f, f. 59.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 550 (holotype, SEL).

Voucher: Colombia, Lehmann 6283 (K).

Pleurothallis lacrima K.W.Holcomb, Pleurothallidinae 1(4): 1. 2022.

Illustration: Holcomb 2022b, f. 1-3.

Distribution: *COL.

Type: Colombia? [uncertain origin], Holcomb s.n. (holotype, GEO-18024)

Note: Described from a cultivated specimen from uncertain origin.

Pleurothallis lamellaris Lindl., Fol. Orchid. ~Pleurothallis~ 8. 1859. Humboldtia lamellaris (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1999a, f. 69; Luer 1996c, f. 33.

Distribution: BOL COL ECU PER.

Type: Bolivia, Bridges s.n. (holotype, K).

Voucher: Colombia, Escobar 8393 (MO)

Pleurothallis languida Luer, Orquideología 21: 95. 1998. Acronia languida (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1998, p. 96, f. 11 (p. 98); Luer 1998b, f. 10.

Distribution: *COL.

Type: Colombia, Luer 17564 (holotype, MO).

Pleurothallis latipetala Garay, Orchid J. 1: 218. 1952. Acronia latipetala (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 148. 2005.

Illustration: Wilson et al. 2018b, f. 1h; Luer 2005, f. 112 (p. 240).

Distribution: *COL.

Type: Colombia, von Sneidern 1602 (holotype, S; isotypes AMES).

Pleurothallis laurifolia Kunth, Nov. Gen. Sp. (quarto ed.) 1: 364. 1816. Humboldtia laurifolia (Kunth) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1991e, f. 489; Luer 1998b, f. 53.

Distribution: COL ECU.

Type: Colombia, Kunth s.n. (holotype, P; isotypes, P).

Note: This name has remained mostly obscure in literature and was not monographed by Luer. However, three good specimens are available at P. It is our interpretation that P. laurifolia is an earlier name for Pleurothallis chloroleuca Lindl.

Pleurothallis leopardina Luer, Phytologia 49(3): 209. 1981.

Illustration: Luer 1999a, f. 41; Luer 1996c, f. 10.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6190 (holotype, SEL).

Voucher: Colombia, Lehmann 7142 (K).

Pleurothallis lilijaeFoldats, Acta Bot. Venez. 3: 379. 1968. Acronia lilijae (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 150, f. 115a. 2005. Zosterophyllanthos lilijae (Foldats) Szlach. & Marg., Polish Bot. J. 46: 119. 2001.

Illustration: Luer 2005, f. 115a (p. 240), f. 115b (p. 241).

Distribution: BOL COL ECU VEN.

Type: Venezuela, Steyermark & Dunsterville 98549 (holotype, VEN; isotype AMES).

Voucher: Colombia, Luer 12540 (MO).

Pleurothallis lindenii Lindl., Ann. Mag. Nat. Hist. 12: 397. 1843. Humboldtia lindenii (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer & Escobar 1996, p. 35; Luer 1999a, f. 70.

Distribution: COL ECU PER VEN.

Type: Venezuela, Linden 630 (holotype, K; isotypes, AMES, BM-000082379, K-L, P, W).

Voucher: Colombia, Luer 10189 (MO).

Pleurothallis linguifera Lindl., Fol. Orchid. ~Pleurothallis~ 10, n. 41. 1859. Acronia linguifera (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 151, f. 116. 2005. Humboldtia linguifera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Zosterophyllanthos linguifer (Lindl.) Szlach. & Marg., Polish Bot. J. 46(2): 119. 2001.

Illustration: Luer 2004, f. 10; Luer 2005, f. 116 (p. 241).

Distribution: BOL COL ECU PER.

Type: Bolivia, Bridges s.n. (holotype, K).

Voucher: Colombia, Luer 7561 (SEL).

Pleurothallis litotes Luer, Selbyana 5(2): 170. 1979.

Illustration: Luer 1999a, f. 42.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 2700 (holotype, SEL).

Voucher: Colombia, Luer 2997 (SEL).

Pleurothallis lobata Luer, Orquideología 20: 214. 1996. Acronia lobata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1996f, 213; Luer 1998b, f. 37.

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K).

Pleurothallis londonoi Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 61(4): 6. 1996. Ancipitia londonoi (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1996b, pl. 24.

Distribution: *COL.

Type: Colombia, Londoño et al. 83 (holotype, HUA).

Pleurothallis lopeziiLuer & R.Escobar, Orquideología 21: 102. 1998. Acronia lopezii (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1998, f. 13 (p. 98), p. 103; Luer 1998b, f. 12.

Distribution: *COL.

Type: Colombia, Escobar 8291 (holotype, JAUM; isotype, MO).

Pleurothallis loranthophylla Rchb.f., Bot. Zeitung (Berlin) 10(39): 674. 1852. Humboldtia loranthophylla (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis punctata (H.Karst.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 6: 63. 1919, nom. illeg. Rhynchopera loranthophylla (Rchb.f.) Szlach. & Marg., Polish Bot. J. 46(2): 118. 2001. Rhynchopera punctata H.Karst., Auswahl Gew. Venez. 21, t. 7. 1848. Non Pleurothallis punctata Ker Gawl. [1824].

Illustration: Luer 1998a, f. 5 (p. 93).

Distribution: BOL COL COS ECU PAN PER VEN.

Type: Venezuela, Wagener 148 (holotype, W).

Voucher: Colombia, Purdie s.n. (K).

Pleurothallis lueriana Karremans & Rodr.-Mart., Lankesteriana 17(2): 255. 2017.

Illustration: Luer 2004, f. 9 (p. 242); Karremans et al. 2017, f. 3, f. 5.

Distribution: *COL.

Type: Colombia, Rodríguez-Martínez 220 (holotype, COL).

Pleurothallis lunaris Luer & R.Escobar, Orquideología 14(2): 158. 1981. Acronia lunaris (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 153, f. 119. 2005. Zosterophyllanthos lunaris (Luer & R.Escobar) Szlach. & Kulak, Richardiana 6: 190. 2006.

Illustration: Luer & Escobar 1981a, p. 159; Luer 2005, f. 119 (p. 242).

Distribution: *COL.

Type: Colombia, Escobar 1156 (holotype, JAUM).

Pleurothallis macrocardia Rchb.f., Bonplandia (Hannover) 3: 72. 1855. Acronia macrocardia (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 155. 2005. Humboldtia macrocardia (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 2005, f. 122 (p. 242).

Distribution: BOL COL ECU.

Type: Colombia, Wagener s.n. (holotype, W).

Pleurothallis macrophylla Kunth, Nov. Gen. Sp. (quarto ed.) 1: 365. 1816. Elongatia macrophylla (Kunth) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Humboldtia macrophylla (Kunth) Kuntze, Revis. Gen. Pl. 2: 667. 1891.

Illustration: Luer 1994, pl. 6 (p. 83); Uribe-Velez et al. 2022a, p. 3-7; Uribe-Velez et al. 2022b, p. 4-6.

Distribution: *COL.

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Pleurothallis magnifica Luer & R.Escobar, Orquideología 16(2): 169. 1984.

Illustration: Luer & Escobar 1984b, p. 170; Luer 1999a, f. 71.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8374 (holotype, SEL).

Pleurothallis manicosa Luer & R.Escobar, Orquideología 20: 66. 1996.

Illustration: Luer & Escobar 1996, p. 57 (top right), p. 67; Luer 1999a, f. 18.

Distribution: *COL.

Type: Colombia, Luer 17554 (holotype, JAUM; isotype, MO).

Pleurothallis marthae Luer & R.Escobar, Orquideología 20: 68. 1996. Acronia marthae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 157. 2005. Zosterophyllanthos marthae (Luer & R.Escobar) Szlach., Ill. Field Guide Orchids Yotoco Forest Reserve 279. 2011, nom. superfl. Zosterophyllanthos marthae (Luer & R.Escobar) Szlach. & Kulak, 134. 2006.

Illustration: Luer & Escobar 1996, p. 69; Luer 2005, f. 125 (p. 243).

Distribution: *COL.

Type: Colombia, Luer 16689 (holotype, JAUM; isotype, MO).

Pleurothallis mark-wilsonii J.S.Moreno, Gal-Tar & Sierra-Ariza, Phytotaxa 568(3): 285. 2022.

Illustration: Moreno et al. 2022c, f. 1, f. 2.

Distribution: *COL.

Type: Colombia, Galindo-Tarazona & Mora 3927 (holotype, CUVC).

Pleurothallis matudana C.Schweinf., Bot. Mus. Leafl. 5(6): 102-103. 1938. Acronia matudana (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 158-159. 2005. Zosterophyllanthos matudanus (C.Schweinf.) Szlach. & Marg., Polish Bot. J. 46(2): 120. 2001.

Illustration: Luer 2005, f. 127a, f. 127b (p. 244).

Distribution: BOL COL COS ECU ELS GUA HON MEX NIC PER.

Type: Mexico, Matuda 1577 (holotype, AMES).

Voucher: Colombia, Franco 7968 (COL).

Pleurothallis mahechae J.S. Moreno, Sierra-Ariza & L.C.Pina, Harvard Pap. Bot. 27(2): 213. 2022.

Illustration: Wilson et al. 2022, f. 23-26.

Distribution: *COL.

Type: Colombia, Santafé Millán & Suarez Romero s.n. (holotype, HUEB-TRA-3855).

Pleurothallis medusa Luer, Lindleyana 12(1): 51. 1997.

Illustration: Luer 1998b, f.59.

Distribution: COL ECU.

Type: Unknown, O-1613 (holotype, W).

Voucher: Colombia, Rincón 274 (TOLI).

Note: The type locality was assumed to be San José, Costa Rica, but Colombia is more likely. The collector, who signed only with a Greek Phi, is unknown but suggested to be Kalbreyer.

Pleurothallis megalorhina Luer & R.Escobar, Orquideología 20: 71. 1996. Acronia megalorhina (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 159. 2005.

Illustration: Luer & Escobar 1996, p. 72; Luer 2005, f. 129 (p. 244).

Distribution: *COL.

Type: Colombia, Luer et al. 11389 (holotype, MO).

Pleurothallis membracidoides Luer, Phytologia 47(2): 77. 1980. Ancipitia membracidoides (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 254. 2004.

Illustration: Luer 1989, pl. 15.

Distribution: *COL.

Type: Colombia, Luer et al. 4580 (holotype, SEL).

Pleurothallis microcardia Rchb.f., Bonplandia (Hannover) 3: 72. 1855. Acronia microcardia (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 160. 2005. Humboldtia microcardia (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 2005, f. 130, f. 131, f. 132 (p. 245).

Distribution: BOL COL ECU VEN.

Type: Colombia, Wagener 610 (holotype, W; isotype, K).

Pleurothallis millei Schltr., Repert. Spec. Nov. Regni Veg. 14: 131. 1914 [1915]. Acronia millei (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 162. 2005. Zosterophyllanthos millei Szlach. & Kulak, Richardiana 6: 190. 2006.

Illustration: Luer 2005, f. 133 (p. 245).

Distribution: COL ECU.

Type: Ecuador, Mille 35 (holotype, B, destroyed). Ecuador, Luer & Kent 273 [neotype, SEL (Luer 2005)].

Voucher: Colombia, Ramos 6104 (COL).

Pleurothallis miranda Luer, Phytologia 46(6): 369. 1980. Mirandopsis miranda (Luer) Szlach. & Marg., Polish Bot. J. 46(2): 117. 2001 [2002].

Illustration: Luer 2007, pl. 22.

Distribution: COL ECU.

Type: Ecuador, Luer 4372 (holotype, SEL).

Voucher: Colombia, Funck 1441 (MPU).

Pleurothallis mocoana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 49. 1924. Specklinia mocoana (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Distribution: *COL.

Type: Colombia, Hopp 69 (syntype, B, destroyed). Colombia, Hopp 77 (syntype, B, destroyed).

Note: Nothing remains of the type material and the placement of this species is uncertain. The description suggests a species of Acianthera, but it could also be a Specklinia.

Pleurothallis mundiflorae K.W.Holcomb ex Karremans, sp. nov.

Pleurothallis mundiflorae K.W.Holcomb, Pleurothallidinae 2(1): 1. 2023, nom. inval.

Illustration: Holcomb 2023, f. 1, f. 2

Distribution: COL ECU?

Type: Ecuador? [uncertain origin], Holcomb s.n. (holotype, GEO-18302).

Voucher: A photographic voucher of a Colombian specimen reproduced in Holcomb 2023, f. 2.

Note: This taxon was published invalidly because the type is a mixture of two localities of the two gatherings discussed in the text, a specimen of uncertain origin allegedly from Ecuador and an in situ photograph of a specimen from Valle de Sibundoy in Colombia.

Pleurothallis mundula Luer & R.Escobar, Orquideología 16(1): 26. 1983.

Illustration: Luer & Escobar 1983, p. 27; Luer 1999a, f. 73.

Distribution: *COL.

Type: Colombia, Luer et al. 7502 (holotype, SEL; isotypes, COL, JAUM).

Pleurothallis narinoensis Rykacz. & Kolan., Polish Bot. J. 61(2): 250. 2016.

Illustration: Rykaczewski & Kolanowska 2016, f. 1.

Distribution: *COL.

Type: Colombia, Garay et al. 424 (holotype, MO).

Pleurothallis nasiterna Luer, Phytologia 49(3): 212. 1981.

Illustration: Luer 1999a, f. 20.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6287 (holotype, SEL).

Voucher: Colombia, André s.n. (K).

Pleurothallis navilinguis Rchb.f., Bonplandia (Hannover) 3: 72. 1855. Humboldtia navilinguis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Distribution: *COL.

Type: Colombia, Wagener s.n. (holotype, W).

Note: An obscure taxon, no type specimen located.

Pleurothallis neorinkei A.Doucette, Orquideología 33(2): 130. 2016. Acronia rinkeiLuer, Harvard Pap. Bot. 16: 316, f. 6. 2011.

Illustration: Luer 2011, f. 6; Doucette et al.. 2016a, f. 6.

Distribution: COL ECU?

Type: Ecuador, Rinke 0701 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Doucette et al. 2016a, f. 6.

Pleurothallis nossax Luer & R.Escobar, Orquideología 14(2): 160. 1981. Acronia nossax (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 164. 2005. Zosterophyllanthos nossax (Luer & R.Escobar) Szlach. & Kulak, Richardiana 6: 191. 2006.

Illustration: Luer & Escobar 1981a, p. 161; Luer 2005, f. 138 (p. 247).

Distribution: *COL.

Type: Colombia, Escobar et al. 1520 (holotype, JAUM).

Pleurothallis notabilis Luer & R.Escobar, Orquideología 14(2): 162. 1981.

Illustration: Luer & Escobar 1981a, p. 163; Luer 1998b, f. 60.

Distribution: *COL.

Type: Colombia, Luer 2085 (holotype, SEL; isotype, JAUM).

Pleurothallis nox-mediaLuer & R.Escobar, Orquideología 21: 104. 1998. Acronia nox-media (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1998, p. 105; Luer 1998b, f. 14.

Distribution: *COL.

Type: Colombia, Luer 17987 (holotype, MO).

Pleurothallis nuda (Klotzsch) Rchb.f., Ann. Bot. Syst. 6: 184. 1861. Lindleyalis nuda (Klotzsch) Luer & Thoerle, Harvard Pap. Bot. 17(2): 368. 2012. Restrepia nuda Klotzsch, Allg. Gartenzeitung 20: 275. 1852.

Illustration: Luer 1975a, f. 10 (p. 77); Luer 2000a, f. 4 (p. 113).

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Wagener 91 (holotype, W-R0025589).

Voucher: Colombia, Linden s.n. (W).

Pleurothallis nutans Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 165. 1924.

Illustration: Luer & Escobar 1983, p. 21; Luer 1999a, f. 74.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed). Colombia, Luer et al. 7922 [neotype (selected here), SEL; isoneotype, JAUM].

Note: Luer (1999) selected Luer et al. 7922 as neotype, but failed to specify which one of two cited specimens (SEL and JAUM). It is important to note that this is the type number of Pleurothallis decora Luer & R.Escobar, which therefore is rendered conspecific.

Pleurothallis octavioi Luer & R.Escobar, Orquideología 14(2): 164. 1981. Acronia octavioi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 165. 2005. Zosterophyllanthos octavioi (Luer & R.Escobar) Szlach. & Kulak, Richardiana 6: 191. 2006.

Illustration: Luer & Escobar 1981a, p. 165; Luer 1991e, f. 492; Luer 2005, f. 140 (p. 247); Wilson et al. 2018b, f. 1i.

Distribution: *COL.

Type: Colombia, Luer et al. 3725 (holotype, SEL).

Pleurothallis odobenicepsLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 31: 38, pl. 17. 1989. Ancipitia odobeniceps (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004.

Illustration: Luer 1989, pl. 17.

Distribution: *COL.

Type: Colombia, Zarucchi et al. 5950 (holotype, MO).

Pleurothallis orecta Luer & R.Escobar, Orquideología 20(1): 75. 1996. Acronia orecta (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1996, p. 57 (left middle), p. 76; Luer 1998b, f.38.

Distribution: *COL.

Type: Colombia, Escobar 5106 (holotype, JAUM; isotype, MO).

Pleurothallis orthostachys Luer & R.Escobar, Orquideología 16(2): 171. 1994.

Illustration: Luer & Escobar 1984b, p. 172; Luer 1999a, f. 75.

Distribution: COL ECU.

Type: Colombia, Escobar & Valencia 2622 (holotype, SEL).

Pleurothallis ottocarii Rodr.-Mart., Rinc.-Useche & Karremans, Syst. Bot. 40(1): 75. 2015.

Illustration: Wilson et al. 2018a, f. 1c.

Distribution: *COL.

Type: Colombia, Rodríguez-Martínez & RincónUseche 057 (holotype, COL).

Pleurothallis paraniesseniae J.M.H.Shaw, Orchid Rev. 122(1308): 77. 2014. Acronia niesseniaeLuer, Harvard Pap. Bot. 16(2): 314, 5. 2011.

Illustration: Luer 2011, f. 5.

Distribution: *COL.

Type: Colombia, Niessen 02/11 (holotype, MO).

Pleurothallis pedunculata (Link, Klotzsch & Otto) Rchb.f., Linnaea 22: 822. 1849. Humboldtia pedunculata (Link, Klotzsch & Otto) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Rhynchopera pedunculata Link, Klotzsch & Otto, Icon. Pl. Rar. 2: 103, t. 41. 1844.

Illustration: Luer 1998a, f. 7 (p. 93).

Distribution: COL VEN.

Type: Venezuela, Otto s.n. (holotype, W).

Voucher: Colombia, Escobar 8270 (MO).

Pleurothallis penduliflora Kraenzl., Repert. Spec. Nov. Regni Veg. 1: 177. 1905.

Illustration: Luer & Escobar 1981a, p. 135; Luer 1999a, f. 22, f. 23.

Distribution: COL ECU PER.

Type: Peru, Weberbauer 2118 [holotype, B, destroyed; lectotype, MOL (selected here); photo of type, AMES, F]. Peru, Vargas 18426 [neotype, AMES; isoneotype, CUZ (Luer 1999a); rejected here].

Voucher: Colombia, Escobar et al. 956 (holotype, JAUM).

Note: The holotype of this taxon was lost at Berlin. Luer (1999) selected a neotype based on a different collection, but an isotype and photographs of the holotype are available for lectotypification. The neotype is here rejected in favor of selecting the isotype as lectotype.

Pleurothallis penicillata Luer, Phytologia 47(2): 78. 1980. Colombiana penicillata (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer 1989, pl. 3 (p. 61).

Distribution: *COL.

Type: Colombia, Luer et al. 4610 (holotype, SEL).

Pleurothallis perijaensis Dunst., Selbyana 2(2/3): 210. 1978. Acronia perijaensis (Dunst.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 171. 2005. Zosterophyllanthos perijaensis (Dunst.) Szlach. & Marg., Polish Bot. J. 46: 120. 2001.

Illustration: Luer 2005, f. 153 (p. 250).

Distribution: COL ECU VEN.

Type: Venezuela, Steyermark & Dunsterville 105666 (holotype, VEN; isotype, AMES).

Voucher: Colombia, Escobar 7778 (COL).

Pleurothallis petroanaSierra-Ariza, Lankesteriana 23(1): 39. 2023.

Illustration: Sierra-Ariza 2023, f. 3, f. 4, f. 5b, f. 6a, f. 7e.

Distribution: *COL.

Type: Colombia, Sierra-Ariza & Bohorquez 194 (holotype, TOLI)

Pleurothallis phalangifera (C.Presl) Rchb.f., Ann. Bot. Syst. 6: 168. 1861. Pleurothallis phalangifera (C.Presl) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 141. 1921, nom. superfl. Acronia phalangifera C.Presl, Reliq. Haenk. 1(2): 104. 1827. Humboldtia phalangifera (C.Presl) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 1998b, f. 16.

Distribution: COL ECU PER VEN.

Type: Peru, Haenke s.n. (holotype, PR?).

Voucher: Colombia, Linden 1271 (K, W).

Pleurothallis phratria Luer & Hirtz, Lindleyana 11(3): 179. 1996.

Illustration: Luer 1999a, f. 77; Luer 1996c, f. 48.

Distribution: COL ECU.

Type: Ecuador, Hirtz & Dodson 2028 (holotype, MO).

Voucher: Colombia, Hartmann 115 (CUVC).

Pleurothallis phyllocardioides Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 193. 1923. Acronia phyllocardioides (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 174-176. 2005. Zosterophyllanthos phyllocardioides (Schltr.) Szlach. & Kulak, Richardiana 6(4): 192. 2006, nom. superfl. Zosterophyllanthos phyllocardioides (Schltr.) Szlach. & Marg., Polish Bot. J. 46(2): 120. 2001.

Illustration: Luer 2005, f. 158a, f. 158b, f.158c (p. 252).

Distribution: BOL COL COS ECU GUA GUY NIC PAN PER VEN.

Type: Costa Rica, Brenes 290 [holotype, B, destroyed; isotypes, AMES-56387, NY; drawing of type, AMES].

Voucher: Colombia, Gentry 56778 (MO).

Pleurothallis pileataLuer & R.Escobar, Orquideología 15(2-3): 142. 1982. Acronia pileata (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 176. 2005.

Illustration: Luer & Escobar 1982, p. 143; Luer 2005, f. 160 (p. 253).

Distribution: *COL.

Type: Colombia, Luer 6723 (holotype, SEL).

Pleurothallis platypetala Luer & R.Escobar, Orquideología 20: 78. 1996. Acronia platypetala (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 177. 2005.

Illustration: Luer & Escobar 1996, p. 77; Luer 2005, f. 161 (p. 253).

Distribution: *COL.

Type: Colombia, Luer et al. 12539 (holotype, MO).

Pleurothallis platysepala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 111. 1920. Acronia platysepala (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 178. 2005.

Illustration: Luer & Escobar 1984b, p. 163; Luer 1991e, f. 490; Luer 2005, f. 162 (p. 253); Wilson et al. 2018b, f. 1l.

Distribution: *COL.

Type: Colombia, Lehmann 6930 [holotype, B, destroyed; lectotype (selected here), K-000584312; isolectotype, K-000584310].

Pleurothallis poculifera Luer & R.Escobar, Orquideología 14(2): 166. 1981.

Illustration: Luer & Escobar 1981a, p. 167; Luer 1999a, f. 43.

Distribution: *COL.

Type: Colombia, Luer 3008 (holotype, SEL).

Pleurothallis polysticta Luer, Selbyana 5(2): 176. 1979. Acronia polysticta (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.17.

Distribution: *COL.

Type: Panama, Luer 2799 (holotype, SEL).

Voucher: Colombia, Luer 16889 (MO).

Pleurothallis prolaticollaris Luer, Selbyana 3(3,4): 368. 1977. Acronia prolaticollaris (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 179. 2005. Pleurothallis prolaticollaris Luer, Selbyana 3(3,4): 368. 1977.

Illustration: Luer 2005, f. 164 (p. 254).

Distribution: COL ECU.

Type: Ecuador, Luer & Kent 334 (holotype, SEL).

Voucher: Colombia, Monsalve 1420 (MO).

Pleurothallis pruinosa Lindl., Edwards’s Bot. Reg. 28: Misc. 75-76. 1842. Humboldtia pruinosa (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 1999a, f. 44, Stenzel 2004, f. 31; Luer 1975f, f. 68; Misas Urreta 2005, p. 553.

Distribution: BRA COL COS CUB DOM ECU ELS GUA GUY HON JAM L.A. NIC PAN PER PUE SUR VEN.

Type: Suriname, Splitgerber 527 (holotype, K; isotype, W).

Voucher: Colombia, Cuatrecasas 14049 (AMES).

Pleurothallis pulvinaris Luer & R.Escobar, Orquideología 16(2): 176. 1984.

Illustration: Luer & Escobar 1984b, p. 177; Luer 1999a, f. 25

Distribution: *COL.

Type: Colombia, Escobar et al. 2685 (holotype, SEL; isotype, JAUM).

Pleurothallis punctulata Rolfe, Gard. Chron., ser. 3 4(105): 756. 1888. Humboldtia punctulata (Rolfe) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Talpinaria punctulata (Rolfe) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 1998a, f. 2 (p. 100).

Distribution: COL VEN.

Type: Colombia, Veitch s.n. (holotype, K).

Pleurothallis quadricaudata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 112. 1920. Acronia quadricaudataLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b. f.40.

Distribution: *COL.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 39, No. 149)].

Pleurothallis quadrifida (Lex.) Lindl., Edwards’s Bot. Reg. 28: Misc. 70. 1842. Dendrobium quadrifidum Lex., Nov. Veg. Descr. 2(Orchid. Opusc.): 40. 1825. Humboldtia quadrifida (Lex.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Lalexia quadrifida (Lex.) Luer, Harvard Pap. Bot. 16: 358. 2011. Loddigesia quadrifida (Lex.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 251. 2006. Specklinia quadrifida (Lex.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Stelis quadrifida (Lex.) Solano & Soto Arenas, Icon. Orchid. (Mexico) 5-6: xi. 2002 [2003].

Illustration: Luer 2000a, f. 34; Luer 1975c, f. 38; Stenzel 2004, f. 17.

Distribution: COL COS CUB DOM ELS GUA HAI HON JAM L.A. MEX NIC PUE VEN.

Type: Mexico, Hilton 13643 [neotype, ARIZ12480 (Meneguzzo & van den Berg 2020)].

Voucher: Colombia, Yepes 211 (UTMC).

Pleurothallis quaternaria Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 31. 1998. Acronia quaternaria (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.41.

Distribution: *COL.

Type: Colombia, Luer 18418 (holotype, MO).

Pleurothallis queremalensisRinc.-Useche, Rodr.-Mart. & Karremans, Lankesteriana 17(2): 257.2017.

Illustration: Karremans et al. 2017, f. 4, f. 5.

Distribution: *COL.

Type: Colombia, Rodríguez-Martínez & RincónUseche 092 (holotype, VALLE).

Pleurothallis questionis Luer & R.Escobar, Orquideología 10: 79. 1996.

Illustration: Luer & Escobar 1996, p. 57 (bottom left), p. 80; Luer 1999a, f. 85.

Distribution: *COL.

Type: Colombia, Luer et al. 16803 (holotype, MO).

Pleurothallis radula Luer, Selbyana 5(2): 177-178. 1979. Acronia radula (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 179. 2005. Zosterophyllanthos radula (Luer) Szlach. & Marg., Polish Bot. J. 46(2): 120. 2001.

Illustration: Luer 2005, f. 165 (p. 254); Pupulin et al. 2021, f. 2(26), f. 7, f. 47, f. 48a-c.

Distribution: COL COS.

Type: Costa Rica, Luer et al. 4251 (holotype, SEL; isotype, K).

Voucher: Colombia, Luer et al. 10097 (MO, US).

Pleurothallis ramificans Luer, Phytologia 46(6): 371. 1980.

Illustration: Luer 1999a, f. 26.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3920 (holotype, SEL).

Voucher: Colombia, Lehmann 7159 (AMES, K).

Pleurothallis ramiromedinaeThoerle & Hirtz, OrchideenJ. 6(1): 6. 2018.

Illustration: Thoerle et al. 2018, p. 6-8.

Distribution: COL ECU.

Type: Ecuador, Hirtz 10943 (holotype, Univ. Católica de Ibarra).

Voucher: No Colombian voucher appears to be available, but see Thoerle et al. (2018).

Pleurothallis ramosii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 32. 1998. Acronia ramosii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f. 42.

Distribution: *COL.

Type: Colombia, Ramos et al.1224 (holotype, CUCV; isotype, MO).

Pleurothallis reginaeGaray, Svensk Bot. Tidskr. 47: 210. 1953. Acronia reginae (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 181. 2005. Zosterophyllanthos reginae (Garay) Szlach. & Kulak, Richardiana 6: 192. 2006.

Illustration: Garay 1953, f. 7.

Distribution: *COL.

Type: Colombia, von Sneidern 811 (holotype, S-R-4971).

Pleurothallis restrepioides Lindl., Companion Bot. Mag. 2: 356. 1836. Elongatia restrepioides (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Humboldtia restrepioides (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Stelis restrepioides (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001.

Illustration: Luer 1975a, f. 14 (p. 85); Luer 1994, pl. 7 (p. 85).

Distribution: COL ECU PER.

Type: Peru, Mathews s.n. (holotype, K; isotype, BM-000082426).

Voucher: Colombia, Escobar 5020 (MO).

Pleurothallis revoluta (Ruiz & Pav.) Garay, Caldasia 8(40): 520. 1962. Humboldtia revoluta Ruiz & Pav., Fl. Peruv. Prodr. 235. 1794. Stelis revoluta (Ruiz & Pav.) Willd., Sp. Pl. 4: 140. 1806.

Illustration: Luer 1999a, f. 27; Luer 1975f, f. 71; Pupulin 2013, f. 36c.

Distribution: BOL COL ECU L.A. PER VEN.

Type: Peru, Ruiz & Pavón (holotype, MA).

Voucher: Colombia, Escobar 5020 (MO).

Pleurothallis rhodoglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 17: 22-23. 1922. Acronia rhodoglossa (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 182. 2005. Zosterophyllanthos rhodoglossus (Schltr.) Szlach. & Marg., Polish Bot. J. 46(2): 120. 2001.

Illustration: Luer 2005, f. 169a, f. 169b (p. 255).

Distribution: COL COS ECU PAN.

Panama, Powell 182 [holotype, B, destroyed; lectotype (Christenson 1991), MO-956015; isolectotype, AMES].

Voucher: Colombia, Luer 8452 (SEL).

Pleurothallis rowleei Ames, Sched. Orch. 1: 11-12. 1922. Acronia rowleei (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.18; Misas Urreta 2005, p. 558, p. 559.

Distribution: COL COS ECU PAN.

Type: Costa Rica, Rowlee 236 (holotype, AMES).

Voucher: Colombia, von Sneidern 1603 (S, AMES).

Pleurothallis ruberrima Lindl., Orchid. Linden. 1, n. 5. 1846. Acronia ruberrima (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 184. 2005. Humboldtia ruberrima (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Zosterophyllanthos ruberrimus (Lindl.) Szlach. & Kulak, Richardiana 6: 192. 2006.

Illustration: Luer 2005, f. 172a, f. 172b, f.172c (p. 256); Luer 1975f, f. 72.

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Linden 670 (holotype, K-L).

Voucher: Colombia, Schneider 404 (S).

Pleurothallis rugosaLuer & R.Escobar, Orquideología 15(2-3): 147. 1982. Acronia rugosa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 185. 2005.

Illustration: Luer & Escobar 1982, p. 148; Luer 2005, f. 174 (p. 257).

Distribution: *COL.

Type: Colombia, Luer et al. 3053 (holotype, SEL).

Pleurothallis ruscaria Luer, Phytologia 47: 79. 1980. Colombiana ruscaria (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer 1989, pl. 4 (p. 63); Wilson et al. 2018a, f. 1d.

Distribution: COL ECU.

Type: Colombia, Luer et al. 4611 (holotype, SEL).

Pleurothallis ruscifolia (Jacq.) R.Br., Hort. Kew. (ed. 2) 5: 211. 1813. Dendrobium ruscifolium (Jacq.) Sw., Nova Acta Regiae Soc. Sci. Upsal. 6: 84. 1799. Epidendrum ruscifolium Jacq., Enum. Syst. Pl.: 29. 1760. Humboldtia ruscifolia (Jacq.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 1975a, f. 15 (p. 87); Stenzel 2004, f. 34; Luer 1998b, f.61; Misas Urreta 2005, p. 550, p. 551, p. 560, p. 561.

Distribution: BOL BRA COL COS CUB DOM ECU GUA HAI JAM L.A. PAN PER PUE VEN.

Type: Martinique, Plumier s.n. [lectotype, W (Szlachetko et al. 2012)].

Voucher: Colombia, Luer 11379 (MO).

Pleurothallis sagittilabia Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 32, f. 43. 1998. Acronia sagittilabia (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1999b, p. 131; Luer 1998b, f. 43.

Distribution: *COL.

Type: Colombia, Giraldo & Olver 557 (holotype, TULV; isotype, MO).

Pleurothallis sandemanii Luer, Orquideología 22: 222. 1996. Talpinaria sandemanii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 1996f, 223; Luer 1998a, f. 3 (p. 100).

Distribution: *COL.

Type: Colombia, Sandeman 5872 (holotype, K).

Pleurothallis sanluisiiFoldats, Acta Bot. Venez. 3: 390. 1968. Acronia sanluisii (Foldats) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.19.

Distribution: COL VEN.

Type: Venezuela, Steryermark 98995 (holotype, VEN; isotype, M).

Voucher: Colombia, Luer 18560A (MO).

Pleurothallis sannioLuer & R.Escobar, Orquideología 21(1): 106-108. 1998. Acronia sannio (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 187. 2005.

Illustration: Luer & Escobar 1998, p. 107; Luer 2005, f. 178 (p. 258).

Distribution: *COL.

Type: Colombia, Luer 17624 (holotype, MO).

Pleurothallis scabrilinguis Lindl., Companion Bot. Mag. 2: 354. 1836. Acronia scabrilinguis (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 188. 2005. Humboldtia scabrilinguis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Zosterophyllanthos scabrilinguis (Lindl.) Szlach. & Marg., Polish Bot. J. 46: 120. 2001.

Illustration: Luer 2005, f. 180 (p. 259); Luer 1975f, f. 76.

Distribution: COL ECU PER.

Type: Peru, Matthews 1904 (holotype, K-L000584193; isotypes, BR, K-L, W).

Voucher: Colombia, De Wilde 4317 (MO).

Pleurothallis secunda Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 49, pl. 85. 1836. Humboldtia secunda (Poepp. & Endl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis secunda var. secunda

Illustration: Luer 1999a, f. 31; Luer 1975f, f. 78.

Distribution: COL ECU PER.

Type: Peru, Poeppig 1746 (holotype, W).

Voucher: Colombia, Zarucchi & Bedoya 4461 (HUA, MO).

Pleurothallis sergioi Luer & R.Escobar, Orquideología 22(3): 272. 2003. Acronia sergioi (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 191. 2005.

Illustration: Luer 2003c, p. 274; Luer 2005, f. 184 (p. 260).

Distribution: *COL.

Type: Colombia, Luer 2958 (holotype, SEL).

Pleurothallis semiscabra Lindl., Orchid. Linden. 2. 1846. Acronia semiscabra (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 190. 2005. Humboldtia semiscabra (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Zosterophyllanthos semiscaber (Lindl.) Szlach. & Kulak, Richardiana 6: 193. 2006.

Illustration: Luer 2005, f. 183 (p. 259).

Distribution: COL VEN.

Type: Venezuela, Linden 669 (holotype, K-L; isotypes, G, W).

Voucher: Colombia, Ortiz-Valdivieso 131 (HPUJ).

Pleurothallis serpens Luer & R.Escobar, Orquideología 16(1): 31. 1983.

Illustration: Luer & Escobar 1983, p. 33; Luer 1999a, f. 79.

Distribution: *COL.

Type: Colombia, Luer et al. 7625 (holotype, SEL; isotypes, COL, JAUM).

Pleurothallis serricardia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 55. 1924. Acronia serricardia (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 191. 2005. Zosterophyllanthos serricardia (Schltr.) Szlach. & Kulak, Richardiana 6: 193. 2006.

Illustration: Luer 2005, f. 185 (p. 260).

Distribution: *COL.

Type: Colombia, Hopp 172 (holotype, B, destroyed).

Pleurothallis sextonii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 108-109. 2002. Acronia sextonii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 2002a, f. 20 (p. 119).

Distribution: *COL.

Type: Colombia, Luer 20050 (holotype, MO).

Pleurothallis sigynes Luer, Selbyana 3(1,2): 186. 1976. Acronia sigynes (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 191. 2005. Zosterophyllanthos sigynes (Luer) Szlach. & Kulak, Richardiana 6: 193. 2006.

Illustration: Luer 2005, f. 186 (p. 260).

Distribution: COL ECU.

Type: Ecuador, Luer & Kent 295 (holotype, SEL).

Voucher: Colombia, Escobar 8303 (MO).

Pleurothallis silverstoneiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 44: 122. 1992. Colombiana silverstonei (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 249. 2006.

Illustration: Luer 1992, pl. 8 (p. 123).

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 2667 (holotype, CUVC; isotype, MO).

Pleurothallis siphoglossa Luer & R.Escobar, Orquideología 20: 82. 1996. Acronia siphoglossa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 192. 2005.

Illustration: Luer & Escobar 1996, p. 83, p. 89; Luer 2005, f. 187 (p. 260).

Distribution: COL ECU VEN.

Type: Colombia, Luer et al. 10104 (holotype, MO).

Pleurothallis sirene Rchb.f., Bonplandia (Hannover) 3: 71. 1855. Humboldtia sirene (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Lindleyalis sirene (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 258. 2004.

Illustration: Luer 2000a, f. 6 (p. 114).

Distribution: COL VEN.

Type: Colombia, Wagener 558 (holotype, W; isotype, K-L).

Pleurothallis solaris K.W.Holcomb, Pleurothallidinae 1(6): [1]. 2022.

Illustration: Holcomb 2022a, p. [2]. p. [3]

Distribution: *COL.

Type: Colombia? [uncertain origin], Holcomb s.n. (holotype, GEO-18283)

Note: Described from a cultivated specimen from uncertain origin, but an in situ photograph is presented.

Pleurothallis somnolenta Luer, Orquideología 20: 225. 1996. Acronia somnolenta (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1996f, 226; Luer 1998b, f. 46.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4334 (holotype, CUCV; isotype, MO).

Pleurothallis sotarae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 115. 1920. Acronia sotarae (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 193. 2005. Zosterophyllanthos sotarae (Schltr.) Szlach. & Kulak, Richardiana 6: 193. 2006.

Illustration: Luer 2005, f. 189 (p. 261).

Distribution: *COL.

Type: Colombia, Lehmann 6219 [holotype, B, destroyed; lectotype, K; isolectotype, AMES (lLuer 2005)].

Pleurothallis spathulipetala Luer, Orquideología 20(2): 227. 1996. Ancipitia spathulipetala (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004.

Illustration: Luer 1996f, 228.

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K).

Pleurothallis stricta Luer, Selbyana 5(2): 181. 1979. Acronia stricta (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.20.

Distribution: COL ECU.

Type: Ecuador, Luer 4113 (holotype, SEL).

Voucher: Colombia, Luer 2937 (SEL).

Pleurothallis strobilifera F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 440. 1898.

Illustration: Luer 1999a, f. 81.

Distribution: *COL.

Type: Colombia, Lehmann 8199 [(holotype, B, destroyed; lectotype, K; isolectotypes, AMES, K (Luer 1999a)].

Pleurothallis subtilis C.Schweinf., Fieldiana, Bot. 28(1): 188. 1951. Acronia subtilis (C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 196. 2005. Zosterophyllanthos subtilis (C.Schweinf.) Szlach. & Marg., Polish Bot. J. 46: 121. 2001.

Illustration: Luer 2005, f. 194 (p. 262).

Distribution: COL VEN.

Type: Venezuela, Steyermark 56030 (holotype, F; isotypes, AMES, NY, VEN).

Voucher: Colombia, Escobar 821 (AMES).

Pleurothallis supervacanea Rchb.f., Hamburger GartenBlumenzeitung 16: 180. 1860. Acronia supervacanea (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1981a, p. 133; Luer 1998b, f. 21.

Distribution: *COL.

Type: Colombia, Patin? sub Schiller s.n. (holotype, W-R-0045922?; tracing of type, W-R-0053809!).

Note: Luer (1998b) cites the type collection as being made by Patin near Medellín. It is unclear to us how this association was made, the sheets we inspected have no annotation and the protologue cites the material as being cultivated by Stange in the collection of Schiller.

Pleurothallis talpinaria Rchb.f., Flora 69: 554. 1886. Talpinaria bivalvis H.Karst., Fl. Columb. 1: 153, t. 76. 1859.

Illustration: Luer 1998a, f. 4 (p. 100); Wilson et al. 2017b, f. 1-4, 9, 10, 12.

Distribution: COL VEN.

Type: Colombia, Karsten 6 (holotype, W-0075627! isotype, W-R-0012220).

Pleurothallis tanyrhina Luer & R.Escobar, Orquideología 14(2): 176. 1981.

Illustration: Luer & Escobar 1981a, p. 177; Luer 1999a, f. 45.

Distribution: *COL.

Type: Colombia, Luer 2082 (holotype, SEL; isotype, JAUM).

Pleurothallis taurus Luer, Phytologia 47(2): 80. 1980.

Illustration: Luer 1999a, f. 33.

Distribution: COL ECU.

Type: Colombia, Luer et al. 3077 (holotype, SEL).

Pleurothallis tenuisepala Mark Wilson, Lankesteriana 19(2): 82-83. 2019.

Illustration: Wilson et al. 2019, f. 2a, 3a, 4a, 5a, 6-9.

Distribution: *COL.

Type: Colombia, Wilson & Niessen PL0963 (holotype, COCO-spirit).

Note: This species substitutes the concept of P. luctuosa in Colombia.

Pleurothallis tetragona Luer & R.Escobar, Orquideología 14(2): 178. 1981. Ancipitia tetragona (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 255. 2004.

Illustration: Luer & Escobar 1981a, p. 179; Luer 1989, pl. 21; Wilson et al. 2017d, f. 7, f. 8b.

Distribution: *COL.

Type: Colombia, Escobar 983 (holotype, JAUM).

Pleurothallis tetroxysLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 31: 66. 1989. Colombiana tetroxys (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer 1989, pl. 6 (p. 67); Wilson et al. 2018a, f. 1f.

Distribution: *COL.

Type: Colombia, Luer 12497 (holotype, MO).

Pleurothallis thoerleae (Luer) J.M.H. Shaw, Orchid Review Suppl., 124(1314): 38. 2016. Acronia thoerleae Luer, Harvard Pap. Bot. 17(2): 337. 2012.

Illustration: Luer & Thoerle 2012b, f. 6.

Distribution: COL ECU.

Type: Ecuador, Thoerle 181 (holotype, MO; isotype, AMES).

Voucher: Colombia, Vieira 389 (JAUM).

Pleurothallis titan Luer, Selbyana 3(3-4): 400. 1977. Acronia titan (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 198, f. 199. 2005.

Illustration: Wilson et al. 2018b, f. 1n; Luer 2005, f. 199 (p. 263); Sierra-Ariza et al. 2022, f. 6.

Distribution: COL PAN.

Type: Panama, Dressler s.n. (holotype, SEL).

Voucher: Colombia, Paz et al. 643 (COL, MO).

Pleurothallis torrana Luer, Orquideología 20(2): 231-233. 1996.

Illustration: Luer 1996f, 232; Luer 1999a, f. 83; Wilson et al. 2017e, f. 2.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4334 (holotype, CUVC; isotype, MO).

Pleurothallis tridentata Klotzsch, Allg. Gartenzeitung 8: 289. 1840. Acronia tridentata (Klotzsch) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 200. 2005. Humboldtia tridentata (Klotzsch) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Zosterophyllanthos tridentatus (Klotzsch) Szlach. & Marg., Polish Bot. J. 46: 121. 2001.

Illustration: Wilson et al. 2018b, f. 1o; Luer 2005, f. 203a (p. 264), f. 203b, f. 203c (p. 265).

Distribution: COL ECU VEN.

Type: Venezuela, Otto s.n. (holotype, K).

Voucher: Forero et al. 6820 (COL, MO).

Pleurothallis trifurcata Luer & Hirtz, Lindleyana 11(3): 190. 1996.

Illustration: Luer 1999a, f. 36; Luer 1996c, f. 62.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11140 (holotype, MO).

Voucher: Colombia, Mora 3091 (AMES, PSO).

Pleurothallis troglodytes Luer, Selbyana 7: 125-126. 1982. Acronia troglodytes (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 201. 2005. Zosterophyllanthos troglodytes (Luer) Szlach. & Kulak, Richardiana 6: 194. 2006.

Illustration: Luer 2005, f. 204 (p. 265).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 7396 (holotype, SEL).

Voucher: Colombia, Dodson 18055 (MO).

Pleurothallis truncata Lindl., Companion Bot. Mag. 2: 355. 1836. Acronia truncata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 115: 257. 2009. Humboldtia truncata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Orbis truncata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 308. 2005.

Illustration: Luer 1975a, f. 21 (p. 99); Luer 1998b, f.2.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Barbosa CB8634 (COL).

Pleurothallis tryssa Luer, Selbyana 1(3): 303. 1975. Acronia tryssa Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1975a, f. 11 (p. 79), Luer 1998b, f.22.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 223 (holotype, SEL).

Voucher: Colombia, Luer 12500 (SEL).

Pleurothallis tuzae Luer, Orquideología 21(2): 132-134. 1999.

Illustration: Luer 1999b, p. 133; Luer 1999a, f. 37.

Distribution: *COL.

Type: Colombia, Lehmann s.n. H.K.60 (holotype, K).

Pleurothallis undulata Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 48. 1836. Acronia undulata (Poepp. & Endl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 203. 2005. Humboldtia undulata (Poepp. & Endl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Zosterophyllanthos undulatus (Poepp. & Endl.) Szlach. & Marg., Polish Bot. J. 46: 121. 2001.

Illustration: Luer 2005, f. 207 (p. 266).

Distribution: COL PER.

Type: Peru, Poeppig 1431 (holotype, W).

Voucher: Colombia, Díaz-Piedrahita 1016 (COL).

Pleurothallis urceolata Luer, Phytologia 46(6): 373. 1980.

Illustration: Luer 1999a, f. 38.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3228 (holotype, SEL).

Voucher: Colombia, Betancur 14856 (COL).

Pleurothallis uviferaLuer & R.Escobar, Orquideología 16(1): 36. 1983.

Illustration: Luer & Escobar 1983, p. 37; Luer 1999a, f. 84.

Distribution: *COL.

Type: Colombia, Luer et al. 6634 (holotype, SEL).

Pleurothallis variabilis Luer, Selbyana 5(2): 187. 1979. Acronia variabilis (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 204. 2005.

Illustration: Luer 2005, f. 210 (p. 267).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3909 (holotype, SEL).

Voucher: Colombia, Dodson & Escobar 18108 (MO).

Pleurothallis vieirae Luer & R.Escobar, Orquideología 14(2): 182. 1981. Acronia vieirae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer & Escobar 1981a, p. 183; Luer 1998b, f. 23; Misas Urreta 2005, p. 582, p. 583.

Distribution: *COL.

Type: Colombia, Luer 2037 (holotype, SEL; isotype, JAUM).

Pleurothallis villahermosaeSierra-Ariza, Rincón-González & Villanueva, Lankesteriana 22(1): 28. 2022.

Illustration: Sierra-Ariza et al. 2022, f. 1c, f. 2, f. 3.

Distribution: *COL.

Type: Colombia, Sierra-Ariza 54 (holotype, TOLI; isotype, JBB).

Pleurothallis vinealis Luer & R.Escobar, Orquideología 16(3): 30. 1986. Acronia vinealis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 205. 2005. Zosterophyllanthos vinealis (Luer & R.Escobar) Szlach. & Marg., Polish Bot. J. 46: 121. 2001.

Illustration: Luer & Escobar 1986, p. 31; Luer 2005, f. 211 (p. 267).

Distribution: COL ECU.

Type: Colombia, Luer & Escobar 10364 (holotype, MO).

Pleurothallis volcanica Luer, Selbyana 3(3-4): 406-407. 1977. Acronia volcanica (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 58. 2005.

Illustration: Luer 1998b, f.24.

Distribution: COL COS PAN.

Type: Panama, Luer & Butcher 1260 (holotype, SEL).

Voucher: Colombia, Hammel et al. 16562 (MO).

Note: A specimen collected exactly on the Panama/ Colombia border was cited in Kolanowska (2014).

Pleurothallis xanthochlora Rchb.f., Linnaea 22: 823. 1849. Humboldtia xanthochlora (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 1999a, f. 47; Kolanowska & Szlachetko 2020b, f. 4; Karremans et al. 2021, f. 3.

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Moritz 1054 (holotype, W; isotypes, AMES, BM-000082375, K-L-000584197).

Voucher: Colombia, Schneider 44 (AMES, S).

Pleurothallopsis Porto & Brade

Pleurothallopsis clausa (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Restrepiella clausa (Luer & R.Escobar) Braas & H.Mohr, Orchideen 33: 154. 1982. Restrepiopsis clausa Luer & R.Escobar, Selbyana 2: 202. 1978.

Illustration: Luer 1991a, pl.2 (p.93).

Distribution: COL ECU.

Type: Colombia, Luer 2330 (holotype, JAUM; isotype, SEL).

Pleurothallopsis inaequalis (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Restrepiopsis inaequalis Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 94. 1991.

Illustration: Luer 1991a, pl.3 (p.95).

Distribution: COL ECU.

Type: Ecuador, Luer 13211 (holotype, MO).

Voucher: Colombia, Luer 2042 (SEL).

Pleurothallopsis insons (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Restrepiopsis insons Luer & R.Escobar, Selbyana 7: 77. 1982.

Illustration: Luer 1991a, pl.4 (p.97).

Distribution: *COL.

Type: Colombia, Luer 7437 (holotype, SEL).

Pleurothallopsis lehmannii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001.

Restrepiopsis lehmannii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 98. 1991.

Illustration: Luer 1991a, pl.5 (p.99).

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K; H.K. 377).

Pleurothallopsis microptera (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001.

Pleurothallis microptera Schltr., Repert. Spec. Nov. Regni Veg. 7: 108. 1920. Restrepiella microptera (Schltr.) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 4: 266, t.. 1966. Restrepiopsis microptera (Schltr.) Luer, Selbyana 2: 200. 1978.

Illustration: Luer 1991a, pl.6 (p.101); Mansfeld 1929, pl. 37 (n. 142).

Distribution: COL ECU.

Type: Colombia, Lehmann 6922 [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 37, No. 142)].

Pleurothallopsis niesseniae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 154. 2010. Restrepiopsis niesseniae Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 79: 130, f. 1. 2000.

Illustration: Luer 2000a, f. 1 (p. 139L).

Distribution: *COL.

Type: Colombia, Niessen & Uribe 114 (holotype, MO).

Pleurothallopsis norae (Garay & Dunst.) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001.

Restrepiella norae Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 388. 1976. Restrepiopsis norae (Garay & Dunst.) Luer, Selbyana 2: 200. 1978.

Illustration: Luer 1991a, pl. 9 (p. 107).

Distribution: COL VEN.

Type: Venezuela, Dunsterville 1232 (holotype, AMES).

Voucher: Colombia, Luer 2332 (SEL, JAUM).

Pleurothallopsis powersii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Restrepiella powersii (Luer) Braas & H.Mohr, Orchideen 33: 154. 1982. Restrepiopsis powersii Luer, Selbyana 2: 204. 1978.

Illustration: Luer 1991a, pl. 11 (p.111).

Distribution: *COL.

Type: Colombia, Powers s.n. (holotype, SEL).

Pleurothallopsis rinkeiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 146, f. 18. 2010.

Illustration: Luer 2010, f. 18 (p. 153).

Distribution: *COL.

Type: Colombia, Rinke s.n. (holotype, MO).

Pleurothallopsis striata (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Restrepiopsis striata Luer & R.Escobar, Orquideología 16: 43. 1983.

Illustration: Luer & Escobar 1983, p. 44; Luer 1991a, pl. 13 (p.115).

Distribution: COL VEN.

Type: Colombia, Luer et al. 7734 (holotype, SEL).

Pleurothallopsis tubulosa (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Pleurothallis tubulosa Lindl., Fol. Orchid. ~Pleurothallis~ 19. 1859. Humboldtia tubulosa (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Restrepiella tubulosa (Lindl.) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 4: 266-267. 1966. Restrepiopsis tubulosa (Lindl.) Luer, Selbyana 2(2-3): 200. 1978.

Illustration: Luer 1991a, pl. 14 (p.117). Fig. 13F.

Distribution: COL COS ECU PAN VEN.

Type: Ecuador, Jameson s.n. [lectotype, K-000079816 (selected here). Ecuador, Jameson s.n. (syntype, K-000079818)].).

Voucher: Colombia, Lehmann 8226 (K).

Note: Although not immediately apparent, Lindley cites two materials by Jameson “Valley of Lloa, on trees” and “forests at 12,000” and both specimens are indeed present in his herbarium. The first is selected as lectotype.

Pleurothallopsis ujarrensis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 255. 2001. Humboldtia ujarrensis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1859. Pleurothallis ujarrensis (Rchb.f.) Lindl., Fol. Orchid. ~Pleurothallis~ 19. 1859. Restrepia ujarrensis Rchb.f., Bonplandia (Hannover): 225. 1855. Restrepiella ujarrensis (Rchb.f.) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 4: 266. 1966. Restrepiopsis ujarrensis (Rchb.f.) Luer, Selbyana 2(2-3): 200. 1978.

Illustration: Luer 1991a, pl. 15 (p.120).

Distribution: COL COS GUA NIC PAN.

Type: Costa Rica, Oersted s.n. [607] (holotype, W-R-0029376; isotype, K-L-000079867).

Voucher: Colombia, Escobar 2579 (COL).

Porroglossum Schltr.

Porroglossum actrix Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 152. 1991.

Illustration: Karremans & Vieira-Uribe 2020, p. 245.

Distribution: *COL.

Type: Colombia, Escobar 4050 (holotype, MO; isotype, JAUM).

Porroglossum agile Luer, Phytologia 46(6): 374.

Illustration: Luer 1987, pl. 1 (35); Kolanowska et al. 2013, p. 79; Karremans & Vieira-Uribe 2020, p. 235.

Distribution: COL PER.

Type: Peru, Luer 5254 (holotype, SEL).

Voucher: Colombia, Medina 150 (MEDEL).

Porroglossum echidna (Rchb.f.) Garay, Svensk Bot. Tidskr. 47: 201. 1953. Masdevallia echidna Rchb.f., Bonplandia (Hannover) 3: 69. 1855. Scaphosepalum echidna (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 219. 1920.

Illustration: Luer 1987, pl. 7 (47); Karremans & Vieira-Uribe 2020, p. 239.

Distribution: *COL.

Type: Colombia, Wagener s.n. (holotype, W-R0050793).

Porroglossum eduardii (Rchb.f.) Sweet, Amer. Orchid Soc. Bull. 41: 519. 1972. Masdevallia eduardii Rchb.f., Gard. Chron. n.s., 14: 778. 1880.

Illustration: Luer 1987, pl. 8; Karremans & Vieira-Uribe 2020, p. 226.

Distribution: COL ECU.

Type: Colombia, Klaboch s.n. (holotype, W-R0047205).

Porroglossum medinae Kolan. & Szlach., Polish Bot. J. 58(2): 629. 2013.

Illustration: Karremans & Vieira-Uribe 2020, p. 235.

Distribution: *COL.

Type: Colombia, Medina 577 (holotype, MEDEL).

Porroglossum mordax (Rchb.f.) H.R.Sweet, Orquideología 5: 166. 1970. Masdevallia mordax Rchb.f., Flora 69: 559. 1886. Lothiania mordax (Rchb.f.) Kraenzl., Gard. Chron., ser. 3 75: 173. 1924.

Illustration: Luer 1987, pl. 13; Karremans & Vieira-Uribe 2020, p. 228. Fig. 13G.

Distribution: *COL.

Type: Colombia, Lehmann 3620 (holotype, W-R0047205).

Porroglossum muscosum (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 83. 1920. Masdevallia muscosa Rchb.f., Gard. Chron. n.s., 3: 460. 1875.

Illustration: Luer 1987, pl. 14, pl. 15, pl. 17; Karremans & Vieira-Uribe 2020, p. 242.

Distribution: COL ECU VEN.

Type: Colombia, Shuttleworth s.n. (holotype, W-R0050782; isotype, K).

Porroglossum nutibara Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 24: 66, pl. 18. 1987.

Illustration: Luer 1987, pl. 18; Karremans & Vieira-Uribe 2020, p. 243.

Distribution: *COL.

Type: Colombia, Escobar s.n. (holotype, JAUM; isotype, MO).

Porroglossum olivaceum Sweet, Orquideología 5: 167. 1970.

Illustration: Luer 1987, pl. 19-21; Karremans & Vieira-Uribe 2020, p. 245.

Distribution: COL ECU.

Type: Ecuador, Drew E-536 (holotype, AMES).

Voucher: Colombia, Luer 4616 (SEL).

Porroglossum parsonsiiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 253-254. 2006.

Illustration: Luer 2006, f. 12. Fig. 13H; Karremans & Vieira-Uribe 2020, p. 240.

Distribution: *COL.

Type: Colombia, Luer 20985 (holotype, MO).

Porroglossum rodrigoi H.R.Sweet, Orquideología 9: 129. 1974.

Illustration: Luer 1987, pl. 25; Karremans & Vieira-Uribe 2020, p. 244.

Distribution: *COL.

Type: Colombia, Escobar 1055 (holotype, AMES).

Porroglossum sergioi P.Ortiz, Orquideología 10: 215. 1975.

Illustration: Luer 1987, pl. 27; Karremans & Vieira-Uribe 2020, p. 243.

Distribution: *COL.

Type: Colombia, Ortiz 689 (holotype, COL).

Note: Originally described as P. sergii.

Porroglossum tripollex Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 72: 114-115. 1998.

Illustration: Luer 1998, f. 8 (p. 119); Karremans & Vieira-Uribe 2020, p. 230.

Distribution: COL ECU.

Type: Ecuador, Teague 271 (holotype, MO).

Voucher: Colombia, Ortiz 1390 (HPUJ).

Pseudolepanthes (Luer) Archila

Pseudolepanthes bihuae M.F.Monteros & Baquero, Lankesteriana 21(1): 5-6. 2021.

Illustration: Karremans & Vieira-Uribe 2020, p. 254 (top right); Monteros et al. 2021, f. 1, f. 2. Fig. 13I.

Distribution: COL ECU.

Type: Ecuador, Monteros 203 (holotype, QCNE).

Voucher: Colombia, Vieira 253 (JAUM).

Pseudolepanthes bricenoensis (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 78. 2000 [2009]. Pseudolepanthes bricenoensis (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 65. 2014 [2015], nom. superfl. Trichosalpinx bricenoensis Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 105. 1997.

Illustration: Luer 1997, f. 93; Karremans & VieiraUribe 2020, p. 253. Fig. 13J.

Distribution: *COL.

Type: Colombia, Luer et al. 14196 (holotype, MO; isotype, JAUM).

Pseudolepanthes calceolaris (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 78. 2000 [2009]. Pseudolepanthes calceolaris (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 65. 2014 [2015], nom. superfl. Trichosalpinx calceolaris Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 105-106. 1997.

Illustration: Luer 1997, f. 94; Karremans & Vieira-Uribe 2020, p. 255, p. 256.

Distribution: *COL.

Type: Colombia, Luer 16723 (holotype, MO).

Pseudolepanthes colombiae Archila, Revista Guatemal. 3(1): 79. 2000 [2009]. Pseudolepanthes colombiae Archila, Revista Guatemal. 17(1): 66. 2014 [2015], nom. superfl. Trichosalpinx pseudolepanthes Luer & R.Escobar, Orquideología 16(2): 183. 1984.

Illustration: Luer & Escobar 1984b, p. 184, p. 185; Luer 1997, f. 97; Karremans & Vieira-Uribe 2020, p. 252.

Distribution: *COL.

Type: Colombia, Escobar & Valencia 2623 (holotype, SEL).

Pseudolepanthes decorata (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 78. 2000 [2009].

Pseudolepanthes decorata (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 65. 2014 [2015], nom. superfl. Trichosalpinx decorata Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 106. 1997.

Illustration: Luer 1997, f. 95.

Distribution: *COL.

Type: Colombia, Luer 16738 (holotype, MO).

Pseudolepanthes ectopa (Luer & R.Escobar) Archila, Revista Guatemal. 3(1): 79. 2000 [2009].

Pseudolepanthes ectopa (Luer & R.Escobar) Archila, Revista Guatemal. 17(1): 65. 2014 [2015], nom. superfl. Trichosalpinx ectopa Luer, Orquideología 18(2): 166-168. 1991. Trichosalpinx ectopa Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 106. 1997. nom. superfl.

Illustration: Luer 1991f, p. 167; Luer 1997, f. 96; Karremans & Vieira-Uribe 2020, p. 257.

Distribution: *COL.

Type: Colombia, Luer et al. 14202 (holotype, MO).

Pseudolepanthes ramosii (Luer) Archila, Revista Guatemal. 3(1): 79. 2000 [2009].

Pseudolepanthes ramosii (Luer) Archila, Revista Guatemal. 17(1): 66. 2014 [2015], nom. superfl. Trichosalpinx ramosii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 108. 1997.

Illustration: Luer 1997, f. 98.

Distribution: *COL.

Type: Colombia, Ramos et al. 1306 (holotype, CUVC; isotype, MO).

Pseudolepanthes silverstonei (Luer) Archila, Revista Guatemal. 3(1): 79. 2000 [2009]. Pseudolepanthes silverstonei (Luer) Archila, Revista Guatemal. 17(1): 66. 2014 [2015], nom. superfl. Trichosalpinx silverstonei Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 108-109. 1997.

Illustration: Luer 1997, f. 99; Karremans & Vieira-Uribe 2020, p. 254.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4324 (holotype, CUVC; isotype, MO).

Pseudolepanthes spathulata (Luer) Archila, Revista Guatemal. 3(1): 79. 2000 [2009]. Pseudolepanthes spathulata (Luer) Archila, Revista Guatemal. 17(1): 67. 2014 [2015], nom. superfl. Trichosalpinx spathulata Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 109. 1997.

Illustration: Luer 1997, f. 100; Karremans & Vieira-Uribe 2020, p. 258.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4323 (holotype, CUVC; isotype, MO).

Pseudolepanthes uvaria (Luer) Archila, Revista Guatemal. 3(1): 80. 2000 [2009]. Pseudolepanthes uvaria (Luer) Archila, Revista Guatemal. 17(1): 67. 2014 [2015], nom. superfl. Trichosalpinx uvaria Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 109-110. 1997.

Illustration: Luer 1997, f. 101; Karremans & Vieira-Uribe 2020, p. 251.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4322 (holotype, CUVC; isotype, MO).

Restrepia Kunth

Restrepia antennifera Kunth, Nov. Gen. Sp. (quarto ed.) 1: 367, pl. 94. 1816.

Illustration: Luer 1996a, pl. 2, pl. 3.

Distribution: COL ECU VEN.

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Restrepia aristulifera Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 5: 258. 1972.

Illustration: Luer 1996a, pl. 4; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Venezuela, Dunsterville 1090 (holotype, AMES).

Voucher: Colombia, Killip & Smith 20313, 20318 (AMES, NY).

Restrepia aspasicensis Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Illustration: Luer 1996a, pl. 5; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Colombia, Wagener s.n. (holotype, W; isotype, K).

Restrepia brachypus Rchb.f., Flora 69: 554. 1886.

Illustration: Luer 1996a, pl. 6, pl. 7; Luer 1996e, p. 150 (top, bottom left); Ortiz & Uribe 2007 (CD Edition).

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Lehmann 3215 (holotype, W).

Restrepia chameleon Luer & R.Escobar, Orquideología 20: 120. 1996.

Illustration: Luer 1996a, pl. 8; Luer 1996e, p. 122, p. 153; Ortiz & Uribe 2007 (CD Edition). Fig. 13K.

Distribution: *COL.

Type: Colombia, Luer 13130 (holotype, JAUM; isotype, MO).

Restrepia chocoënsis Garay, Orquideología 8(3): 181. 1973.

Illustration: Luer 1996a, pl. 9; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Escobar 1054 (holotype, AMES).

Restrepia chrysoglossa Luer & R.Escobar, Orquideología 20: 123. 1996.

Illustration: Luer 1996a, pl. 10; Luer 1996e, p. 124, p. 154 (bottom left).

Distribution: *COL.

Type: Colombia, Luer 17874 (holotype, JAUM; isotype, MO).

Restrepia citrinaLuer & R.Escobar, Orquideología 16(1): 40. 1983.

Illustration: Luer & Escobar 1983, p. 41; Luer 1996a, pl. 11; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Luer et al. 6624 (holotype, SEL; isotype, JAUM).

Restrepia contorta (Ruiz & Pav.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 59: 50. 1996.

Humboldtia contorta Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 235. 1798. Stelis contorta (Ruiz & Pav.) Pers., Syn. Pl. 2: 524. 1807.

Illustration: Luer 1996a, pl. 14, pl. 15, pl. 16, pl. 17, pl. 18; ; Pupulin 2013, f. 38A.

Distribution: COL ECU PER VEN.

Type: Peru, Ruiz & Pavón s.n. (holotype, MA; isotype, W).

Voucher: Colombia, Linden 1270 (K, BR, W).

Restrepia cuprea Luer & R.Escobar, Orquideología 20: 130. 1996.

Illustration: Luer 1996a, pl. 19; Luer 1996e, p. 131, p. 151 (bottom left); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Luer 16677 (holotype, JAUM; isotype, MO).

Restrepia echinata Luer & R.Escobar, Orquideología 20: 135. 1996.

Illustration: Luer 1996a, pl. 22; Luer 1996e, p. 137, p. 154 (top left); Ortiz & Uribe 2007 (CD Edition).

Distribution: COL PER.

Type: Colombia, Luer 16660 (holotype, JAUM; isotype, MO).

Restrepia echo Luer & R.Escobar, Orquideología 20: 138. 1996.

Illustration: Luer 1996a, pl. 23; Luer 1996e, p. 140, p. 155 (bottom left); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Escobar 6001 (holotype, JAUM; isotype, MO).

Note: According to Luer (1996a) the type is Luer 16702, but the protologue indicates Escobar 6001 (Luer 1996e).

Restrepia elegans H.Karst., Allg. Gartenzeitung 15: 202. 1847.

Illustration: Luer 1996a, pl. 24, pl. 25; Luer 1996e, p. 154 (top right); Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Venezuela, Karsten s.n. (lectotype, W; isolectotype, K).

Voucher: Colombia, Luer 14445 (MO).

Restrepia escobariana Luer, Orquideología 20: 144. 1996.

Illustration: Luer 1996a, pl. 27; Luer 1996e, p. 145, p. 149 (top left, bottom); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Luer 16656 (holotype, JAUM; isotype, MO).

Restrepia falkenbergii Rchb.f., Gard. Chron., n.s. 13: 232. 1880.

Illustration: Luer 1996a, pl. 28; Luer 1996e, p. 154 (bottom right); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Falkenberg s.n. (lectotype, W, illustration).

Restrepia flosculata Luer, Selbyana 7(1): 127. 1982.

Illustration: Luer 1996a, pl. 29.

Distribution: COL ECU.

Type: Ecuador, Dodson & Thien 1600 (holotype, SEL).

Voucher: Escobar s.n. (JAUM).

Restrepia fritillina Luer & V.N.M.Rao, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 110. 2007.

Illustration: Luer 2007, f. 5 (p. 114).

Distribution: *COL.

Type: Colombia, Rao 299 (holotype, MO).

Restrepia guttulata Lindl., Companion Bot. Mag. 2: 357. 1836.

Illustration: Luer 1996a, pl. 30, pl. 31, pl. 32, pl. 33.

Distribution: COL ECU PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Luer 5213 (SEL).

Restrepia limbata Luer & R.Escobar, Selbyana 7: 76. 1982.

Illustration: Luer 1996a, pl. 39.

Distribution: *COL.

Type: Colombia, Escobar 1187 (holotype, SEL; isotype, JAUM).

Restrepia metae Luer, Orquideología 20: 159. 1996.

Illustration: Luer 1996a, pl. 41; Luer 1996e, p. 161.

Distribution: *COL.

Type: Colombia, Renz 3610 (holotype, Herb. J. Renz).

Restrepia muscifera (Lindl.) Rchb.f. ex Lindl., Fol. Orchid. Restrepia 2. 1859. Pleurothallis muscifera Lindl., Edwards’s Bot. Reg. 28: Misc. 79. 1842.

Illustration: Luer 1996a, pl. 43, pl. 44, pl. 45; Luer 1996e, p. 155 (top left); Ortiz & Uribe 2007 (CD Edition).

Distribution: COL COS ECU ELS GUA HON MEX NIC PAN.

Type: Guatemala, Skinner s.n. (holotype, K).

Voucher: Colombia, Escobar 3328 (JAUM).

Restrepia nittiorhyncha (Lindl.) Garay, Bot. Mus. Leafl. 21: 253. 1967. Humboldtia nittiorhyncha (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis nittiorhyncha Lindl., Fol. Orchid. ~Pleurothallis~ 20. 1859.

Illustration: Luer 1996a, pl. 46; Luer 1996e, p. 151 (top left); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Schlim s.n. (holotype, K).

Restrepia pandurata Rchb.f., Gard. Chron., ser. 3 3: 244. 1888.

Illustration: Luer 1996a, pl. 47; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Wendland s.n. (holotype, W).

Restrepia pelyx Luer & R.Escobar, Selbyana 7(1): 76. 1982.

Illustration: Luer 1996a, pl. 48; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Colombia, Escobar 241 (holotype, SEL; isotype, JAUM).

Restrepia purpurea Luer & R.Escobar, Orquideología 20: 162. 1996.

Illustration: Luer 1996a, pl. 49; Luer 1996e, p. 150 (bottom right), p. 163.

Distribution: *COL.

Type: Colombia, Luer 17545 (holotype, JAUM; isotype, MO).

Restrepia sanguinea Rolfe, Bull. Misc. Inform. Kew 1896: 44. 1896.

Illustration: Luer 1996a, pl. 53; Ortiz & Uribe 2007 (CD Edition); Rincón-González & Vieira-Uribe 2019, f. 36.

Distribution: *COL.

Type: Colombia, Charlesworth & Co. s.n. (holotype, K; isotype, AMES-00103796).

Restrepia santanderensis N.Gut. & K.Gil-Amaya, Phytotaxa 598(4): 293-300. 2023.

Illustration: Gutiérrez Morales et al. 2023, f. 1, f. 2, f. 3c-d.

Distribution: *COL.

Type: Colombia, Calderón et al. 191 (holotype, JBB).

Restrepia seketii Luer & R.Escobar, Orquideología 20: 175. 1996.

Illustration: Luer 1996a, pl. 55; Luer 1996e, p. 156 (top left, bottom), p. 177.

Distribution: *COL.

Type: Colombia, Escobar 3325 (holotype, JAUM).

Note: According to Luer (1996a) the type is Luer 16658A, but the protologue indicates Escobar 3325 (Luer 1996e)

Restrepia tabeae H.Mohr, Leafl. Schlechter Inst. 2: 10, t. 1-2. 1996.

Illustration: Luer 1996a, pl. 56; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, unknown collector (holotype, Herb. Schltr. Instit; isotype, W).

Restrepia trichoglossa F.Lehm. ex Sander, Sander’s Orch. Guide 215. 1901.

Illustration: Luer 1996a, pl. 58, pl. 59, pl. 60; Luer 1996e, p. 152 (bottom).

Distribution: COL COS ECU GUA MEX PAN PER.

Type: Colombia, Lehmann B.T. 1312 (lectotype, K; isolectotypes, AMES, W).

Restrepia tsubotae Luer & R.Escobar, Orquideología 20: 178. 1996.

Illustration: Luer 1996a, pl. 61; Luer 1996e, p. 152 (top right), p. 179; Ortiz & Uribe 2007 (CD Edition). Fig. 13L.

Distribution: *COL.

Type: Colombia, Luer 17976 (holotype, JAUM; isotype, MO).

Restrepiella Garay & Dunst.

Restrepiella doucetteana Doucette ex L.Salm, OrchideenJ. 24(4): 172. 2017.

Illustration: Salm 2017, p. 174-175.

Distribution: COL.

Type: Colombia, Doucette 343 (holotype, WIS).

Scaphosepalum Pfitzer

Scaphosepalum antenniferum Rolfe, Gard. Chron., ser. 3 7: 709. 1890.

Illustration: Luer 1988, pl. 5 (p. 37).

Distribution: COL ECU PER.

Type: Colombia, Charlesworth, Shuttleworch & Co. s.n. (holotype, K; isotype, W).

Scaphosepalum bicolor Luer & R.Escobar, Orquideología 14: 186. 1981.

Illustration: Luer & Escobar 1981a, p. 187; Luer 1988, pl. 7 (p. 41).

Distribution: *COL.

Type: Colombia, Escobar 1650 (holotype, JAUM).

Scaphosepalum bicristatum Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 110. 2002.

Illustration: Luer 2002a, f. 23 (p. 120).

Distribution: *COL.

Type: Colombia, Hirtz 7629 (holotype, MO).

Scaphosepalum breve (Rchb.f.) Rolfe, J. Bot. 28: 136. 1890. Masdevallia brevis Rchb.f., Gard. Chron. 2: 588. 1883.

Illustration: Luer 1988, pl. 8-10 (p. 43-45).

Distribution: BOL COL ECU GUY VEN.

Type: Guyana, Veitch & Sons 444 (holotype, K; isotype, W).

Voucher: Colombia, Wallis 343 (W).

Scaphosepalum decorum Luer & R.Escobar, Orquideología 15: 149. 1982.

Illustration: Luer & Escobar 1982, p. 150; Luer 1988, pl. 14-15 (p. 53-54).

Distribution: *COL.

Type: Colombia, Luer 3459 (holotype, SEL; isotype, JAUM).

Scaphosepalum gibberosum (Rchb.f.) Rolfe, J. Bot. 28: 136. 1890. Masdevallia gibberosa Rchb.f., Gard. Chron. 1: 8. 1876.

Illustration: Luer 1988, pl. 18-19 (p. 60-61).

Distribution: *COL.

Type: Colombia, Wallis 387 (holotype, W; isotype, K).

Scaphosepalum grande Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 135. 1922.

Illustration: Luer 1988, pl. 20 (p. 63). Fig. 16A.

Distribution: *COL.

Type: Colombia, Kalbreyer 1846 [holotype, B, destroyed; lectotype, W (Luer 1988)].

Scaphosepalum lima (F.Lehm. & Kraenzl.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 220. 1920. Masdevallia lima F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 454. 1899.

Illustration: Luer 1988, pl. 22 (p. 67).

Distribution: *COL.

Type: Colombia, Lehmann 8238 (holotype, K; isotype, G).

Scaphosepalum macrodactylum (Rchb.f.) Rolfe, J. Bot. 28: 136. 1890. Masdevallia macrodactyla Rchb.f., Gard. Chron. 571. 1872.

Illustration: Luer 1988, pl. 23 (p. 69).

Distribution: COL ECU.

Type: Colombia, Veitch & Sons 202 (holotype, W).

Scaphosepalum microdactylum Rolfe, Bull. Misc. Inform. Kew 1893(82-83): 335. 1893 [1894].

Illustration: Luer 1988, pl. 24-28 (p. 69-76); Karremans et al. 2016, f. 4d. f. 6e, f. 8e.

Distribution: COL COS ECU GUA HON MEX NIC PAN.

Type: Unknown (holotype, K).

Voucher: Colombia, Luer 3461 (SEL).

Scaphosepalum odontochilum Kraenzl., Bull. Misc. Inform. Kew 113. 1925.

Illustration: Luer 1988, pl. 29 (p. 78), pl. 30 (p. 79). Fig. 16B.

Distribution: COL ECU.

Type: Colombia, Lehmann 172 (holotype, K; isotype, AMES).

Scaphosepalum ophidion Luer, Phytologia 49(3): 222-223. 1981.

Illustration: Luer 1988, pl. 31 (p. 81), Baquero 2017, fig. 2.

Distribution: COL ECU.

Type: Ecuador, Hirtz et al. 4381 (holotype, SEL).

Voucher: Colombia, Luer et al. 4607 (SEL).

Scaphosepalum panduratum Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 134. 1997.

Illustration: Luer 1997, f. 49 (p. 136).

Distribution: *COL.

Type: Colombia, Escobar 8226 (holotype, MO).

Scaphosepalum pulvinare (Rchb.f.) Rolfe, J. Bot. 28: 137. 1890. Masdevallia pulvinaris Rchb.f., Gard. Chron. 1: 200. 1880.

Illustration: Luer 1988, pl. 33 (p. 85).

Distribution: COL ECU.

Type: Colombia, Calvert s.n. (holotype, W).

Scaphosepalum swertiifolium (Rchb.f.) Rolfe, J. Bot. 28: 137. 1890. Masdevallia swertiifolia Rchb.f., Gard. Chron., n.s. 2: 390. 1880.

Illustration: Luer 1988, pl. 35-38 (p. 89-93). Fig. 16C.

Distribution: COL ECU.

Type: Colombia, Lehmann 544 (holotype, W).

Scaphosepalum tiaratum Luer, Phytologia 49: 223. 1981.

Illustration: Luer 1988, pl. 39 (p. 95).

Distribution: *COL.

Type: Colombia, Luer 5255 (holotype, SEL).

Scaphosepalum verrucosum (Rchb.f.) Pfitzer, Nat. Pflanzenfam. (ed. 2) 2(6): 139. 1888. Humboldtia verrucosa (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Masdevallia verrucosa Rchb.f., Linnaea 22: 819. 1849. Pleurothallis verrucosa (Rchb.f.) Rchb.f., Bonplandia (Hannover) 2: 24. 1854.

Illustration: Luer 1988, pl. 42 (p. 101).

Distribution: *COL.

Type: Colombia, Schlim 1439 (holotype, W; isotypes, G, P).

Specklinia Lindl.

Specklinia acrisepala (Ames & C.Schweinf.) Pridgeon & M.W.Chase, Lindleyana 16(4): 256. 2001. Pleurothallis acrisepala Ames & C.Schweinf., Sched. Orch. 8: 22-23. 1925. Sarcinula acrisepala (Ames & C.Schweinf.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 203.

Illustration: Luer 2006, f. 156; Misas Urreta 2005, p. 510, p. 511; Karremans et al. 2016, f. 6g; Karremans & Vieira-Uribe 2020, p. 293, p. 300; Karremans et al. 2020, f. 1.

Distribution: COL COS ECU HON PAN.

Type: Panama, Powell 420 (holotype, AMES).

Voucher: Colombia, Lellinger & de la Sota 17 (US).

Specklinia areldii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 256. 2001. Pleurothallis areldii Luer, Selbyana 2(4): 383-384. 1978. Sarcinula areldii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 204-205, f. 158. 2006.

Illustration: Luer 2006, f. 158; Misas Urreta 2005, p. 516, p. 517; Karremans & Vieira-Uribe 2020, p. 295.

Distribution: COL PAN.

Type: Panama, Luer 1653 (holotype, SEL).

Voucher: Colombia, Luer 18443 (MO).

Specklinia brighamii (S. Watson) Pridgeon & M.W.Chase, Lindleyana 16(4): 256. 2001. Pleurothallis brighamii S. Watson, Proc. Amer. Acad. Arts 23(2): 285-286. 1888. Sarcinula brighamii (S. Watson) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 206-207. 2006.

Illustration: Stenzel 2004, f. 6; Luer 2006, f. 161; Karremans et al. 2016, f. 6g; Karremans & VieiraUribe 2020, p. 294; Karremans et al. 2020, f. 4.

Distribution: BEL COL COS CUB ECU ELS GUA HAI HON JAM L.A. MEX NIC PAN.

Type: Guatemala, Watson s.n. (holotype, AMES).

Voucher: Colombia, Duque 2038 (JAUM).

Specklinia cactantha (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Pleurothallis cactantha Luer, Selbyana 3(1-2): 72-73. 1976. Sylphia cactantha (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 228. 2006.

Illustration: Luer 2006, f. 180; Karremans & Vieira-Uribe 2020, p. 295.

Distribution: COL PAN.

Type: Panama, Dressler 743 (holotype, SEL).

Voucher: Colombia, Luer et al. 16845 (MO).

Specklinia calderae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004. Pleurothallis calderae Luer, Orquideología 22(1): 53-56. 2001. Sarcinula calderae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 207-208. 2006.

Illustration: Luer 2001b, p. 55; Luer 2006, f. 162. Fig. 16D.

Distribution: *COL.

Type: Colombia, Luer 14273 (holotype, MO).

Specklinia calyptrostele (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Pleurothallis calyptrostele Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 23. 1923.

Illustration: Luer 2006, f. 2; Luer 1975e, f. 86; Misas Urreta 2005, p. 526, p. 527; Karremans et al. 2016, f. 7f; Karremans & Vieira-Uribe 2020, p. 290, p. 301. Fig. 16E.

Distribution: COL COS PAN.

Type: Costa Rica, Wercklé 148 [holotype, B, destroyed; lectotype, AMES (Luer 2006)].

Voucher: Colombia, Misas Urreta 305 (HPUJ).

Specklinia campylotyle (P.Ortiz) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Acostaea campylotyle P.Ortiz, Orquideología 13(3): 240, 246. 1979.

Illustration: Ortiz 1979b, p. 241, p. 243 (top, left); Luer 1987, pl. 1; Vieira-Uribe & Karremans 2018a, f. 2b; Karremans & Vieira-Uribe 2020, p. 280.

Distribution: *COL.

Type: Colombia, Ortiz & Restrepo 247 (holotype, COL).

Specklinia caulophryne S.Vieira-Uribe & Karremans, Orquideología 35(2): 165. 2018.

Illustration: Vieira-Uribe & Karremans 2018a, f. 1, f. 2a, f. 3; Karremans & Vieira-Uribe 2020, p. 279.

Distribution: *COL.

Type: Colombia, Vieira et al. 26 (holotype, JAUM).

Specklinia colombiana (Garay) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Acostaea colombiana Garay, Orquideología 9: 112. 1974. Acostaea costaricensis subsp. colombiana (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 24: 10. 1988.

Illustration: Luer 1975d, f. 42; Ortiz 1982, p. 165, p. 167; Luer 1987, pl. 3, pl. 4; Karremans et al. 2016, f. 4f, f. 7i; Karremans & Vieira-Uribe 2020, p. 277, p. 278, p. 300.

Distribution: COL COS ECU PAN.

Type: Colombia, Arango s.n. (holotype, AMES).

Specklinia corniculata (Sw.) Steud., Nomencl. Bot. (ed. 2) 1: 489. 1840. Sarcinula corniculata (Sw.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 209-211, f. 165. 2006. Cymbidium corniculatum (Sw.) Spreng., Syst. Veg. 3: 722. 1826. Dendrobium corniculatum (Sw.) Sw., Nova Acta Regiae Soc. Sci. Upsal. 6: 83. 1799. Epidendrum corniculatum Sw., Prodr. Fl. Ind. Orient. 1: 123. 1788. Humboldtia corniculata (Sw.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis corniculata (Sw.) Lindl., Edwards’s Bot. Reg. 28: Misc. 83. 1842.

Illustration: Stenzel 2004, f. 8; Luer 2006, f. 165; Karremans & Vieira-Uribe 2020, p. 272.

Distribution: BEL COL COS CUB FRE GUA GUY HAI HON JAM NIC PAN VEN.

Type: Jamaica, Swartz s.n. (holotype, S-R-1935; isotypes, BM [not seen], LD-1732006, M-0226530, UPS [not seen], W-R-0026609).

Voucher: Colombia, Monsalve 1003 (CUVC).

Note: Kolanowska (2014) presumably selected an isotype at W as lectotype, without specifying nor annotating any particular sheet or specimen. Regardless, it is debatable if this taxon truly requires a lectotype given that there is an annotated specimen in Swartz’s herbarium at S. The latter, in any case, would certainly have priority as lectotype.

Specklinia coronula (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004. Pleurothallis coronula Luer, Orquideología 21(3): 325-329.. 2000, nom. superfl. Pleurothallis coronula Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 171. 1999. Sarcinula coronula (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 211. 2006.

Illustration: Luer 1999a, f. 23a; Luer 2000d, p. 326; Luer 2006, f. 166; Karremans & Vieira-Uribe 2020, p. 282.

Distribution: COL ECU.

Type: Colombia, Luer 18824 (holotype, MO).

Specklinia costaricensis (Rolfe) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Pleurothallis costaricensis Rolfe, Bull. Misc. Inform. Kew 1917(2): 80. 1917.

Illustration: Luer 2006, f. 4; Karremans et al. 2016, f. 7g; Karremans & Vieira-Uribe 2020, p. 285.

Distribution: COL COS ECU PAN.

Type: Costa Rica, Lankester s.n. [Lectotype, K; (Luer 2006)]. Costa Rica, Lankester 21 [Lectotype, K-000463188, Luer (2023)].

Voucher: Colombia, Lehmann 7080 (AMES, W).

Note: A lectotype for this taxon was initially selected by Luer (2006), but it not clear which of the many specimens of this species collected by Lankester conserved at Kew was actually meant. One was afterwards specified by Luer (2023).

Specklinia cycesis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004. Pleurothallis cycesis Luer & R.Escobar, Orquideología 20: 49. 1996. Sarcinula cycesis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 212. 2006.

Illustration: Luer & Escobar 1996, p. 51; Luer 2006, f. 167; Misas Urreta 2005, p. 530, p. 531; Karremans & Vieira-Uribe 2020, p. 282.

Distribution: *COL.

Type: Colombia, Luer 14290 (holotype, MO).

Specklinia dunstervillei Karremans, Pupulin & Gravend., PLoS ONE 10(7): e13197(5). 2015.

Illustration: Karremans et al. 2015a, f. 1A-B, f. 3-8; Luer 1986, pl. 18; Misas Urreta 2005, p. 534, p. 535; Karremans et al. 2016, f. 4h; Karremans & Vieira-Uribe 2020, p. 268.

Distribution: COL COS PAN VEN.

Type: Venezuela, Karremans & Gravendeel 5966 (holotype, JBL-spirit; isotype, L-spirit).

Voucher: Colombia, Misas Urreta 291 (HPUJ).

Note: The Colombian material of this taxon have been previously misidentified as Specklinia endotrachys and S. remotiflora, neither is known to occur in the country.

Specklinia fulgens (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Humboldtia fulgens (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis fulgens Rchb.f., Gard. Chron., n.s. 4(95): 516. 1875. Sarcinula fulgens (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 213-214. 2006.

Illustration: Luer 2006, f. 170; Karremans et al. 2016, f. 6h; Karremans & Vieira-Uribe 2020, p. 271, p. 302.

Distribution: COL COS PAN.

Type: Costa Rica, Endrés 2512 [lectotype, W-R0025509/W-0020353 (Luer 2023)]

Voucher: Colombia, Juncosa 1692 (JAUM).

Specklinia glandulosa (Ames) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Pleurothallis glandulosa Ames, Sched. Orch. 6: 60-61. 1923. Sarcinula glandulosa (Ames) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 214-215. 2006.

Illustration: Luer 2006, f. 171; Karremans et al. 2015b, f. 1E, f. 2C, f. 9, f. 10, f. 18E; Karremans & Vieira-Uribe 2020, p. 264.

Distribution: BEL COL COS GUY HON PAN SUR.

Type: Panama, Powell 306 (holotype, AMES; isotype, MO).

Voucher: Colombia, Prance 28011 (COL).

Specklinia gracillima (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001. Humboldtia gracillima (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis gracillima Lindl., Fol. Orchid. ~Pleurothallis~ 35. 1859.

Illustration: Luer 1975a, f. 9 (p. 75); Luer 2006, f. 11; Karremans & Vieira-Uribe 2020, p. 286.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Specklinia grobyi (Bateman ex Lindl.) F.Barros, Hoehnea 10: 110. 1983 [1984]. Humboldtia grobyi (Bateman ex Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis grobyi Bateman ex Lindl., Edwards’s Bot. Reg. 21: t. 1797. 1835. Specklinia grobyi (Bateman ex Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001, nom. superfl.

Illustration: Luer 2006, f. 13; Misas Urreta 2005, p. 542, p. 543; Karremans et al. 2016, f. 5c, f. 6a; Karremans & Vieira-Uribe 2020, p. 284; Bogarín & Pupulin 2021, f. 13h, f. 51.

Distribution: BEL BOL BRA COL COS ECU ELS FRE GUA GUY MEX NIC PAN PER SUR VEN.

Type: Guyana, Bateman s.n. (holotype, K).

Specklinia leptantha (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004. Pleurothallis leptantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 107. 1920. Sarcinula leptantha (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 215-216. 2006.

Illustration: Luer 2006, f. 173.

Distribution: *COL.

Type: Colombia, Smith 2554 [holotype, B, destroyed; lectotype, AMES, (Luer 2006); isolectotypes, AMES, CM, NY, US].

Specklinia microphylla (A.Rich. & Galeotti) Pridgeon & M.W.Chase, Lindleyana 16(4): 258. 2001. Humboldtia microphylla (A.Rich. & Galeotti) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis microphylla A.Rich. & Galeotti, Ann. Sci. Nat., Bot., sér. 3, 3: 17. 1845.

Illustration: Luer 2006, f. 18; Karremans & Vieira-Uribe 2020, p. 285 (third row left).

Distribution: BEL COL COS ECU HON JAM MEX NIC PAN.

Type: Mexico, Galeotti 5175 (holotype, W).

Voucher: Colombia, Cárdenas-López 2069 (JAUM).

Specklinia morganii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004. Pleurothallis morganii Luer, Lindleyana 11(3): 171. 1996.

Illustration: Luer 2006, f. 21; Luer 1996c, f. 38; Misas Urreta 2005, p. 577, p. 576; Karremans & Vieira-Uribe 2020, p. 290.

Distribution: COL ECU.

Type: Ecuador, Luer 4834 (holotype, SEL).

Voucher: Colombia, Misas Urreta 342 (HPUJ).

Specklinia picta (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001. Humboldtia picta (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis picta Lindl., Edwards’s Bot. Reg. 21: sub. t. 1797. 1835.

Illustration: Cogniaux 1896, f. 6 (p. 116); Luer 2006, f. 28; Karremans & Vieira-Uribe 2020, p. 286. Fig. 16F.

Distribution: BRA BOL COL COS ECU GUA GUY PAN PER SUR.

Type: Guyana, Loddiges s.n. (holotype, K).

Specklinia purpurella (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001. Pleurothallis purpurella Luer, Orquideología 21(3): 335-337. 2000, nom. superfl. Pleurothallis purpurella Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 176. 1999. Sarcinula purpurella (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 217. 2006.

Illustration: Luer 1999a, f. 31a; Luer 2000d, p. 336; Luer 2006, f. 175.

Distribution: *COL.

Type: Colombia, Lehmann 1043 (holotype, K).

Specklinia recula (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Pleurothallis recula Luer, Lindleyana 11(2): 92. 1996.

Illustration: Luer 2006, f. 31; Misas Urreta 2005, p. 556, p. 557; Karremans et al. 2016, f. 5d; Karremans & Vieira-Uribe 2020, p. 282, p. 298.

Distribution: COL COS ECU PAN.

Type: Ecuador, Hirtz & Ortega 5364 (holotype, MO).

Specklinia scolopax (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001. Pleurothallis scolopax Luer & R.Escobar, Orquideología 14(2): 172. 1981. Sarcinula scolopax (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 217. 2006.

Illustration: Luer & Escobar 1981a, p. 173; Luer 2006, f. 176; Misas Urreta 2005, p. 564, p. 565; Karremans & Vieira-Uribe 2020, p. 291.

Distribution: *COL.

Type: Colombia, Escobar 1518 (holotype, JAUM).

Specklinia simmleriana (Rendle) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Pleurothallis simmleriana Rendle, J. Bot. 38(451): 274-275. 1900. Sarcinula simmleriana (Rendle) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 218. 2006.

Illustration: Luer 2006, f. 177; Karremans et al. 2016, f. 7l; Karremans & Vieira-Uribe 2020, p. 294; Karremans et al. 2020, f. 5.

Distribution: COL COS HON PAN.

Type: Costa Rica, Tonduz s.n. (holotype, BM000082204; isotype, G).

Specklinia striata (H.Focke) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004. Humboldtia striata (H.Focke) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis striata H.Focke, Tijdschr. Wis-Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 4: 63-64. 1851. Sarcinula striata (H.Focke) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 219. 2006.

Illustration: Luer 2006, f. 178; Karremans & Vieira-Uribe 2020, p. 294.

Distribution: COL ECU FRE GUY PAN SUR.

Type: Suriname, Focke 77 [lectotype (selected here), W-R-0025668, middle illustr. ‘23 Maart 1850’; isolectotype, K-L].

Voucher: Colombia, Lozano-Contreras s.n. (COL).

Note: The type sheet at W-R contains several different materials, including actual specimens. However, the specimens carry a note by Focke suggesting they were identified after having described P. striata, and they are in contradiction with the protologue since they show multiple inflorescences and multiple flowers. The middle illustration, which has been selected as lectotype, agrees with the protologue, also regarding preparation dates.

Specklinia tenax (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001. Acostaea tenax Luer & R.Escobar, Orquideología 15: 123. 1982.

Illustration: Luer & Escobar 1982, p. 124; Luer 1987, pl. 6.

Distribution: *COL.

Type: Colombia, Luer & Escobar 6603 (holotype, SEL; isotype, JAUM).

Specklinia trilobata (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 259. 2001. Acostaea trilobata Luer, Selbyana 1(3): 216. 1975.

Illustration: Luer 1987, pl. 7; Luer 1975d, f. 43; Karremans & Vieira-Uribe 2020, p. 280. p. 301.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 465 (holotype, SEL)

Voucher: A photographic voucher of a Colombian specimen reproduced in Karremans & Vieira-Uribe 2020, p. 280, p. 301.

Stelis Sw.

Stelis acuifera Lindl., Fol. Orchid. ~Stelis~ 12. 1852- 1855 [1859].

Illustration: Luer 2002a, f. 88; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Colombia, Moritz s.n. (holotype, K).

Stelis acuminosa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 59. 2016.

Illustration: Luer 2016a, f. 1.

Distribution: *COL.

Type: Colombia, Luer et al. 17660 (holotype, MO).

Stelis adinfinitum Luer & R.Escobar, Harvard Pap. Bot. 22(1): 27. 2017.

Illustration: Luer 2017a, f. 1.

Distribution: *COL.

Type: Colombia, Luer 2871 (holotype, SEL).

Stelis aggregata Luer & R.Escobar, Harvard Pap. Bot. 21(2): 193. 2016.

Illustration: Luer 2016b, f. 1.

Distribution: *COL.

Type: Colombia, Luer et al. 17632 (holotype, MO).

Stelis alba Kunth, Nov. Gen. Sp. (quarto ed.) 1: 363. 1816.

Illustration: Luer 2018a, f. 1; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1004.

Distribution: *COL.

Type: Colombia, Bonpland 2073 (holotype, P; isotype, P).

Stelis alboviolacea Luer, Harvard Pap. Bot. 22(2): 81. 2017.

Illustration: Luer 2017b, f. 1.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 4532 (holotype, CUVC; isotype, CUVC).

Stelis aleanophila Luer, Harvard Pap. Bot. 22(2): 81. 2017.

Illustration: Luer 2017b, f. 2.

Distribution: *COL.

Type: Colombia, Luer 6739 (holotype, SEL).

Stelis alleyoop Luer & R.Escobar, Harvard Pap. Bot. 21(1): 59. 2016.

Illustration: Luer 2016a, f. 2.

Distribution: *COL.

Type: Colombia, Luer et al. 7517 (holotype, SEL).

Stelis alloinfundibulosa J.M.H.Shaw, Orchid Rev. 122(1308): 77. 2014. Stelis eucalypta Luer, Harvard Pap. Bot. 22(1): 39. 2017, nom. superfl. Stelis infundibulosa Luer, Selbyana 30(1): 22, f. 42. 2009, nom. illeg. Non Stelis infundibulosa (Luer) Pridgeon & M.W.Chase [2001].

Illustration: Luer 2009b, f. 42; Luer 2017a, f. 23.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 4793 (holotype, CUVC).

Stelis amaliae (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 261. 2001. Pleurothallis amaliae Luer & R.Escobar, Orquideología 14(2): 124. 1981. Unciferia amaliae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer & Escobar 1981a, p. 125; Luer 1998b, f.1 (p. 98).

Distribution: *COL.

Type: Colombia, Luer 3006 (holotype, SEL).

Stelis amaliana Luer, Harvard Pap. Bot. 22(2): 112. 2017. Stelis amaliae Luer & R.Escobar, Harvard Pap. Bot. 22(1): 27. 2017, nom. illeg. Non Stelis amaliae (Luer & R.Escobar) Pridgeon & M.W.Chase [2001].

Illustration: Luer 2017a, f. 2.

Distribution: *COL.

Type: Colombia, Luer 8447 (holotype, SEL)

Stelis ambrosia Luer & R.Escobar, Harvard Pap. Bot. 21(2): 193. 2016.

Illustration: Luer 2016b, f. 2, f. 3.

Distribution: *COL.

Type: Colombia, Luer et al. 3071 (holotype, SEL).

Stelis amilotensis Luer & R.Escobar, Harvard Pap. Bot. 22(2): 82. 2017.

Illustration: Luer 2017b, f. 3.

Distribution: *COL.

Type: Colombia, Luer et al. 7479 (holotype, SEL).

Stelis amphigena O.Duque, Orquideología 27(1): 5, 6, Fotos 1-2. 2010.

Illustration: Duque 2010a, f. 1, ph. 1-2.

Distribution: *COL.

Type: Colombia, Duque 1802 (holotype, JAUM).

Stelis anchorilabia O.Duque, Orquideología 27(1): 6, 8, Fotos 3-5. 2010.

Illustration: Duque 2010a, f. 2, ph. 3-5.

Distribution: *COL.

Type: Colombia, Duque 2195 (holotype, JAUM).

Stelis ancistra (Luer & Hirtz) Pridgeon & M.W.Chase, Lindleyana 16(4): 261. 2001. Pleurothallis ancistra Luer & Hirtz, Lindleyana 11(3): 144-145. 1996. Unciferia ancistra (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 265. 2004.

Illustration: Luer 1996c, f. 5, Luer 1998b, f.2 (p.98).

Distribution: COL ECU.

Type: Ecuador, Hirtz 1469 (holotype, MO).

Voucher: Colombia, Lehmann s.n. (K).

Stelis andrei Luer, Selbyana 30(1): 20. 2009.

Illustration: Luer 2009b, f. 38.

Distribution: *COL.

Type: Colombia, André 366 (holotype, K).

Stelis angelicaKarremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe saccata Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 134-136. 2020. Non Stelis saccata Luer & Hirtz [2002].

Illustration: Kolanowska & Szlachetko 2020b, f. 35; Karremans et al. 2021, f. 4.

Distribution: *COL.

Type: Colombia, Schultes & Villarreal 7842 [lectotype, AMES-02084620; isolectotype, AMES02084544 (Karremans et al. 2021)].

Stelis angustifolia Kunth, Nov. Gen. Sp. (quarto ed.) 1: 362. 1816.

Illustration: Luer 2009a, f. 381, f. 399; Luer 2018a, f. 2-4; Luer 2018b, f. 69.

Distribution: COL ECU.

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Stelis anisopetala O.Duque, Orquideología 27(1): 10, 12. 2010.

Illustration: Duque 2010a, f. 3, ph. 6-8.

Distribution: *COL.

Type: Colombia, Duque 2301 (holotype, JAUM).

Stelis antioquiënsis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 83. 1920.

Distribution: *COL.

Type: Colombia, Lehmann 3175 [holotype, B, destroyed; lectotype (selected here), BR0000006592523; isotypes, BM-0000006592523, K-000573887, US-00093536; photo of type, AMES-00090519].

Note: The specimen at K has been indicated as holotype. Not only is this unlikely given that Schltr.’s herbarium was lost at Berlin, but the information and dimensions of the specimen at K don’t fit well with the protologue. An isotype at BR more closely fitting the description is selected as lectotype.

Stelis aprica Lindl., Companion Bot. Mag. 2: 353. 1837.

Illustration: Cogniaux 1896, t. 80 (III); Luer 2009a, f. 352; Luer 2018b, f. 7.

Distribution: BOL BRA COL ECU ELS GUY HON NIC PAN PER SUR VEN.

Type: Brazil, Martius s.n. (holotype, K).

Voucher: Colombia, Viveros 773 (HUQ).

Stelis arbuscula Luer & R.Escobar, Harvard Pap. Bot. 21(1): 60. 2016.

Illustration: Luer 2016a, f. 3.

Distribution: *COL.

Type: Colombia, Luer et al. 9029 (holotype, SEL).

Stelis argentata Lindl., Edwards’s Bot. Reg. 28: Misc. 64. 1842.

Illustration: Cogniaux 1896, t. 80 (II), t. 81 (I); Luer 2009a, f. 353-353b; Luer 2018b, f. 8, Luer 2018c, f. 1, f. 2; Misas Urreta 2005, p. 674; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1005.

Distribution: BOL BRA COL COS ECU HON FRE GUA GUY MEX NIC PAN PER SUR VEN.

Type: Guyana, Schomburgk 427 [lectotype, K (Christenson 1996); isolectotype, US].

Voucher: Colombia, Silverstone-Sopkin 8302 (MO).

Stelis arrecta Luer & R.Escobar, Harvard Pap. Bot. 21(2): 195. 2016.

Illustration: Luer 2016b, f. 4.

Distribution: *COL.

Type: Colombia, Luer et al. 10362 (holotype, MO).

Stelis ascendens Lindl., Companion Bot. Mag. 2: 353. 1836.

Illustration: Luer 2002a, f. 38; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Ecuador, Hall s.n. (holotype, K).

Voucher: Colombia, Dodson 15281 (MO).

Stelis asseris O.Duque, Orquideología 20(3): 329-330. 1997.

Illustration: Duque 1997, f. 1, ph. 1-3 (pl. I); Duque 2010b, f. 18, ph. 48-50; Luer 2018a, f. 6-7.

Distribution: *COL.

Type: Colombia, Duque 1400 (holotype, JAUM).

Stelis atomacea Luer & R.Escobar, Harvard Pap. Bot. 21(1): 60. 2016.

Illustration: Luer 2016a, f. 4.

Distribution: *COL.

Type: Colombia, Luer et al. 3693 (holotype, SEL).

Stelis atra Lindl., Orchid. Linden. 3, n. 15. 1846. Stelis hallii var. atra (Lindl.) Garay & C.Schweinf., Fieldiana, Bot. 33: 11. 1970.

Illustration: Luer 2004, f. 152 (p. 182); Luer 2009a, f. 152a (p. 205), Luer 2017a, f. 13; Luer 2018b, f. 9; Luer 2018c, f. 9, f. 10; Romero-González & Carnevalli 2000, p. 1006.

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Linden 687 (holotype, K).

Voucher: Colombia, Vélez 1600 (HUQ).

Stelis atroviolacea Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Illustration: Luer 2007, f. 256; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1007.

Distribution: COL ECU VEN.

Type: Colombia, Wagener s.n. (holotype, W).

Stelis attenuata Lindl., Fol. Orchid. ~Stelis~ 2. n. 7. 1852-1855 [1859].

Illustration: Luer 2007, f. 251; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU PER.

Type: Colombia, Purdie s.n. (holotype, K).

Stelis aurea (Lindl.) Karremans, Lankesteriana 13(3): 328. 2014. Pleurothallis aurea Lindl., Ann. Mag. Nat. Hist. 12(79): 397. 1843.

Illustration: Cogniaux 1896, f. 1 (p. 110); Luer 1975a, f. 1 (p. 59); Luer 1999a, f. 1 (p. 126).

Distribution: BOL BRA COL ECU PER VEN.

Type: Venezuela, Linden 634 (holotype, K-L000334396; isotypes, F-0046447F, G-00168961, LE-00001408, W-R-0053798, W-R-0012745, W-R-0046081).

Voucher: Colombia, García-Barriga & Jaramillo 19885 (AMES, COL).

Stelis auriculata O.Duque, Orquideología 27(1): 14, 16. 2010.

Illustration: Duque 2010a, f. 5, ph. 13-14.

Distribution: *COL.

Type: Colombia, Villegas s.n. (holotype, JAUM).

Stelis aviceps Lindl., Fol. Orchid. ~Stelis~ 16, n. 121. 1852-1855 [1859].

Illustration: Luer 2009a, f. 310; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1008.

Distribution: BOL COL ECU GUY PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Viveros 396 (HUQ).

Stelis barbuda O.Duque, Orquideología 20(3): 330-331. 1997.

Illustration: Duque 1997, f. 2, ph. 4-6 (pl. I).

Distribution: *COL.

Type: Colombia, Duque 1896 (holotype, JAUM).

Stelis barkleyi (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe barkleyi Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 109-110. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 5.

Distribution: *COL.

Type: Colombia, Barkley & Bouthillette 38C057 (holotype, AMES; drawing, UGDA).

Stelis batillacea (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 261. 2001. Crocodeilanthe batillacea (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis batillacea Luer, Selbyana 3(1-2): 58. 1976.

Illustration: Luer 1998a, f. 6.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 853 (holotype, SEL).

Voucher: Colombia, Luer 3142 (SEL).

Stelis baudoensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 29. 2017.

Illustration: Luer 2017a, f. 3; Misas Urreta 2005, p. 688.

Distribution: *COL.

Type: Colombia, de Wilde 3939 (holotype, SEL).

Stelis bella O.Duque, Orquideología 20(3): 331-332. 1997.

Illustration: Duque 1997, f. 3, ph. 7-9 (pl. I); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1592 (holotype, JAUM).

Stelis bialaria Luer & R.Escobar, Harvard Pap. Bot. 22(2): 82. 2017.

Illustration: Luer 2017b, f. 4.

Distribution: *COL.

Type: Colombia, Luer et al. 10351 (holotype, MO).

Stelis bialata (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe bialata Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 110-111. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 6.

Distribution: *COL.

Type: Colombia, Mora 3256 (holotype, AMES; drawing, UGDA).

Stelis bicolor Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 143, 177. 2004.

Illustration: Luer 2004, f. 134 (p. 177).

Distribution: COL ECU.

Type: Ecuador, Hirtz 8420 (holotype, MO).

Voucher: Colombia, Duque 2338 (JAUM).

Stelis bicornis Lindl., Fol. Orchid. ~Stelis~ 18, no. 133. 1852-1855 [1859].

Illustration: Luer 2009a, f. 311, f. 311a.

Distribution: COL ECU PER.

Type: Ecuador, Jameson 490 (holotype, K; isotypes, AMES, BM, G).

Voucher: Colombia, Duque 2039 (JAUM).

Stelis bigibba Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 84. 1920.

Illustration: Luer 2018a, f. 8.

Distribution: *COL.

Type: Colombia, Lehmann 6834 (holotype, B, destroyed; isotype, AMES). [holotype, B, destroyed; lectotype, AMES-00084071 (Luer 2018a); isolectotypes, LE-00001398, NY-00009386, K000573882].

Stelis biserrula Lindl., Fol. Orchid. ~Stelis~ 16, n. 120. 1852-1855 [1859].

Illustration: Luer 2004, f. 119 (p. 137); Ortiz & Uribe 2007 (CD Edition); Duque 2010, pl. 5 (8, 9), pl. 6 (1).

Distribution: COL COS ECU PAN PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Misas Urreta 281 (HPUJ).

Stelis blandita Luer & R.Escobar, Harvard Pap. Bot. 22(2): 82. 2017.

Illustration: Luer 2017b, f. 5.

Distribution: *COL.

Type: Colombia, Luer et al. 7606 (holotype, SEL).

Stelis bogotensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 152. 1924.

Distribution: *COL.

Type: Colombia, Hopf s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis bordoncillensis (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe bordoncillensis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 112. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 8.

Distribution: *COL.

Type: Colombia, Medina et al. S18/12 (holotype, JAUM).

Stelis boyacensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 29. 2017.

Illustration: Luer 2017a, f. 4.

Distribution: *COL.

Type: Colombia, Luer et al. 7559 (holotype, SEL).

Stelis braccata Rchb.f. & Warsz., Bonplandia (Hannover) 2: 114. 1854. Apatostelis braccata (Rchb.f. & Warsz.) Garay, Bot. Mus. Leafl. 27: 187. 1979.

Illustration: Luer 2009a, f. 355, f. 355a; Luer 2018c, f. 3; Romero-González & Carnevalli 2000, p. 1033.

Distribution: COL ECU VEN.

Type: Colombia, Warszewicz s.n. (holotype, W; isotype, AMES; fragment of isotype, AMES-spirit).

Stelis bracteolenta Luer & R.Escobar, Harvard Pap. Bot. 21(2): 195. 2016.

Illustration: Luer 2016b, f. 5. Fig. 15D.

Distribution: *COL.

Type: Colombia, Luer et al. 10369 (holotype, SEL).

Stelis brevilabris Lindl., Ann. Mag. Nat. Hist. 5: 107. 1845.

Illustration: Luer 2002a, f. 99; Luer 2018c, f. 4; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU PER.

Type: Colombia, Hartweg 1411 (holotype, K; isotypes, BR, F, G, LD, LE, W).

Stelis bucaramangae (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 261. 2001. Pleurothallis bucaramangae Luer & R.Escobar, Orquideología 20: 38. 1996. Crocodeilanthe bucaramangae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer & Escobar 1996, p. 39; Luer 1998a, f. 7.

Distribution: *COL.

Type: Colombia, Luer et al. 10107 (holotype, MO).

Stelis calawayiKarremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe dodsonii Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 117. 2020. Non Stelis dodsonii Luer [2002].

Illustration: Kolanowska & Szlachetko 2020b, f. 14.

Distribution: *COL.

Type: Colombia, Dodson & Escobar 18130 (holotype, AMES; drawing, UGDA).

Stelis calceolarisGaray, Bot. Mus. Leafl. 27(7-9): 185. 1979.

Illustration: Garay 1979, f. VIII-c-10 (p. 234); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Karsten s.n. (holotype, LE).

Stelis calculosa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 62. 2016.

Illustration: Luer 2016a, f. 6.

Distribution: *COL.

Type: Colombia, Luer et al. 16838 (holotype, MO).

Stelis caldodsonii Luer & R.Escobar, Harvard Pap. Bot. 21(1): 62. 2016.

Illustration: Luer 2016a, f. 7.

Distribution: *COL.

Type: Colombia, Dodson & Escobar 15316 (holotype, MO).

Stelis caliensis Luer, Selbyana 30(1): 21. 2009.

Illustration: Luer 2009b, f. 40.

Distribution: *COL.

Type: Colombia, André 2642 (holotype, K).

Stelis callosaKarremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe purpurea Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 128-129. 2020. Non Stelis purpurea (Ruiz & Pav.) Willd. [1805].

Illustration: Kolanowska & Szlachetko 2020b, f. 28.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 19831bis (holotype, AMES-102872; drawing, UGDA).

Stelis calopsis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 29. 2017.

Illustration: Luer 2017a, f. 5, 6.

Distribution: *COL.

Type: Colombia, Ecobar 5130 (holotype, SEL).

Stelis calotricha Schltr., Notizbl. Bot. Gart. BerlinDahlem 7: 323. 1919.

Illustration: Misas Urreta 2005, p. 680.

Distribution: BRA COL VEN.

Type: Brazil, von Grossmann s.n. (holotype, B, destroyed).

Voucher: Colombia, Misas Urreta 124 (HPUJ).

Stelis carnalis Luer & R.Escobar, Harvard Pap. Bot. 21(1): 63. 2016.

Illustration: Luer 2016a, f. 8.

Distribution: *COL.

Type: Colombia, Luer et al. 11376 (holotype, MO).

Stelis cassidis (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 261. 2001. Crocodeilanthe cassidis (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia cassidis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis cassidis Lindl., Ann. Mag. Nat. Hist. 15: 384. 1845.

Illustration: Luer 1998a, f. 8, f. 9.

Distribution: COL ECU PER.

Type: Colombia, Hartweg s.n. (holotype, K).

Stelis catenataKarremans, Lankesteriana 13(3): 328. 2014. Anathallis ramulosa (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Humboldtia ramulosa (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis ramulosa Lindl., Fol. Orchid. ~Pleurothallis~ 33. 1859. Specklinia ramulosa (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Non Stelis ramulosa Luer & Dalström [2004].

Illustration: Luer 1999a, f. 21 (p. 131).

Distribution: COL ECU VEN.

Type: Ecuador, Jameson s.n. (holotype, K-L).

Voucher: Colombia, Ariste-Joseph A198 (US).

Stelis caudex Luer & R.Escobar, Harvard Pap. Bot. 22(1): 30. 2017.

Illustration: Luer 2017a, f. 7, 8.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8387 (holotype, SEL).

Stelis caveata O.Duque, Orquideología 27(1): 16, 18. 2010.

Illustration: Duque 2010a, f. 6, ph. 15-17.

Distribution: *COL.

Type: Colombia, Duque 2080 (holotype, JAUM).

Stelis cesarensis (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 136. 2021. Crocodeilanthe cesarensis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 113. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 9.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 20633 (holotype, AMES; drawing, UGDA).

Stelis chalatantha Luer & Hirtz, Harvard Pap. Bot. 21(2): 195. 2016.

Illustration: Luer 2016b, f. 6.

Distribution: *COL.

Type: Colombia, Luer et al. 4597 (holotype, SEL).

Stelis chamaestelis (Rchb.f.) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 4: 292. 1966. Apatostelis chamaestelis (Rchb.f.) Garay, Bot. Mus. Leafl. 27: 187. 1979. Pleurothallis chamaestelis Rchb.f., Linnaea 22: 825. 1850.

Illustration: Luer 2018a, f. 9; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1011.

Distribution: COL VEN.

Type: Venezuela, Schlim 1195 (holotype, G; isotypes, LE, MPU, P).

Voucher: Colombia, Luer et al. 10336 (MO).

Stelis chasmiphora Luer & R.Escobar, Harvard Pap. Bot. 22(2): 84. 2017.

Illustration: Luer 2017b, f. 6.

Distribution: *COL.

Type: Colombia, Luer et al. 3720 (holotype, SEL).

Stelis chiliantha Luer & R.Escobar, Harvard Pap. Bot. 22(2): 84. 2017.

Illustration: Luer 2017b, f. 7.

Distribution: *COL.

Type: Colombia, Luer & Escobar 6599 (holotype, SEL).

Stelis chocoënsis O.Duque, Orquideología 27(1): 20. 2010.

Illustration: Duque 2010a, f. 7, ph. 18-20; Luer 2018a, f. 10.

Distribution: *COL.

Type: Colombia, Duque 2192 (holotype JAUM).

Stelis choerorhyncha (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 261-262. 2001. Pleurothallis choerorhyncha Luer, Orquideología 20: 204. 1996. Crocodeilanthe choerorhyncha (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer 1996f, 205; Luer 1998a, f. 11.

Distribution: *COL.

Type: Colombia, Cuatrecasas 18116 (holotype, US; isotype, AMES).

Stelis ciliaris Lindl., Companion Bot. Mag. 2: 353. 1836 [1837]. Apatostelis ciliaris (Lindl.) Garay, Bot. Mus. Leafl. 27(7-9): 188. 1979.

Illustration: Luer & Toscano 2012, f. 9; Luer 2009a, f. 357; Romero-González & Carnevalli 2000, p. 1021.

Distribution: BEL BRA COL COS ECU GUA HON MEX NIC PAN VEN.

Type: Mexico, Hartweg s.n. (holotype, K).

Voucher: Colombia, Marín & Ávila 5452 (FMB).

Stelis cladophora Luer & R.Escobar, Harvard Pap. Bot. 22(1): 30. 2017.

Illustration: Luer 2017a, f. 9.

Distribution: *COL.

Type: Colombia, Luer et al. 7585 (holotype, SEL).

Stelis cleistogamoides O.Duque, Orquideología 27(1): 22, 24. 2010.

Illustration: Duque 2010a, f. 8, ph. 21.

Distribution: *COL.

Type: Colombia, Duque 1401 (holotype JAUM).

Stelis clematis Luer & R.Escobar, Harvard Pap. Bot. 21(2): 196. 2016.

Illustration: Luer 2016b, f. 7.

Distribution: *COL.

Type: Colombia, Luer 13261 (holotype, MO).

Stelis climacella Luer & R.Escobar, Harvard Pap. Bot. 21(2): 196. 2016.

Illustration: Luer 2016b, f. 8.

Distribution: *COL.

Type: Colombia, Luer et al. 8959 (holotype, SEL).

Stelis clipeus O.Duque, Orquideología 20(3): 333, 338. 1997.

Illustration: Duque 1997, f. 4, ph. 1-2 (pl. II); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 595 (holotype, JAUM).

Stelis coccidifera Luer & R.Escobar, Harvard Pap. Bot. 22(2): 84. 2017.

Illustration: Luer 2017b, f. 8.

Distribution: *COL.

Type: Colombia, Luer et al. 8454 (holotype, SEL).

Stelis cochlearis Garay, Canad. J. Bot. 34: 352. 1956.

Illustration: Luer 2016a, f. 40; Luer 2016b, f. 39; Luer 2018c, f. 5.

Distribution: *COL.

Type: Colombia, Johnson & Barkley 18C804 (holotype, US-2102700; isotypes, AMES, MEDEL).

Stelis cochleata (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe cochleata Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 113-114. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 10.

Distribution: *COL.

Type: Colombia, Pennell 10475 (holotype, AMES40953; drawing, UGDA).

Stelis cochliops Luer & Hirtz, Harvard Pap. Bot. 21(1): 63. 2016.

Illustration: Luer 2016a, f. 9.

Distribution: *COL.

Type: Colombia, Hirtz 8562 (holotype, MO).

Stelis cocornaënsis (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001. Effusiella cocornaënsis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Pleurothallis cocornaënsis Luer & R.Escobar, Orquideología 20: 45. 1996. Specklinia cocornaënsis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004.

Illustration: Luer & Escobar 1996, p. 47, p. 58 (middle left); Luer 2000a, f. 5 (p. 91).

Distribution: *COL.

Type: Colombia, Escobar 5047 (holotype, JAUM; isotype, MO).

Stelis coelochilaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 195, 198. 2004.

Illustration: Luer 2009a, f. 358.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 9793 (holotype, MO).

Voucher: Colombia, Duque 2136 (JAUM).

Stelis colombiana Ames, Sched. Orch. 1: 4. 1922.

Illustration: Luer 2018a, f. 11; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Pittier 1093 (holotype, AMES).

Stelis colorata O.Duque, Orquideología 27(1): 24, 26. 2010.

Illustration: Duque 2010a, f. 9, ph. 22-24.

Distribution: *COL.

Type: Colombia, Duque 2215 (holotype JAUM).

Stelis colossus Luer & R.Escobar, Harvard Pap. Bot. 21(2): 198. 2016.

Illustration: Luer 2016b, f. 9.

Distribution: *COL.

Type: Colombia, Luer et al. 16812 (holotype, MO).

Stelis comica O.Duque, Orquideología 20(3): 338-339. 1997.

Illustration: Duque 1997, f. 1 (p. 345), ph. 3-6 (pl. II); Luer 2018a, f. 12; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1232 (holotype, JAUM).

Stelis conchipetala Luer & R.Escobar, Harvard Pap. Bot. 22(2): 86. 2017.

Illustration: Luer 2017b, f. 9.

Distribution: *COL.

Type: Colombia, Luer 6838 (holotype, SEL).

Stelis concinna Lindl., J. Bot. (Hooker) 1: 11. 1834.

Illustration: Luer 2002a, f. 67; Luer 2009a, f. 67b; Luer 2018b, f. 30; Luer 2018c, f. 6; Romero-González & Carnevalli 2000, p. 1019.

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Hall s.n. (holotype, K).

Voucher: Colombia, Escobar 5145 (JAUM, MO).

Stelis convoluta (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001. Specklinia convoluta (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 259. 2004. Effusiella convoluta (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Humboldtia convoluta (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis convoluta Lindl., Ann. Mag. Nat. Hist. 15: 107. 1845.

Illustration: Luer 1999b, p. 123; Luer 2000a, f. 7 (p. 91).

Distribution: *COL.

Type: Colombia, Hartweg 1414 (holotype, K-L; isotypes, AMES-00074175, BM-000082417, BR-0000006585389, G-00168959,K-000584190,K-001382013,K-001382012, LD-1404157, W-R-0012245).

Stelis coraeFoldats, Acta Bot. Venez. 3: 401. 1968. Apatostelis corae (Foldats) Garay, Bot. Mus. Leafl. 27: 188. 1980.

Illustration: Foldats 1968, f. 31; Romero-González & Carnevalli 2000, p. 1012.

Distribution: COL VEN.

Type: Venezuela, Steyermark & Dunsterville 98725 (holotype, VEN).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition).

Stelis cordilabia (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe cordilabia Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 114-115. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 11.

Distribution: *COL.

Type: Colombia, Garay 970 (holotype, AMES; drawing, UGDA).

Stelis crassilabia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 87. 1920.

Illustration: Romero-González & Carnevalli 2000, p. 1014.

Distribution: COL VEN.

Type: Colombia, Lehmann 2102 [holotype, B, destroyed; lectotype (selected here), K-000573874; isolectotypes, BM-000084121, BR0000006594091, US-00093553].

Stelis crassisepala Luer & R.Escobar, Harvard Pap. Bot. 22(1): 33. 2017.

Illustration: Luer 2017a, f. 10.

Distribution: *COL.

Type: Colombia, Luer et al. 3052 (holotype, SEL).

Stelis craticula Luer & R.Escobar, Harvard Pap. Bot. 22(1): 33. 2017.

Illustration: Luer 2017a, f. 11.

Distribution: *COL.

Type: Colombia, Luer et al. 14206 (holotype, SEL).

Stelis crenulata O.Duque, Orquideología 27(1): 28, 30. 2010.

Illustration: Duque 2010a, f. 10, ph. 25-27.

Distribution: *COL.

Type: Colombia, Duque 2115 (holotype, JAUM).

Stelis crescentiicola Schltr., Repert. Spec. Nov. Regni Veg. 16(468-473): 442. 1920.

Distribution: COL COS ELS NIC PAN.

Type: Panama, Lehmann 4540 [holotype, B, destroyed; lectotype, US-00093554 (814880), Luer (2023); isolectotype, LE-00001389; illustration, AMES].

Voucher: Colombia, Duque 2139 (JAUM).

Stelis cruenta O.Duque, Orquideología 27(1): 30, 32, 34, Fotos 28-30. 2010.

Illustration: Duque 2010a, f. 11, ph. 28-30.

Distribution: *COL.

Type: Colombia, Duque 1960 (holotype, JAUM).

Stelis cuatrecasasii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001. Crocodeilanthe cuatrecasasii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis cuatrecasasii Luer, Orquideología 20: 208. 1996.

Illustration: Luer 1996f, 209; Luer 1998a, f. 14.

Distribution: *COL.

Type: Colombia, Cuatrecasas 19264 (holotype, NY).

Stelis cucullata Ames, Sched. Orch. 6: 52-53. 1923. Illustration: Romero-González & Carnevalli 2000, p. 1015.

Distribution: COL COS NIC VEN.

Type: Costa Rica, Lankester 475 (holotype, AMES-26975).

Voucher: Colombia, Ospina 68 (JAUM).

Stelis cuculligera Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 27. 1924.

Distribution: *COL.

Type: Colombia, Hopp 169 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis culmosa Luer & R.Escobar, Harvard Pap. Bot. 22(2): 86. 2017.

Illustration: Luer 2017b, f. 10.

Distribution: *COL.

Type: Colombia, Luer et al. 7601 (holotype, SEL).

Stelis cundinamarcae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 153. 1924.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis cupidinea Luer & R.Escobar, Harvard Pap. Bot. 22(2): 88. 2017.

Illustration: Luer 2017b, f. 11.

Distribution: *COL.

Type: Colombia, Luer 8451 (holotype, SEL).

Stelis cupreata Luer & R.Escobar, Harvard Pap. Bot. 22(1): 33. 2017.

Illustration: Luer 2017a, f. 12.

Distribution: *COL.

Type: Colombia, de Wilde 4435 (holotype, SEL).

Stelis curiosa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 65. 2016.

Illustration: Luer 2016a, f. 11.

Distribution: *COL.

Type: Colombia, Luer 16772 (holotype, MO).

Stelis cyathochila Luer & R.Escobar, Harvard Pap. Bot. 21(2): 198. 2016.

Illustration: Luer 2016b, f. 11.

Distribution: *COL.

Type: Colombia, Luer et al. 10257 (holotype, MO).

Stelis cycloglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 154. 1924.

Distribution: *COL.

Type: Colombia, Hopf s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis dapidis Luer & R.Escobar, Harvard Pap. Bot. 23(2): 146. 2018.

Illustration: Luer 2018c, f. 7.

Distribution: *COL.

Type: Colombia, Luer et al. 6699 (holotype, SEL).

Stelis dapsilis Pridgeon & M.W.Chase, Lindleyana 17(2): 99. 2002. Crocodeilanthe maxima (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis maxima Luer, Selbyana 3(1-2): 140. 1976. Stelis maxima (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001, nom. illeg. Non Stelis maxima Lindl. [1845].

Illustration: Luer 1998a, f. 39.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 651 (holotype, SEL).

Voucher: Colombia, Luteyn 13299 (NY).

Stelis dasysepala Luer & R.Escobar, Harvard Pap. Bot. 21(2): 198. 2016.

Illustration: Luer 2016b, f. 10.

Distribution: *COL.

Type: Colombia, Luer et al. 8749 (holotype, SEL).

Stelis decipiens Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 155. 1924.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis decipula Luer & R.Escobar, Harvard Pap. Bot. 21(1): 65. 2016.

Illustration: Luer 2016a, f. 12.

Distribution: *COL.

Type: Colombia, Luer et al. 4506 (holotype, SEL).

Stelis declivis (Lindl.) Luer, Selbyana 23(1): 44. 2002. Anathallis declivis (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 248. 2001. Humboldtia declivis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis declivis Lindl., Fol. Orchid. ~Pleurothallis~ 36, no. 218. 1859.

Illustration: Luer 2009a, f. 362.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, García-Barriga 19828 (COL).

Stelis dejavu Luer & R.Escobar, Harvard Pap. Bot. 22(2): 88. 2017.

Illustration: Luer 2017b, f. 12.

Distribution: *COL.

Type: Colombia, Luer et al. 6672 (holotype, SEL).

Stelis deltoidea (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe deltoidea Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 116-117. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 13.

Distribution: *COL.

Type: Colombia, Garay et al. 135 (holotype, AMES; drawing, UGDA).

Stelis dentata O.Duque, Orquideología 20(3): 339-340. 1997.

Illustration: Duque 1997, f. 2 (p. 345), ph. 7-9 (pl. II); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1280 (holotype, JAUM).

Stelis depauperata Lindl., Fol. Orchid. ~Stelis~ 16, no. 124. 1852-1855 [1859].

Illustration: Luer 2007, f. 253; Luer 2009a, f. 253a; Misas Urreta 2005, p. 682.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Misas Urreta 258 (HPUJ).

Stelis desautelsii Luer, Harvard Pap. Bot. 22(2): 88. 2017.

Illustration: Luer 2017b, f. 13.

Distribution: *COL.

Type: Colombia, Luer 6765 (holotype, SEL).

Stelis deutrodewildei J.M.H.Shaw, Orchid Rev. Suppl., 127(1326): 46. 2019. Crocodeilanthe dewildeiLuer & Toscano, Harvard Pap. Bot. 23(1): 47. 2018. Stelis dewildei (Luer & Toscano) Karremans, Phytotaxa 406(5): 265. 2019, nom. illeg. Stelis toscanoi Karremans, Lankesteriana 19(3): 289. 2019, nom. superfl. Non Stelis dewildei Luer & R.Escobar [2017].

Illustration: Luer & Toscano de Brito 2018, f. 1.

Distribution: *COL.

Type: Colombia, de Wilde 339 (holotype, MO).

Stelis dewildei Luer & R.Escobar, Harvard Pap. Bot. 22(1): 34. 2017.

Illustration: Luer 2017a, f. 14.

Distribution: *COL.

Type: Colombia, de Wilde 03 (holotype, SEL).

Stelis dialissa Rchb.f., Bonplandia (Hannover) 3: 71. 1855. Dialissa pulchella Lindl., Ann. Mag. Nat. Hist. 15(96): 107. 1845. Non Stelis pulchella Kunth [1816].

Illustration: Luer 2009a, f. 314; Luer 2018c, f. 8; Ortiz & Uribe 2007 (CD Edition). Fig. 14I.

Distribution: COL ECU.

Type: Colombia, Hartweg 1413 (holotype, W; isotypes, BM, BR, G, K, LD, LE).

Stelis digitifera Luer & R.Escobar, Harvard Pap. Bot. 21(1): 67. 2016.

Illustration: Luer 2016a, f. 13.

Distribution: *COL.

Type: Colombia, Luer et al. 14179 (holotype, MO).

Stelis dimidia (Luer) Karremans, Lankesteriana 13(3): 328. 2014. Anathallis dimidia (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 248. 2001. Pleurothallis dimidia Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 109-110. 1999. Specklinia dimidia (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004.

Illustration: Luer 1999a, f. 11 (p. 128).

Distribution: BOL COL PER VEN.

Type: Bolivia, Luer et al. 16497 (holotype, MO).

Voucher: Colombia, Foster et al. 1332 (AMES).

Stelis diprizo Luer, Harvard Pap. Bot. 22(2): 89. 2017.

Illustration: Luer 2017b, f. 15.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 2665 (holotype, MO; isotype, CUVC).

Stelis discolor Rchb.f., Bonplandia (Hannover) 3: 240. 1855.

Illustration: Luer 2009a, f. 363; Duque 2010, pl. 10 (1-3). Fig. 15K.

Distribution: COL ECU PER.

Type: Ecuador, Jameson 691 (holotype, W; isotypes: AMES, BM, BR, G, K).

Voucher: Colombia, Lehmann s.n. (W).

Stelis discors Luer & Hirtz, Harvard Pap. Bot. 22(2): 89. 2017.

Illustration: Luer 2017b, f. 14.

Distribution: *COL.

Type: Colombia, Luer et al. 4563 (holotype, SEL).

Stelis dissidens Luer & R.Escobar, Harvard Pap. Bot. 22(2): 89. 2017.

Illustration: Luer 2017b, f. 16; Misas Urreta 2005, p. 692.

Distribution: *COL.

Type: Colombia, Luer 17584 (holotype, MO).

Stelis dithele Luer & R.Escobar, Harvard Pap. Bot. 22(1): 36. 2017.

Illustration: Luer 2017a, f. 15.

Distribution: *COL.

Type: Colombia, Luer et al. 7464 (holotype, SEL).

Stelis divaricans Luer & R.Escobar, Harvard Pap. Bot. 21(2): 199. 2016.

Illustration: Luer 2016b, f. 12.

Distribution: *COL.

Type: Colombia, Luer et al. 3161 (holotype, SEL).

Stelis divergens Luer & R.Escobar, Harvard Pap. Bot. 22(1): 36. 2017.

Illustration: Luer 2017a, f. 16; Misas Urreta 2005, p. 696.

Distribution: *COL.

Type: Colombia, de Wilde 4500 (holotype, SEL).

Stelis dolichopus Schltr., Orchis 6: 63. 1912.

Distribution: *COL.

Type: Colombia, unknown (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis dressleri Luer, Phytologia 49(3): 227-228. 1981.

Illustration: Luer 2018a, f. 13.

Distribution: COL COS PAN.

Type: Panama, Luer & Dressler 1146 (holotype, SEL).

Voucher: Colombia, Duque 1835 (holotype, JAUM).

Stelis dupliciformis C.Schweinf., Bot. Mus. Leafl. 15: 15, t. 8. 1951.

Distribution: COL PER.

Type: Peru, Vargas 3816 (holotype, AMES-65251).

Voucher: Colombia, Rodriguez 269 (FMB).

Stelis duquei P.Ortiz, Orquideología 28: 15. 2011. Stelis globosa O.Duque, Orquideología 27(2): 124, 127. 2010, nom. illeg. Non Stelis globosa Pridgeon & M.W.Chase [2002].

Illustration: Duque 2010b, f. 14, ph. 39-40.

Distribution: *COL.

Type: Colombia, Duque 2150 (holotype, JAUM).

Stelis dunstervilleorumFoldats, Acta Bot. Venez. 3: 404. 1968.

Illustration: Foldats 1968, f. 32; Romero-González & Carnevalli 2000, p. 1016.

Distribution: COL VEN.

Type: Venezuela, Steyermark & Dunsterville 98529 (holotype, VEN).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition).

Stelis ecmeles Luer, Harvard Pap. Bot. 23(1): 26. 2018.

Illustration: Luer 2018a, f. 14.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 4310 (holotype, MO; isotype, CUVC).

Stelis efsiella Luer, Selbyana 30(1): 21-22. 2009.

Illustration: Luer 2009b, f. 41.

Distribution: *COL.

Type: Colombia, Lehmann 3300 (holotype, K).

Stelis ejuncida Luer & R.Escobar, Harvard Pap. Bot. 22(1): 37. 2017.

Illustration: Luer 2017a, f. 18. Fig. 14G.

Distribution: *COL.

Type: Colombia, Luer et al. 10111 (holotype, SEL).

Stelis elementaria Luer & R.Escobar, Harvard Pap. Bot. 22(2): 91. 2017.

Illustration: Luer 2017b, f. 17.

Distribution: *COL.

Type: Colombia, Luer et al. 7699 (holotype, SEL).

Stelis elegantissima J.M.Murillo, Rom.Cec. & Karremans, Phytotaxa 496(2): 137. 2021. Stelis crascens Luer & R.Escobar, Harvard Pap. Bot. 21(1): 65. 2016, nom. illeg. Non Stelis cracens Luer & Hirtz [2013].

Distribution: *COL.

Type: Colombia, Luer et al. 11373 (holotype, MO).

Stelis ellipsophylla Luer & R.Escobar, Harvard Pap. Bot. 22(1): 37. 2017.

Illustration: Luer 2017a, f. 19.

Distribution: *COL.

Type: Colombia, Luer et al. 17689 (holotype, MO).

Stelis elongata Kunth, Nov. Gen. Sp. (quarto ed.) 1: 363. 1816.

Illustration: Luer 2009a, f. 315, f. 315a; Luer 2018b, f. 40; Ortiz & Uribe 2007 (CD Edition).

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; photo, AMES).

Stelis emarginata (Ruiz & Pav.) Soto Arenas & Solano, Icon. Orchid. (Mexico) 5-6: t. 681. 2002 [2003]. Epidendrum emarginatum Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 247. 1798. Humboldtia emarginata Pav. ex Lindl., Gen. Sp. Orchid. Pl. 6. 1830. Physosiphon emarginatus (Ruiz & Pav.) Lindl., Edwards’s Bot. Reg. 21: sub t. 1797. 1835. Pleurothallis emarginata (Ruiz & Pav.) Lindl., Gen. Sp. Orchid. Pl. 6. 1830.

Illustration: Luer 2007, f. 3 (p. 104).

Distribution: COL GUA MEX PER.

Type: Peru, Pavon 595 (holotype, W; isotypes, K-L; BM-000082027).

Voucher: Colombia, Lehmann 4108 (G).

Stelis embreei Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 14-15. 2002.

Illustration: Luer 2002a, f. 15; Rincón-González et al. 2020a, f. 5.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11949 (holotype, MO).

Voucher: Colombia, Rincón et al. 1277 (TOLI).

Stelis enormis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 37. 2017.

Illustration: Luer 2017a, f. 20.

Distribution: *COL.

Type: Colombia, Luer 17563 (holotype, MO).

Stelis erecta O.Duque, Orquideología 20(3): 341-342. 1997.

Illustration: Duque 1997, f. 3 (p. 345), ph. 1-3 (pl. III); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1677 (holotype, JAUM).

Stelis erucosa (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001. Effusiella erucosa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Pleurothallis erucosa Luer & R.Escobar, Orquideología 21(1): 88. 1998. Specklinia erucosa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004.

Illustration: Luer & Escobar 1998, p. 89; Luer 2000a, f. 12 (p. 92).

Distribution: *COL.

Type: Colombia, Luer 2341 (holotype, SEL).

Stelis escobarii Luer, Harvard Pap. Bot. 22(2): 112. 2017. Stelis rodrigoi Luer, Harvard Pap. Bot. 21(1): 85. 2016, nom. illeg. Non Stelis rogrigoi (Luer) Pridgeon & M.W.Chase [2001].

Illustration: Luer 2016a, f. 44. Fig. 14A.

Distribution: *COL.

Type: Colombia, Escobar 5141 (holotype, MO).

Stelis eublepharis Rchb.f., Bonplandia (Hannover) 3: 240. 1855.

Illustration: Luer 2009a, f. 368, f. 368a; Luer 2018c, f. 11, f. 12; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1017. Fig. 14L.

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Jameson 264 (holotype, K; isotypes, AMES, BR, G, LE, W).

Voucher: Colombia, Duque 1712 (JAUM, MO).

Stelis eugenii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 87. 1920.

Illustration: Mansfeld 1929, Pl. 29 (No. 110); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Langlassé 104 [holotype, B, destroyed; lectotype (selected here), P-00489279; isolectotypes, AMES-00104751, G-00169085, G-00169128, GH-00104750, MPU-018358, P-00489280, US-00093559].

Stelis euglossina Luer & R.Escobar, Harvard Pap. Bot. 21(1): 68. 2016.

Illustration: Luer 2016a, f. 15.

Distribution: *COL.

Type: Colombia, Luer et al. 10191 (holotype, MO).

Stelis euprepes Luer & R.Escobar, Harvard Pap. Bot. 22(1): 41. 2017.

Illustration: Luer 2017a, f. 24.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8378 (holotype, SEL).

Stelis euspatha Rchb.f., Bonplandia (Hannover) 3: 225. 1855.

Illustration: Luer 2009a, f. 392; Luer 2018b, f. 42.

Distribution: BOL COL ECU PER.

Type: Bolivia, Bridges s.n. (holotype, W).

Voucher: Colombia, Jameson s.n. (K).

Stelis euthema Luer & R.Escobar, Harvard Pap. Bot. 21(1): 68. 2016.

Illustration: Luer 2016a, f. 16.

Distribution: *COL.

Type: Colombia, Luer et al. 9041 (holotype, SEL).

Stelis exaltata Luer & R.Escobar, Harvard Pap. Bot. 21(1): 68. 2016.

Illustration: Luer 2016a, f. 17.

Distribution: *COL.

Type: Colombia, Luer et al. 16791 (holotype, MO).

Stelis excentrica Reina-Rodr. & López-Mach., Lankesteriana 20(2): 152. 2020.

Illustration: Reina-Rodríguez et al. 2020, f. 1-3.

Distribution: *COL.

Type: Colombia, Reina-Rodríguez et al. 2957 (holotype, CUVC-spirit; isotype, COL).

Stelis expansa (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001. Crocodeilanthe expansa (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia expansa (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis expansa Lindl., Fol. Orchid. ~Pleurothallis~ 4. 1859.

Illustration: Luer 1998a, f. 19; Luer 1975f, f. 56.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Jameson 308 (G).

Stelis famelica Luer & R.Escobar, Harvard Pap. Bot. 22(1): 41. 2017.

Illustration: Luer 2017a, f. 25. Fig. 15B.

Distribution: *COL.

Type: Colombia, Luer et al. 8768 (holotype, SEL).

Stelis fascinata Luer & R.Escobar, Harvard Pap. Bot. 22(1): 42. 2017.

Illustration: Luer 2017a, f. 26.

Distribution: *COL.

Type: Colombia, Luer et al. 3160 (holotype, SEL).

Stelis fecunda Luer & R.Escobar, Harvard Pap. Bot. 22(2): 91. 2017.

Illustration: Luer 2017b, f. 18.

Distribution: *COL.

Type: Colombia, Luer et al. 9053 (holotype, SEL).

Stelis felix Luer & R.Escobar, Harvard Pap. Bot. 23(1): 28. 2018.

Illustration: Luer 2018a, f. 15.

Distribution: *COL.

Type: Colombia, Escobar 5014 (holotype, MO).

Stelis fendleri Lindl., Fol. Orchid. ~Stelis~ 3. 1852- 1855 [1859].

Illustration: Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1018.

Distribution: COL VEN.

Type: Venezuela, Fendler 1470 (holotype, K-000573867; isotypes, AMES-00104755, BR0000006593766, G-00169082, G-00169083, GH-00104754, MO-1889091, PH-00027781, P-00489281).

Voucher: Colombia, Duque 1720 (JAUM, MO).

Stelis fissurosa Luer & R.Escobar, Harvard Pap. Bot. 22(1): 42. 2017.

Illustration: Luer 2017a, f. 27.

Distribution: *COL.

Type: Colombia, de Wilde 058 (holotype, SEL).

Stelis flacca Rchb.f., Bonplandia (Hannover) 3: 240. 1855.

Illustration: Luer 2009a, f. 316; Luer 2018c, f. 13-16.

Distribution: COL ECU.

Type: Ecuador, Jameson 735 (holotype, K; isotypes, BM, BR,G, LE, W).

Voucher: Colombia, Luer et al. 8803 (SEL).

Stelis flagellifera Luer & R.Escobar, Harvard Pap. Bot. 21(1): 70. 2016.

Illustration: Luer 2016a, f. 18.

Distribution: *COL.

Type: Colombia, Luer et al. 3719 (holotype, SEL).

Stelis flexuella Luer & R.Escobar, Harvard Pap. Bot. 21(1): 70. 2016.

Illustration: Luer 2016a, f. 19.

Distribution: *COL.

Type: Colombia, Luer et al. 4514 (holotype, SEL).

Stelis floresii Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 47. 2002.

Illustration: Luer 2002a, f. 58, f. 66.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 11139 (holotype, MO).

Voucher: Colombia, Salgado-Negret 60286 (COL).

Stelis floripendens O.Duque, Orquideología 27(2): 119-120. 2010.

Illustration: Duque 2010b, f. 12, ph. 31-34.

Distribution: *COL.

Type: Colombia, Duque 2165 (holotype, JAUM).

Stelis fluxflorum Luer & R.Escobar, Harvard Pap. Bot. 22(1): 42. 2017.

Illustration: Luer 2017a, f. 28.

Distribution: *COL.

Type: Colombia, Luer et al. 8876 (holotype, SEL).

Stelis foetida O.Duque, Orquideología 20(3): 342-343. 1997.

Illustration: Duque 1997, f. 4 (p. 345), ph. 4-6 (pl. III); Luer 2002a, f. 97; Luer 2018a, f. 16; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1760 (holotype, JAUM).

Stelis fons-florum (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Crocodeilanthe fonsflorum (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia fons-florum (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis fons-florum Lindl., Fol. Orchid. ~Pleurothallis~ 5, no. 15. 1859.

Illustration: Luer 1998a, f. 21.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Callejas 11330 (HUA).

Stelis formiciferaKarremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe rectangularis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 131-132. 2020. Non Stelis rectangularis Luer & R.Escobar [2016].

Illustration: Kolanowska & Szlachetko 2020b, f. 31.

Distribution: *COL.

Type: Colombia, Garcia-Barriga 18767 (holotype, AMES-112711; drawing, UGDA).

Stelis fornix Luer & R.Escobar, Harvard Pap. Bot. 21(1): 70. 2016.

Illustration: Luer 2016a, f. 20.

Distribution: *COL.

Type: Colombia, Luer et al. 16802 (holotype, MO).

Stelis fosteri (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe fosteri Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 117-118. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 15.

Distribution: *COL.

Type: Colombia, Foster 2055 (holotype, AMES116327; drawing, UGDA).

Stelis franciscana O.Duque, Orquideología 20(3): 343, 346-347. 1997.

Illustration: Duque 1997, f. 1 (p. 350), ph. 7-9 (pl. III); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1890 (holotype, JAUM).

Stelis fredoniensis O.Duque, Orquideología 27(2): 120, 123. 2010.

Illustration: Duque 2010b, f. 13, ph. 35-38.

Distribution: *COL.

Type: Colombia, Duque 2281 (holotype, JAUM).

Stelis frondifera Luer & R.Escobar, Harvard Pap. Bot. 23(1): 45. 2018. Stelis frondifera Luer & R.Escobar, Harvard Pap. Bot. 22(1): 44. 2017, nom. inval.

Illustration: Luer 2017a, f. 29, 30.

Distribution: *COL.

Type: Colombia, Luer et al. 10051 (holotype, SEL).

Stelis frontinoënsis O.Duque, Orquideología 20(3): 347-348. 1997.

Illustration: Duque 1997, f. 3 (p. 350), ph. 1-3 (pl. IV); Luer 2004, f. 127 (p. 139); Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Colombia, Duque 1891 (holotype, JAUM).

Stelis furfuracea F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 447. 1898.

Illustration: Luer 2018a, f. 17; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Lehmann 6253 [holotype, B; lectotype, K (Luer 2018a); isolectotype, LE].

Stelis fusilifera Luer & R.Escobar, Harvard Pap. Bot. 22(2): 92. 2017.

Illustration: Luer 2017b, f. 19.

Distribution: *COL.

Type: Colombia, Luer et al. 3139 (holotype, SEL).

Stelis fusisepala (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe fusisepala Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 118-119. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 16.

Distribution: *COL.

Type: Colombia, Garay et al. 533 (holotype, AMES; drawing, UGDA).

Stelis galapagosensis Luer & R.Escobar, Harvard Pap. Bot. 21(1): 72. 2016.

Illustration: Luer 2016a, f. 21.

Distribution: *COL.

Type: Colombia, Luer et al. 16816 (holotype, MO).

Stelis galeata (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Crocodeilanthe galeata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia galeata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis galeata Lindl., Ann. Mag. Nat. Hist. 15: 107. 1845.

Illustration: Luer 1998a, f. 22, f. 23, f. 24.

Distribution: COL ECU PER VEN.

Type: Ecuador, Hartweg 1408 (holotype, K-L000674237; isotypes: G-00168953, K-000584205, K-000584206, LD-1221064, W-R-0053660; sketch, K-000584207).

Voucher: Colombia, Sander s.n. (K, AMES).

Stelis galerasensis (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Crocodeilanthe galerasensis (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis galerasensis Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 28, f. 26. 1998.

Illustration: Luer 1998a, f. 25.

Distribution: *COL.

Type: Colombia, Luer & Luer 3749 (holotype, SEL).

Stelis garciabarrigae (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe garciabarrigae Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 120-121. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 18.

Distribution: *COL.

Type: Colombia, Garcia-Barriga 20214 (holotype, AMES; drawing, UGDA).

Stelis gargantua Pridgeon & M.W.Chase, Lindleyana 17(2): 99. 2002. Crocodeilanthe gigas (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis gigas Luer & R.Escobar, Orquideología 20(1): 52. 1996. Stelis gigas (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001, nom. illeg. Non Stelis gigas Barb.Rodr. [1881].

Illustration: Luer & Escobar 1996, p. 55, p. 57 (below, right); Luer 1998a, f. 26. Fig. 14B.

Distribution: *COL.

Type: Colombia, Luer et al. 14152 (holotype, MO).

Stelis gelida (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Crocodeilanthe gelida (Lindl.) Carnevali & I.Ramírez, Smithsonian Contr. Bot. 100: 133. 2014. Humboldtia gelida (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Niphantha gelida (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007, nom. inval. Niphantha gelida (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 120: 154. 2010. Pleurothallis gelida Lindl., Edwards’s Bot. Reg. 27: Misc. 91. 1841. Specklinia gelida (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 260. 2004.

Illustration: Cogniaux 1896, f. 3 (p. 112); Luer 2000a, f. 17, f. 18 (p. 94); Stenzel 2004, f. 15.

Distribution: BEL BOL BRA COL COS CUB DOM ECU ELS HAI HON JAM MEX NIC PAN PER PUE USA.

Type: Jamaica, Loddiges s.n. (holotype, K).

Voucher: Colombia, Viveros 75a (HUQ).

Stelis gemmaGaray, Orquideología 4(2): 77. 1969.

Illustration: Garay 1969, p. 78, p. 82 (top right); Luer 2009a, f. 372; Ortiz & Uribe 2007 (CD Edition). Fig. 15F.

Distribution: COL ECU.

Type: Ecuador, Andre 506 (holotype, K).

Voucher: Colombia, Escobar 338 (AMES).

Stelis genychila Garay, Canad. J. Bot. 34: 354. 1956.

Illustration: Luer 2009a, f. 344. Fig. 14F.

Distribution: COL ECU.

Type: Colombia, von Sneidem 1095 (holotype, S).

Stelis gerontica Luer & R.Escobar, Harvard Pap. Bot. 22(2): 92. 2017.

Illustration: Luer 2017b, f. 20.

Distribution: *COL.

Type: Colombia, Escobar 5022 (holotype, SEL).

Stelis gigapetala Luer & R.Escobar, Harvard Pap. Bot. 22(2): 92. 2017.

Illustration: Luer 2017b, f. 21.

Distribution: *COL.

Type: Colombia, Luer et al. 8828 (holotype, SEL).

Stelis giraffina Luer & R.Escobar, Harvard Pap. Bot. 21(1): 72. 2016.

Illustration: Luer 2016a, f. 22.

Distribution: *COL.

Type: Colombia, Luer 17581 (holotype, MO).

Stelis gladiata Lindl., Fol. Orchid. ~Stelis~ 9, no. 61. 1852-1855 [1859].

Illustration: Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Ecuador, Jameson 99 (holotype, K-000573917).

Voucher: Colombia, Mutis 1743 (MA).

Stelis globiflora Rchb.f., Otia Bot. Hamburg. 1: 19, no. 74. 1878.

Illustration: Luer 2009a, f. 318; Ortiz & Medina 2012, f. 15.

Distribution: COL ECU.

Type: Ecuador, Lehmann s.n. (holotype, W).

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz & Medina (2012).

Stelis globosa Pridgeon & M.W.Chase, Lindleyana 17(2): 99. 2002. Crocodeilanthe popayanensis (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis popayanensis F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 438. 1898. Stelis popayanensis (F.Lehm. & Kraenzl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 265. 2001, nom. illeg. Non Stelis popayanensis F.Lehm. & Kraenzl. [1899].

Illustration: Luer 1998a, f. 49.

Distribution: *COL.

Type: Colombia, Lehmann 4514 (holotype, K).

Stelis glochochila Luer & R.Escobar, Harvard Pap. Bot. 21(2): 199. 2016.

Illustration: Luer 2016b, f. 13, f. 14.

Distribution: *COL.

Type: Colombia, Luer et al. 8962 (holotype, SEL).

Stelis glomifera Luer & R.Escobar, Harvard Pap. Bot. 22(1): 44. 2017.

Illustration: Luer 2017a, f. 31, 32.

Distribution: *COL.

Type: Colombia, Luer et al. 8977 (holotype, SEL).

Stelis gloriae O.Duque, Orquideología 27(2): 127-128. 2010.

Illustration: Duque 2010b, f. 15, ph. 41-43.

Distribution: *COL.

Type: Colombia, Duque 2483 (holotype, JAUM).

Stelis glossula Rchb.f., Gard. Chron. 1870(42): 1373. 1870. Apatostelis glossula (Rchb.f.) Garay, Bot. Mus. Leafl. 27(7-9): 188. 1979.

Illustration: Luer 2009a, f. 320; Misas Urreta 2005, p. 684; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL COS ECU ELS HON NIC PAN.

Type: Costa Rica, Veitch s.n. (holotype, W-R0020266!).

Voucher: Colombia, Misas Urreta 132 (HPUJ).

Stelis gongylophora Luer & R.Escobar, Harvard Pap. Bot. 22(1): 46. 2017.

Illustration: Luer 2017a, f. 33.

Distribution: *COL.

Type: Colombia, Luer et al. 13071 (holotype, SEL).

Stelis gracilis Ames, Orchidaceae 2: 266. 1908.

Distribution: BEL COL COS GUA HON MEX NIC PAN.

Type: Guatemala, von Tuerkheim 7681 (holotype, AMES).

Voucher: Colombia, Alzate 569 (FAUC).

Stelis graminea Lindl., Fol. Orchid. ~Stelis~ 11, no. 77. 1852-1855 [1859].

Illustration: Luer 2018a, f. 19.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Luer et al. 13174 (MO).

Stelis grandiflora Lindl., Companion Bot. Mag. 2: 353. 1836.

Illustration: Cogniaux 1896, t. 78 (IV); Luer 2009a, f. 394, f. 419, f. 419a; Luer 2018b, f. 54, f. 55; Luer 2018c, f. 17-20; Romero-González & Carnevalli 2000, p. 1009, p. 1034, p. 1057; Ossenbach & Jenny 2017, f. 18.

Distribution: BOL BRA COL ECU GUY L.A. PER SUR VEN.

Type: Brazil, Descourtilz s.n. [holotype, Descourtilz’s pl. 14, unpublished manuscript (reproduced in Ossenbach & Jenny 2017, f. 18)].

Voucher: Colombia, Duque 883 (JAUM).

Stelis gravida Luer & R.Escobar, Harvard Pap. Bot. 21(1): 73. 2016.

Illustration: Luer 2016a, f. 23, f. 24.

Distribution: *COL.

Type: Colombia, Luer et al. 10079 (holotype, MO).

Stelis guianensis Rolfe, Trans. Linn. Soc. London, Bot. 6: 59. 1901.

Illustration: Luer 2009a, f. 40, f. 41a (p. 211); Luer 2018b, f. 56; Romero-González & Carnevalli 2000, p. 1022.

Distribution: BOL COL ECU FRE GUY VEN.

Type: Venezuela, McCornnell & Quelch 703 (holotype, K).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition).

Stelis gustavoromeroi (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe gustavoromeroi Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 121-122. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 19.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 19842 (holotype, AMES-109279; drawing, UGDA).

Stelis hallii Lindl., J. Bot. (Hooker) 1: 12. 1834.

Illustration: Luer 2007, f. 291; Luer 2009a, f. 291a, f. 291b (p. 212).

Distribution: COL ECU PER.

Type: Colombia, Lehmann s.n. (holotype, W).

Stelis hansenacea Luer & R.Escobar, Harvard Pap. Bot. 21(1): 73. 2016.

Illustration: Luer 2016a, f. 25.

Distribution: *COL.

Type: Colombia, Luer et al. 16796 (holotype, MO).

Stelis hennisiana Schltr., Orchis 6: 64. 1912.

Distribution: *COL.

Type: Colombia, Hennis s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis hercules Luer & R.Escobar, Harvard Pap. Bot. 21(2): 199. 2016.

Illustration: Luer 2016b, f. 15.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8360 (holotype, SEL).

Stelis heteroarcuata J.M.H.Shaw, Orchid Rev. 122(1308): 77. 2014. Stelis arcuata O.Duque, Orquideología 27(1): 12. 2010, nom. illeg. Non Stelis arcuata (Lindl.) Pridgeon & M.W.Chase [2001].

Illustration: Duque 2010a, f. 4, ph. 9-11.

Distribution: *COL.

Type: Colombia, Duque 2450 (holotype, JAUM).

Stelis hiatilabia Luer & R.Escobar, Harvard Pap. Bot. 21(2): 201. 2016.

Illustration: Luer 2016b, f. 16.

Distribution: *COL.

Type: Colombia, Luer et al. 7501 (holotype, SEL).

Stelis hippocrepica Luer & R.Escobar, Harvard Pap. Bot. 21(1): 73. 2016.

Illustration: Luer 2016a, f. 26.

Distribution: *COL.

Type: Colombia, Luer 11327 (holotype, MO).

Stelis hirsutaGaray, Bot. Mus. Leafl. 27(7-9): 184. 1979. Stelis jamesonii var. parviflora Garay, Canad. J. Bot. 34: 352. 1956.

Illustration: Luer 2018a, f. 20.

Distribution: *COL.

Type: Colombia, Grant & Fosburg 9362 (holotype, AMES; isotypes, COL, US, WS).

Stelis hoppii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 29. 1924.

Distribution: *COL.

Type: Colombia, Hopp 123 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis huilensis O.Duque, Orquideología 27(2): 128, 131. 2010.

Illustration: Duque 2010b, f. 16, ph. 44-45.

Distribution: *COL.

Type: Colombia, Duque 2160 (holotype, JAUM).

Stelis humidensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 46. 2017.

Illustration: Luer 2017a, f. 34. Fig. 15L.

Distribution: *COL.

Type: Colombia, Luer 17575 (holotype, SEL).

Stelis hyacinthalis Luer & R.Escobar, Harvard Pap. Bot. 22(2): 94. 2017.

Illustration: Luer 2017b, f. 22.

Distribution: *COL.

Type: Colombia, Luer et al. 7509 (holotype, SEL).

Stelis hylophila Rchb.f., Bonplandia (Hannover) 3: 241. 1855. Apatostelis hylophila (Rchb.f.) Garay, Bot. Mus. Leafl. 27: 189. 1979.

Illustration: Luer 2009a, f. 376; Luer 2018b, f. 59, f. 60; Ortiz & Uribe 2007 (CD Edition); RomeroGonzález & Carnevalli 2000, p. 1024.

Distribution: BOL COL ECU PER VEN.

Type: Ecuador, Jameson 687 (holotype, W; isotypes, BR, G, K, LE).

Voucher: Colombia, Duque 1715 (JAUM, MO).

Stelis hymenantha Schltr., Repert. Spec. Nov. Regni Veg. 10(251-253): 291. 1912.

Illustration: Luer 2009a, f. 377.

Distribution: BRA COL COS ECU GUA HON MEX NIC PAN.

Type: Guatemala, von Türckheim II-1968 [holotype, B, destroyed; lectotype, AMES-55238, photo and illustr. of the type (Luer 2009a); lectotype, AMES-33369, isotype (Luer 2023)].

Voucher: Colombia, Viveros 805 (HUQ).

Note: An isotype and original illustration at AMES have both been selected as lectotypes.

Stelis hypsela Luer, Harvard Pap. Bot. 22(2): 94. 2017.

Illustration: Luer 2017b, f. 23; Misas Urreta 2005, p. 708.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 4357 (holotype, MO; isotype, CUVC).

Stelis hypsitera Luer & R.Escobar, Harvard Pap. Bot. 21(2): 201. 2016.

Illustration: Luer 2016b, f. 17.

Distribution: *COL.

Type: Colombia, Luer 17571 (holotype, MO).

Stelis immersa (Linden & Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Effusiella immersa (Linden & Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Humboldtia immersa (Linden & Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis immersa Linden & Rchb.f., Bonplandia (Hannover) 3: 224. 1855. Specklinia immersa (Linden & Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004.

Illustration: Barringer 1986, f. 12; Luer 2000a, f. 20 (p. 94).

Distribution: COL COS GUA HON MEX NIC PAN VEN.

Type: Colombia, Linden s.n. [lectotype (selected here), K-L-000079814; type sketch, W-R0053656].

Note: Luer annotated the sheet W-R-0053656 as holotype. This sheet consists of an illustration, a rudimentary sketch with two floral segments glued on, and a loose inflorescence apex. The illustration was prepared long after the original description, and it is unclear if the inflorescence and floral segments were added later too. The sketch seems original. Nevertheless, this cannot be the holotype as Reichenbach clearly must have had actual plant material at hand given that the isotype, bearing his writting, at K-L includes a leaft and two full inflorescences with a flower each.

Stelis imperiosa Luer & R.Escobar, Harvard Pap. Bot. 23(1): 45. 2018. Stelis imperialis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 46. 2017, nom. inval.

Illustration: Luer 2017a, f. 35, 36.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8380 (holotype, SEL).

Stelis imraei (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Specklinia imraei (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004. Effusiella imraei (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Humboldtia imraei (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis imraei Lindl., Fol. Orchid. ~Pleurothallis~ 9. 1859.

Illustration: Luer 2000a, f. 21 (p. 95); Luer 1975b, f. 30.

Distribution: BOL COL COS DOM ECU GUY L.A. PAN PER VEN.

Type: Dominica, Imray 272 (K-L-000079856).

Voucher: Colombia, Luer 17549 (MO).

Stelis inamoena Luer & R.Escobar, Harvard Pap. Bot. 22(1): 48. 2017.

Illustration: Luer 2017a, f. 37.

Distribution: *COL.

Type: Colombia, Luer et al. 6653 (holotype, SEL).

Stelis incertaKarremans, Phytotaxa 496(2): 137. 2021. Crocodeilanthe undulata Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 139-140. 2020. Non Stelis undulata Luer & Hirtz [2002].

Illustration: Kolanowska & Szlachetko 2020b, f. 40.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 20692 (holotype, AMES-02084705; drawing, UGDA).

Stelis inclinata O.Duque, Orquideología 20(3): 349-351. 1997.

Illustration: Duque 1997, f. 4 (p. 350), ph. 6-8 (pl. IV); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1914 (holotype, JAUM).

Stelis index Luer & R.Escobar, Harvard Pap. Bot. 21(1): 76. 2016.

Illustration: Luer 2016a, f. 27.

Distribution: *COL.

Type: Colombia, Luer et al. 16805 (holotype, MO).

Stelis infundibulosa (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 263. 2001. Crocodeilanthe infundibulosa (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis infundibulosa Luer, Orquideología 20: 210. 1996.

Illustration: Luer 1996f, 211; Luer 1998a, f. 27.

Distribution: *COL.

Type: Colombia, Fassett 25732 (holotype, US).

Stelis intermedia Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 46, t. 79. 1836.

Illustration: Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1025.

Distribution: BRA COL GUY PER SUR VEN.

Type: Peru, Poeppig 1579 [lectotype, W-0010637 (selected here); isolectotypes, W-0010636, W-0047804, W-0047805, W-0047806, LE00001397, G-00169060, G-00169061, GOET-008692, AMES; type illustr. W-0076437].

Voucher: Colombia, Rivera OAD 1956 (COL).

Stelis ionantha Luer & R.Escobar, Harvard Pap. Bot. 21(1): 76. 2016.

Illustration: Luer 2016a, f. 28.

Distribution: *COL.

Type: Colombia, Luer et al. 14159 (holotype, MO).

Stelis ipialesensis Luer & Hirtz, Harvard Pap. Bot. 22(1): 50. 2017.

Illustration: Luer 2017a, f. 39.

Distribution: *COL.

Type: Colombia, Luer et al. 2738 (holotype, SEL).

Stelis jamesonii Lindl., J. Bot. (Hooker) 1: 11. 1834.

Illustration: Luer 2009a, f. 378, f. 378a; Luer 2018c, f. 21; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Cuatrecasas 18794 (AMES, NY, US).

Stelis jaramilloi (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 139. 2021. Crocodeilanthe jaramilloi Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 122. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 20.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 19831 (holotype, AMES; drawing, UGDA).

Stelis jorgei Luer, Harvard Pap. Bot. 22(2): 94. 2017.

Illustration: Luer 2017b, f. 24.

Distribution: *COL.

Type: Colombia, Giraldo-Gensini & Agredo 558 (holotype, MO; isotype, TULV).

Stelis jurisdicciensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 50. 2017.

Illustration: Luer 2017a, f. 40.

Distribution: *COL.

Type: Colombia, Luer et al. 6655 (holotype, SEL).

Stelis jurisdixii (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Crocodeilanthe jurisdixii (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis jurisdixii Luer & R.Escobar, Orquideología 20: 64. 1996.

Illustration: Luer & Escobar 1996, p. 65; Luer 1998a, f. 29.

Distribution: *COL.

Type: Colombia, Luer et al. 10211 (holotype, MO).

Stelis kefersteiniana (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 17(2): 99. 2002. Humboldtia kefersteiniana (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis kefersteiniana Rchb.f., Bot. Zeitung (Berlin) 10: 673. 1852.

Illustration: Luer 2000a, f. 13, f. 14, f. 15 (p. 93); Luer 1975b, f. 27.

Distribution: BOL COL ECU PER VEN.

Type: Venezuela, Wagener s.n. [L560] (holotype, W-R-0025692).

Voucher: Colombia, Hodge 6801 (AMES).

Stelis killipii (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 139. 2021. Crocodeilanthe killipii Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 122-123. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 21.

Distribution: *COL.

Type: Colombia, Killip & Smith 19430 (holotype, AMES-48055; drawing, UGDA).

Stelis kunturmarkaeKarremans, Phytotaxa 496(2): 139. 2021. Crocodeilanthe cundinamarcae Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 115-116. 2020. Non Stelis cundinamarcae Schltr. [1924].

Illustration: Kolanowska & Szlachetko 2020b, f. 12.

Distribution: *COL.

Type: Colombia, Catrecasas 5712 (holotype, AMES-59093; drawing, UGDA).

Stelis lacertina Luer & R.Escobar, Harvard Pap. Bot. 21(2): 203. 2016.

Illustration: Luer 2016b, f. 18.

Distribution: *COL.

Type: Colombia, Luer 16761 (holotype, MO).

Stelis laevigata (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Crocodeilanthe laevigata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia laevigata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis laevigata Lindl., Ann. Mag. Nat. Hist. 15: 106. 1845.

Illustration: Luer 1998a, f. 30, f. 31.

Distribution: COL ECU PER.

Type: Colombia, Hartweg 1406 (holotype, K-L; isotypes, AMES-00074378, AMES-00104094, BM-000074409, BR-0000008434876, BR0000008434883, G, K-000674238, K-000584209, K-000584208, LE-00001417, LD-1212858, P-00485529, P00485528, W-R, W-R-0053653, W-R-0053632; type illustr., K-000584210).

Stelis lagarantha Luer & R.Escobar, Harvard Pap. Bot. 21(2): 203. 2016.

Illustration: Luer 2016b, f. 19.

Distribution: *COL.

Type: Colombia, Luer et al. 10146 (holotype, MO).

Stelis lalinensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 50. 2017.

Illustration: Luer 2017a, f. 41.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8492 (holotype, SEL).

Stelis lamellata Lindl., Companion Bot. Mag. 2: 353. 1836.

Illustration: Luer 2009a, f. 380-380b.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K; isotype, W).

Voucher: Colombia, Luer et al. 13844 (MO).

Stelis lanceolata (Ruiz & Pav.) Willd., Sp. Pl. 4: 139. 1805. Humboldtia lanceolata Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 237. 1798.

Illustration: Luer 2009a, f. 383; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1026. Fig. 14K.

Distribution: COL ECU PER VEN.

Type: Peru, Pavon s.n. (holotype, MA).

Voucher: Colombia, Serralde 81 (COL).

Stelis lancifera Luer & R.Escobar, Harvard Pap. Bot. 21(1): 77. 2016.

Illustration: Luer 2016a, f. 29.

Distribution: *COL.

Type: Colombia, Luer & Luer 3733 (holotype, SEL).

Stelis lancipetala (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 139. 2021. Crocodeilanthe lancipetala Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 123-124. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 22.

Distribution: *COL.

Type: Colombia, Vogel 075A (holotype, AMES; drawing, UGDA).

Stelis langlassei Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 89. 1920.

Illustration: Luer 2009a, f. 323-323b; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Colombia, Langlassé 101 [holotype, B, destroyed; lectotype (Luer 2009a), K; isolectotypes, AMES, G].

Stelis lankesteri Ames, Sched. Orch. 3: 4-5. 1923.

Illustration: Ortiz & Uribe 2007 (CD Edition).

Distribution: COL COS PAN.

Type: Costa Rica, Lankester 365 (holotype, AMES-22847).

Voucher: Colombia, Ortiz-Valdivieso 94 (HPUJ).

Stelis laplanadensis Luer & R.Escobar, Harvard Pap. Bot. 21(2): 203. 2016.

Illustration: Luer 2016b, f. 20.

Distribution: *COL.

Type: Colombia, Luer 12515 (holotype, MO).

Stelis lappacea Luer & Teague, Harvard Pap. Bot. 21(1): 77. 2016.

Illustration: Luer 2016a, f. 30.

Distribution: *COL.

Type: Colombia, Luer 13224 (holotype, MO).

Stelis latisepala C.Schweinf., Bot. Mus. Leafl. 19: 199. 1961.

Illustration: Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1023.

Distribution: COL VEN.

Type: Venezuela, Steyermark 75712 (holotype, AMES-69505).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition) and cited by Duque 2010.

Stelis lehmanneptis (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Effusiella lehmanneptis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 106. 2007. Pleurothallis lehmanneptis Luer & R.Escobar, Orquideología 21: 100. 1998. Specklinia lehmanneptis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004.

Illustration: Luer & Escobar 1998, p. 101; Luer 2000a, f. 23 (p. 95).

Distribution: *COL.

Type: Colombia, Luer 18449 (holotype, MO).

Stelis lehmanniana (Schltr.) Karremans, Phytotaxa 203(3): 293. 2015. Crocodeilanthe lehmanniana (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis endotrachys F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 439. 1899, nom. illeg. Pleurothallis lehmanniana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 235. 1920. Non Pleurothallis endotrachys Rchb.f. [1876].

Illustration: Luer 1998a, f. 34.

Distribution: COL ECU.

Type: Colombia, Lehmann 4510 (holotype, B, destroyed). Colombia, Luer 12552 [neotype, MO (Luer 1998a)].

Stelis lehmannii Pridgeon & M.W.Chase, Lindleyana 17(2): 99. 2002. Effusiella petiolaris (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007. Pleurothallis petiolaris Luer, Orquideología 20: 220. 1996. Specklinia petiolaris (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004. Stelis petiolaris (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 265. 2001, nom. illeg. Non Stelis petiolaris Schltr. [1924].

Illustration: Luer 1996f, 221; Luer 2000a, f. 30 (p. 97).

Distribution: *COL.

Type: Colombia, Lehmann 9099 (holotype, K).

Stelis lentiginosa Lindl., Fol. Orchid. ~Stelis~ 8, no. 58. 1852-1855 [1859].

Illustration: Luer 2009a, f. 384; Luer 2018c, f. 22; Ortiz & Uribe 2007 (CD Edition); RomeroGonzález & Carnevalli 2000, p. 1028. Fig. 15G.

Distribution: COL ECU PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Schneider Wc 126 (AMES).

Stelis leprina Luer & R.Escobar, Harvard Pap. Bot. 21(2): 205. 2016.

Illustration: Luer 2016b, f. 21.

Distribution: *COL.

Type: Colombia, de Wilde 005 (holotype, MO).

Stelis leptorhiza Luer & R.Escobar, Harvard Pap. Bot. 21(2): 205. 2016.

Illustration: Luer 2016b, f. 22.

Distribution: *COL.

Type: Colombia, Luer 16919 (holotype, MO).

Stelis lesliegarayiKarremans, Phytotaxa 496(2): 139. 2021. Crocodeilanthe garayi Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 119-120. 2020. Non Stelis garayi (Dunst.) Carnevali & I. Ramírez [1990].

Illustration: Kolanowska & Szlachetko 2020b, f. 17.

Distribution: *COL.

Type: Colombia, Garay et al. 467 (holotype, AMES; drawing, UGDA).

Stelis liberalis Luer & J. Portilla, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 159, 186. 2004.

Illustration: Luer 2004, f. 166 (p. 186); Luer 2018c, f. 23, f. 24. Fig. 15E.

Distribution: COL ECU.

Type: Ecuador, Luer 19734 (holotype, MO).

Voucher: Colombia, Luer 17589 (MO).

Stelis ligulata (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Crocodeilanthe ligulata (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia ligulata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis ligulata Lindl., Fol. Orchid. ~Pleurothallis~ 29. 1859.

Illustration: Luer 1998a, f. 35, f. 36, f. 37. Fig. 15A.

Distribution: BOL COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Luer et al. 6658 (SEL).

Stelis lijiae Luer & R.Escobar, Harvard Pap. Bot. 21(1): 77. 2016.

Illustration: Luer 2016a, f. 31.

Distribution: *COL.

Type: Colombia, Luer 17602 (holotype, MO).

Stelis lindenii Lindl., Orchid. Linden. 3, no. 17. 1846.

Illustration: Luer 2009b, f. 39; Luer 2018a, f. 21- 23; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Venezuela, Linden 679 (holotype, K).

Voucher: Colombia, Silverstone-Sopkin 8000 (CUVC, MO).

Stelis londonnii O.Duque, Orquideología 20(3): 352-353. 1997.

Illustration: Duque 1997, f. 1 (p. 363), ph. 1-3 (pl. V); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1424 (holotype, JAUM).

Stelis longipetala O.Duque, Orquideología 20(3): 353-355. 1997.

Illustration: Duque 1997, f. 2 (p. 363), ph. 4-6 (pl. V); Luer 2018a, f. 24; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1903 (holotype, JAUM).

Stelis longiracemosa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 30. 1924.

Distribution: *COL.

Type: Colombia, Hopp 118 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis longirepens Carnevali & J.L.Tapia, Orchids Venez. (ed. 2) 1146. 2000.

Illustration: Luer 2018c, f. 25; Romero-González & Carnevalli 2000, p. 1029.

Distribution: COL VEN.

Type: Venezuela, Diaz 125 (holotype, VEN; isotype, PORT).

Voucher: Colombia, Luer 17604 (MO).

Stelis lopezii Luer, Selbyana 30(1): 22-23. 2009.

Illustration: Luer 2009b, f. 43.

Distribution: *COL.

Type: Colombia, Lopez Figueiras 8391 (holotype, COL).

Stelis loxensis Lindl., Pl. Hartw. 16: 3. 1953.

Illustration: Luer 2009a, f. 386; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU.

Type: Ecuador, Hartweg 38 (holotype, K).

Voucher: Colombia, Duque 2427 (JAUM).

Stelis luisii Luer, Harvard Pap. Bot. 22(2): 96. 2017.

Illustration: Luer 2017b, f. 25.

Distribution: *COL.

Type: Colombia, Giraldo-Gensini & Agredo 425 (holotype, MO; isotype, TULV).

Stelis lumbricosa O.Duque, Orquideologia 20: 354. 1997.

Illustration: Duque 1997, p. 363, ph. 7-9 (pl. V); Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 585 (holotype, JAUM).

Stelis lutea Lindl., Fol. Orchid. ~Stelis~ 7. 1852-1855 [1859].

Illustration: Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Venezuela, Fendler 1461 (holotype, K-000573907).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition).

Stelis lynnianaLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 5. 2007.

Illustration: Luer 2007, f. 208; Luer 2018c, f. 26. Fig. 15C.

Distribution: COL ECU.

Type: Ecuador, O’Shaughnessy 02856 (holotype, MO).

Voucher: Colombia, Luer 18439 (MO).

Stelis macropoda Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 89. 1920.

Illustration: Luer 2018a, f. 25-26.

Distribution: *COL.

Type: Colombia, Lehmann 4519 [holotype, B, destroyed; lectotype (selected here), AMES00287416; isolectotype, K-000573945].

Stelis maderoi Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 90. 1920.

Illustration: Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1030.

Distribution: COL VEN.

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 30, No. 114)].

Stelis magdalenae (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Crocodeilanthe magdalenae (Rchb.f.) Toscano, Harvard Pap. Bot. 23(1): 54. 2018. Humboldtia magdalenae (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis magdalenae Rchb.f., Bonplandia (Hannover) 3: 72. 1855.

Illustration: Luer 1999a, f. 3 (p. 94).

Distribution: BOL COL ECU.

Type: Colombia, Wagener s.n. (holotype, W).

Stelis magnesialis Luer & Hirtz, Harvard Pap. Bot. 22(1): 51. 2017.

Illustration: Luer 2017a, f. 42.

Distribution: *COL.

Type: Colombia, Luer et al. 4520 (holotype, SEL).

Stelis magnipetala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 31. 1924. Apatostelis magnipetala (Schltr.) Garay, Bot. Mus. Leafl. 27: 189. 1979.

Distribution: *COL.

Type: Colombia, Hopp 47 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis major Rchb.f., Bonplandia (Hannover) 2: 23. 1854.

Illustration: Luer 2018a, f. 27-28; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1031.

Distribution: COL VEN.

Type: Venezuela, Wagener s.n. (holotype, K).

Voucher: Colombia, St. John 20879 (US, AMES).

Stelis majuscula Luer, Harvard Pap. Bot. 22(1): 51. 2017.

Illustration: Luer 2017a, f. 43.

Distribution: *COL.

Type: Colombia, Uribe 385 (holotype, AMWS).

Stelis mantae (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe mantae Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 124-125. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 23.

Distribution: *COL.

Type: Colombia, Mora 3085 (holotype, AMES106118; drawing, UGDA).

Stelis marginata Luer & R.Escobar, Harvard Pap. Bot. 21(2): 205. 2016.

Illustration: Luer 2016b, f. 23.

Distribution: *COL.

Type: Colombia, Luer et al. 8359 (holotype, SEL).

Stelis maxima Lindl., Ann. Mag. Nat. Hist. 15(96): 106-107. 1845.

Illustration: Luer 2009a, f. 324; Luer 2018c, f. 27; Ortiz & Uribe 2007 (CD Edition); RomeroGonzález & Carnevalli 2000, p. 1032.

Distribution: COL ECU.

Type: Ecuador, Hartweg 1407 (holotype, K; isotypes: G, LD, LE, W).

Voucher: Colombia, Cuatrecasas 20912 (AMES).

Stelis meerenbergensis Luer & R.Escobar, Harvard Pap. Bot. 22(2): 96. 2017.

Illustration: Luer 2017b, f. 26.

Distribution: *COL.

Type: Colombia, Luer et al. 8485 (holotype, SEL).

Stelis megaloglossa Luer, Phytologia 49(3): 231. 1981.

Illustration: Luer 2007, f. 215; Luer 2018c, f. 28.

Distribution: COL ECU.

Type: Colombia, Luer et al. 4528 (holotype, SEL).

Stelis melanopus (F.Lehm. & Kraenzl.) Karremans, Lankesteriana 13(3): 329. 2014. Pleurothallis melanopus F.Lehm. & Kraenzl., Bot. Jaarb. 26: 443. 1899.

Illustration: Luer 1999a, f. 30 (p. 133).

Distribution: BOL COL.

Type: Unknown, Lehmann s.n. (holotype, ?).

Voucher: Colombia, Lehmann 4515 (K, US).

Stelis menippe Luer & R.Escobar, Harvard Pap. Bot. 21(2): 206. 2016.

Illustration: Luer 2016b, f. 24.

Distribution: *COL.

Type: Colombia, Luer et al. 9025 (holotype, SEL).

Stelis meridana (Rchb.f.) Karremans, Lankesteriana 13(3): 329. 2014. Anathallis meridana (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 249. 2001. Humboldtia meridana (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis meridana Rchb.f., Linnaea 22: 826. 1849. Specklinia meridana (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 1999a, f. 18 (p. 130).

Distribution: COL VEN.

Type: Venezuela, Moritz 1063 [lectotype, W (Luer 1999a)].

Voucher: Colombia, Cuatrecasas 25327 (US).

Stelis mesohybos Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 155. 1924.

Distribution: *COL.

Type: Colombia, Hopf s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis microchila Schltr., Repert. Spec. Nov. Regni Veg. 9(214-216): 289. 1911.

Illustration: Luer 2009a, f. 391, f. 391a; Misas Urreta 2005, p. 676.

Distribution: BEL COL COS ELS GUA MEX PAN.

Type: Guatemala, von Türckheim 700 (holotype, B, destroyed; lectotype, AMES-22591 (Luer 2009a); isolectotypes, BR, G; lectotype, AMES-72173 (Luer 2023)].

Voucher: Colombia, Misas Urreta 125 (HPUJ).

Note: An isotype and original illustration at AMES have both been selected as lectotypes.

Stelis microphylla Lindl., Fol. Orchid. ~Stelis~ 10, no. 74. 1852-1855 [1859].

Illustration: Luer 2018a, f. 29.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Lehmann 6039 (AMES).

Note: We agree with Luer 2018a rather than Luer 2018b.

Stelis miranda Luer & R.Escobar, Harvard Pap. Bot. 21(2): 206. 2016.

Illustration: Luer 2016b, f. 25.

Distribution: *COL.

Type: Colombia, Luer et al. 7633 (holotype, SEL).

Stelis mocoana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 33. 1924.

Distribution: *COL.

Type: Colombia, Hopp 159 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis molecula Luer & R.Escobar, Harvard Pap. Bot. 21(2): 208. 2016.

Illustration: Luer 2016b, f. 26.

Distribution: *COL.

Type: Colombia, Luer et al. 10218 (holotype, MO).

Stelis monetaria Luer & R.Escobar, Harvard Pap. Bot. 21(1): 79. 2016.

Illustration: Luer 2016a, f. 32.

Distribution: *COL.

Type: Colombia, Luer 16637 (holotype, MO).

Stelis montserratii (Porsch) Karremans, Lankesteriana 13(3): 329. 2014. Pleurothallis montserratii Porsch, Oesterr. Bot. Z. 158. 1905.

Illustration: Luer 1999a, f. 23 (p. 131); Loefgren 1918, pl. XXVI; Santos et al. 2019, f. 1o, f. 10.

Distribution: BOL BRA COL ECU PER.

Type: Brazil, Wettstein & Schniffner s.n. (holotype, W).

Voucher: Colombia, Uribe 4856 (AMES, COL).

Stelis mordica Luer & R.Escobar, Harvard Pap. Bot. 21(2): 208. 2016.

Illustration: Luer 2016b, f. 27.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8411 (holotype, SEL).

Stelis morganii Dodson & Garay, Icon. Pl. Trop. 4: t. 328. 1980.

Illustration: Luer 2009a, f. 393, f. 393a.

Distribution: COL ECU.

Type: Ecuador, Dodson et al. 7796 (holotype, SEL).

Stelis moritzii (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001. Crocodeilanthe moritzii (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia moritzii (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis moritzii Rchb.f., Linnaea 22: 824. 1849.

Illustration: Luer 1998a, f. 41; Kolanowska & Szlachetko 2020b, f. 1.

Distribution: COL ECU VEN.

Type: Venezuela, Moritz 854 (holotype, W).

Voucher: Colombia, Medina T. et al. 462 (JAUM).

Stelis morula Luer & R.Escobar, Harvard Pap. Bot. 22(1): 53. 2017.

Illustration: Luer 2017a, f. 44.

Distribution: *COL.

Type: Colombia, Luer et al. 12545 (holotype, SEL).

Stelis mucronipetala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 33. 1924.

Illustration: Luer 2018a, f. 30-31.

Distribution: *COL.

Type: Colombia, Hopp 135 (holotype, B, destroyed). Colombia, Luer 8496 [neotype, SEL (Luer 2018a)].

Stelis multirostris (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 17(2): 1000. 2002. Pleurothallis multirostris Rchb.f., Linnaea 41: 49. 1877.

Illustration: Luer 2000a, f. 34 (p. 98); Stenzel 2004, f. 32.

Distribution: COL COS CUB DOM ELS GUA HAI HON JAM MEX NIC PUE VEN.

Type: Jamaica, unknown collector (holotype, K; illustr., W).

Stelis muscosa Lindl., Fol. Orchid. ~Stelis~ 7, no. 48. 1852-1855 [1859].

Distribution: BOL BRA COL ECU GUY L.A. SUR VEN.

Type: Venezuela, Fendler 1468 (holotype, K-000573949).

Voucher: Colombia, Ospina 61-28 (JAUM).

Stelis mystrion Luer & R.Escobar, Harvard Pap. Bot. 21(1): 79. 2016.

Illustration: Luer 2016a, f. 33, f. 34.

Distribution: *COL.

Type: Colombia, Luer et al. 14180 (holotype, MO).

Stelis nemoralis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 53. 2017.

Illustration: Luer 2017a, f. 45.

Distribution: *COL.

Type: Colombia, Luer et al. 7689 (holotype, SEL).

Stelis nexipous Garay, Bot. Mus. Leafl. 18(5): 194. 1958.

Illustration: Luer 2009a, f. 346, f. 346a.

Distribution: COL ECU.

Type: Ecuador, Asplund 19638 (holotype, S; isotype, AMES).

Voucher: Colombia, Duque 1165 (JAUM).

Stelis nexosa Luer & R.Escobar, Harvard Pap. Bot. 21(2): 208. 2016.

Illustration: Luer 2016b, f. 28.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8491 (holotype, SEL).

Stelis niessen-andreae Luer, Harvard Pap. Bot. 22(2): 96. 2017.

Illustration: Luer 2017b, f. 27.

Distribution: *COL.

Type: Colombia, Luer 18432 (holotype, MO).

Stelis niesseniae (Luer) Karremans, Phytotaxa 406(5): 265. 2019. Effusiella niesseniae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007. Pleurothallis niesseniae Luer, Orquideología 22(1): 59-61. 2001. Specklinia niesseniae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 262. 2004.

Illustration: Luer 2001b, p. 61.

Distribution: *COL.

Type: Colombia, Niessen 121 (holotype, MO).

Stelis nitens Rchb.f., Bonplandia (Hannover) 2: 22. 1854.

Illustration: Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1036.

Distribution: COL ECU VEN.

Type: Venezuela, Wagener s.n. (holotype, W; illustr., AMES-00104852).

Voucher: Colombia, Wagener s.n. (W, AMES00287033).

Stelis nutans Lindl., Fol. Orchid. ~Stelis~ 15, no. 109. 1852-1855 [1859].

Illustration: Luer 2009a, f. 327, f. 327a; Luer 2018b, f. 83.

Distribution: BOL COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K; isotypes, W).

Voucher: Colombia, Rivas 35 (HUQ).

Stelis obescula Luer & R.Escobar, Harvard Pap. Bot. 22(1): 53. 2017.

Illustration: Luer 2017a, f. 46.

Distribution: *COL.

Type: Colombia, Luer et al. 7600 (holotype, SEL).

Stelis oblector Luer & R.Escobar, Harvard Pap. Bot. 21(2): 210. 2016.

Illustration: Luer 2016b, f. 29.

Distribution: *COL.

Type: Colombia, Luer et al. 3087 (holotype, SEL).

Stelis oblonga (Ruiz & Pav.) Willd., Sp. Pl. 4: 139. 1805. Humboldtia oblonga Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 236. 1798.

Illustration: Luer 2002a, f. 50; Luer 2009a, f. 50, f. 50a (p. 220); Luer 2018c, f. 31; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1038; Pupulin 2013, f. 48.

Distribution: COL ECU PER VEN.

Type: Peru, Pavón s.n. (holotype, MA; isotype, W).

Voucher: Colombia, Pittier 1299 (US-531497, AMES-25099).

Stelis oblongifolia Lindl., Fol. Orchid. ~Stelis~ 12, no. 89. 1852-1855 [1859].

Illustration: Luer 2009a, f. 396, f. 396a; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Stelis ocreosa Luer & R.Escobar, Harvard Pap. Bot. 21(2): 210. 2016.

Illustration: Luer 2016b, f. 30.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8391 (holotype, SEL).

Stelis octavioi Luer & R.Escobar, Harvard Pap. Bot. 22(2): 98. 2017.

Illustration: Luer 2017b, f. 28.

Distribution: *COL.

Type: Colombia, Luer et al. 3722 (holotype, SEL).

Stelis odontopetala Luer & Hirtz, Harvard Pap. Bot. 22(2): 98. 2017.

Illustration: Luer 2017b, f. 29.

Distribution: *COL.

Type: Colombia, Luer et al. 4549 (holotype, SEL).

Stelis oligoblephara Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 34. 1924.

Distribution: *COL.

Type: Colombia, Hopp 133 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis opercularis Luer, Phytologia 49(3): 233. 1981.

Illustration: Luer 2009a, f. 329.

Distribution: COL ECU VEN.

Type: Ecuador, Dodson et al. 5500 (holotype, SEL).

Stelis ophiodontodes Luer & R.Escobar, Harvard Pap. Bot. 21(2): 210. 2016.

Illustration: Luer 2016b, f. 31.

Distribution: *COL.

Type: Colombia, Escobar 5142 (holotype, SEL).

Stelis oreibator Luer & R.Escobar, Harvard Pap. Bot. 21(2): 211. 2016.

Illustration: Luer 2016b, f. 32.

Distribution: *COL.

Type: Colombia, Luer et al. 12549 (holotype, SEL).

Stelis orphana Luer, Harvard Pap. Bot. 22(2): 100. 2017.

Illustration: Luer 2017b, f. 30.

Distribution: *COL.

Type: Colombia, Luer 20217 (holotype, MO).

Stelis oscarii Luer, Harvard Pap. Bot. 22(2): 112. 2017. Stelis duquei Luer & R.Escobar, Harvard Pap. Bot. 21(1): 67. 2016, nom. illeg. Stelis occulta O.Duque, Orquideología 27(2): 131-132. 2010, nom. inval. Non Stelis duquei P.Ortiz [2011].

Illustration: Duque 2010b, f. 17, ph. 46-47; Luer 2016a, f. 14.

Distribution: *COL.

Type: Colombia, Luer 14328 (holotype, MO).

Stelis ospinae (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe ospinae Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 125-126. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 24.

Distribution: *COL.

Type: Colombia, Ospina-Hernández 70 (holotype, AMES-111319; drawing, UGDA).

Stelis otara Luer & R.Escobar, Harvard Pap. Bot. 21(2): 211. 2016.

Illustration: Luer 2016b, f. 33.

Distribution: *COL.

Type: Colombia, Luer et al. 8961A (holotype, SEL).

Stelis ovatipetala (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe ovatipetala Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 126. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 25.

Distribution: *COL.

Type: Colombia, Schneider 241 (holotype, AMES116222; drawing, UGDA).

Stelis oxysepala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 156. 1924.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis ozota Luer & R.Escobar, Harvard Pap. Bot. 21(2): 213. 2016.

Illustration: Luer 2016b, f. 34.

Distribution: *COL.

Type: Colombia, Luer et al. 3107 (holotype, SEL).

Stelis pachoi Luer & R.Escobar, Harvard Pap. Bot. 21(2): 213. 2016.

Illustration: Luer 2016b, f. 35.

Distribution: *COL.

Type: Colombia, Luer 17603 (holotype, MO).

Stelis pachyphyta Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 55. 2002.

Illustration: Luer 2002a, f. 74; Ortiz & Medina 2012, f. 16. Fig. 14J.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 10848 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz & Medina (2012).

Stelis pachypus F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 447. 1899. Crocodeilanthe pachypus (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis pachypus (F.Lehm. & Kraenzl.) Garay, Canad. J. Bot. 34: 254. 1956.

Illustration: Luer 1998a, f. 44.

Distribution: BOL COL ECU PER.

Type: Colombia, Lehmann 3176 [holotype, B, destroyed; lectotype (selected here), K-000584305; isolectotypes, AMES-00104870, G-00168935, BR0000006594787, NY-00009405, US00093578).

Stelis pachystachya Lindl., Fol. Orchid. ~Stelis~ 13. 1852-1855 [1859].

Illustration: Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Purdie s.n. (holotype, K-000573938).

Stelis pachystele Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 29. 1920.

Illustration: Luer 2018a, f. 32.

Distribution: *COL.

Type: Colombia, Lehmann 6922 [holotype, B, destroyed; lectotype (selected here), K-000573932; isolectotypes, K-000573933, GH-00104890, LE00001402, NY-00009406].

Stelis pachythrix Luer & R.Escobar, Harvard Pap. Bot. 21(2): 213. 2016.

Illustration: Luer 2016b, f. 36.

Distribution: *COL.

Type: Colombia, Luer et al. 10114 (holotype, SEL).

Stelis palifera Luer & R.Escobar, Harvard Pap. Bot. 22(1): 55. 2017.

Illustration: Luer 2017a, f. 47.

Distribution: *COL.

Type: Colombia, Luer et al. 7679 (holotype, SEL).

Stelis palimmeces Luer & R.Escobar, Harvard Pap. Bot. 21(1): 79. 2016.

Illustration: Luer 2016a, f. 35.

Distribution: *COL.

Type: Colombia, Luer et al. 16832a (holotype, MO).

Stelis papaquerensis Rchb.f., Linnaea 22: 822. 1849.

Illustration: Cogniaux 1896, t. 80 (I), t. 81 (IV, V); Misas Urreta 2005, p. 686.

Distribution: BRA COL FRE GUY L.A. SUR VEN.

Type: Brazil, Beyrich s.n. (holotype, W).

Voucher: Colombia, Misas Urreta 287 (HPUJ).

Stelis papilio O.Duque, Orquideología 20(3): 358, 361. 1997.

Illustration: Duque 1997, p. 363, ph. 1-3 (pl. VI); Luer 2018a, f. 33; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 419 (holotype, JAUM).

Stelis papiliopsis O.Duque, Orquideología 20(3): 361-362. 1997.

Illustration: Duque 1997, p. 367; ph. 4-6 (pl. VI); Luer 2018a, f. 33; Ortiz & Uribe 2007 (CD Edition).

Distribution: *COL.

Type: Colombia, Duque 1366 (holotype, JAUM).

Stelis papillipetala (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe papillipetala Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 127. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 26.

Distribution: *COL.

Type: Colombia, Schneider 304 (holotype, AMES111732; drawing, UGDA).

Stelis papposa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 80. 2016.

Illustration: Luer 2016a, f. 36.

Distribution: *COL.

Type: Colombia, Luer et al. 8755 (holotype, SEL).

Stelis paradoxa Luer & R.Escobar, Harvard Pap. Bot. 21(2): 215. 2016.

Illustration: Luer 2016b, f. 37.

Distribution: *COL.

Type: Colombia, Luer 14263 (holotype, SEL).

Stelis paraguasensis Luer, Selbyana 30(1): 23. 2009.

Illustration: Luer 2009b, f. 44; Luer 2018c, f. 33- 36.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin et al. 7955 (holotype, CUVC).

Stelis pardipes Rchb.f., Beitr. Orchid.-K. C. Amer. 96. 1866. Stelis triseta var. pardipes (Rchb.f.) C.Schweinf., Bot. Mus. Leafl. 15(1): 24-25. 1951.

Illustration: Luer 2009a, f. 330, f. 330a, f. 321, f. 321a; Luer 2018b, f. 90, f. 91; Luer 2018c, f. 37, f. 38; Ortiz & Uribe 2007 (CD Edition).

Distribution: BOL COL COS ECU ELS GUA HON MEX NIC PAN PER.

Type: Costa Rica, Wendland 899 (holotype, W).

Voucher: Colombia, Duque 1592 (JAUM).

Stelis parviflora (Ruiz & Pav.) Pers., Syn. Pl. 2: 524. 1807. Humboldtia parviflora Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 236. 1798.

Illustration: Pupulin 2013, f. 49.

Distribution: COL PER.

Type: Peru, Ruiz & Pavón s.n. (holotype, MA; isotype, BM-000082442).

Voucher: Colombia, Valencia et al. 383 (JBB).

Stelis parvilabris Lindl., Ann. Mag. Nat. Hist. 15: 385. 1845.

Illustration: Luer 2018a, f. 34; Ortiz & Uribe 2007 (CD Edition); Romero-González & Carnevalli 2000, p. 1039. Fig. 14E.

Distribution: COL VEN.

Type: Colombia, Hartweg s.n. (holotype, K).

Stelis pasminoi Luer, Harvard Pap. Bot. 22(2): 100. 2017.

Illustration: Luer 2017b, f. 31.

Distribution: *COL.

Type: Colombia, Pasmiño & Posso 31 (holotype, MO; isotype, PSO).

Stelis paulalleniiKarremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe allenii Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 107-108. 2020. Non Stelis allenii L.O.Williams [1942].

Illustration: Kolanowska & Szlachetko 2020b, f. 3.

Distribution: *COL.

Type: Colombia, Allen 3003 (holotype, AMES; drawing, UGDA).

Stelis pauxilla Luer & R.Escobar, Harvard Pap. Bot. 21(1): 80. 2016.

Illustration: Luer 2016a, f. 37.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8365 (holotype, MO).

Stelis pendularis O.Duque, Orquideología 27(2): 138, 141. 2010.

Illustration: Duque 2010b, f. 19, ph. 51-53.

Distribution: *COL.

Type: Colombia, Duque 2442 (holotype, JAUM).

Stelis pendulata O.Duque, Orquideología 27(2): 141-142. 2010.

Illustration: Duque 2010b, f. 20, ph. 54-56.

Distribution: *COL.

Type: Colombia, Duque 2280 (holotype, JAUM).

Stelis pennelliana (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 265. 2001. Crocodeilanthe pennelliana (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Pleurothallis pennelliana Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 43, f. 47. 1998.

Illustration: Luer 1998a, f. 47; Kolanowska & Szlachetko 2020b, f. 42.

Distribution: *COL.

Type: Colombia, Pennell 2043 (holotype, US; isotype, AMES).

Stelis perbona Luer & R.Escobar, Harvard Pap. Bot. 21(1): 82. 2016.

Illustration: Luer 2016a, f. 38.

Distribution: *COL.

Type: Colombia, Luer et al. 8901 (holotype, SEL).

Stelis pertenius Luer & R.Escobar, Harvard Pap. Bot. 21(1): 82. 2016.

Illustration: Luer 2016a, f. 39; Misas Urreta 2005, p. 702.

Distribution: *COL.

Type: Colombia, Luer de Wilde s.n. (holotype, MO).

Stelis petiolaris Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 36. 1924.

Distribution: *COL.

Type: Colombia, Hopp 48 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis pili-papillosa O.Duque, Orquideología 27(2): 142. 2010.

Illustration: Duque 2010b, f. 21, ph. 57-59.

Type: Colombia, Duque 1957 (holotype, JAUM).

Stelis pilicrepa Luer & R.Escobar, Harvard Pap. Bot. 22(1): 55. 2017.

Illustration: Luer 2017a, f. 48.

Distribution: *COL.

Type: Colombia, Luer et al. 7686 (holotype, SEL).

Stelis pilifera (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 265. 2001. Crocodeilanthe pilifera (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia pilifera (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pilifera Lindl., Fol. Orchid. ~Pleurothallis~ 9. 1859.

Illustration: Luer 1998a, f. 48.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Valoyes s.n. (IIAP).

Stelis pittieri (Schltr.) Rojas-Alv. & Karremans, Phytotaxa 406(5): 266. 2019. Pleurothallis pittieri Schltr., Repert. Spec. Nov. Regni Veg. 3(42-43): 247. 1907. Stelis pittieri Schltr. ex Knuth, Repert. Spec. Nov. Regni Veg. Beih. 43: 227. 1927, nom. inval.

Illustration: Luer 1998a, f. 20.

Distribution: BOL COL COS ECU L.A. PAN PER VEN.

Type: Costa Rica, Pittier 2067 (holotype, B, destroyed). Costa Rica, Pittier 2023 [neotype, AMES-23666 (Luer 1998a)].

Voucher: Colombia, Lehmann 3746 (BR).

Stelis platystachya Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 6: 424. 1976.

Illustration: Luer 2009a, f. 400; Romero-González & Carnevalli 2000, p. 1040.

Distribution: COL ECU VEN.

Type: Colombia, Foster 1774 (holotype, AMES; isotype, COL).

Stelis pleistantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 36. 1924. Apatostelis pleistantha (Schltr.) Garay, Bot. Mus. Leafl. 27: 190. 1979.

Illustration: Luer 2018a, f. 35.

Distribution: *COL.

Type: Colombia, Hopp 139 (holotype, B, destroyed).

Stelis pollerecta Luer & R.Escobar, Harvard Pap. Bot. 21(2): 215. 2016.

Illustration: Luer 2016b, f. 38.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8393 (holotype, SEL).

Stelis polyclada Lindl., Fol. Orchid. ~Stelis~ 10, no. 69. 1852-1855 [1859].

Illustration: Luer 2018a, f. 36.

Distribution: COL ECU.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Hopp 151 (B, destroyed).

Note: We agree with Luer 2018a rather than Luer 2018b.

Stelis popayanensis F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26(3-4): 448. 1899.

Illustration: Luer 2018a, f. 37.

Distribution: *COL.

Type: Colombia, Lehmann 6817 (holotype, K).

Stelis porpax Rchb.f., Bonplandia (Hannover) 2: 23. 1854.

Illustration: Luer 2009a, f. 402; Romero-González & Carnevalli 2000 (p. 1041); Ortiz & Uribe 2007 (CD Edition).

Distribution: COL ECU VEN.

Type: Venezuela, Wagener s.n. (holotype, W).

Voucher: Colombia, Ortiz 427 (HPUJ).

Stelis portillae Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 168, 190. 2004.

Illustration: Luer 2004, f. 183 (p. 190); Ortiz & Medina 2012, f. 17.

Distribution: COL ECU.

Type: Ecuador, Hirtz 8022 (holotype, MO).

Voucher: A photographic voucher of a Colombian specimen reproduced in Ortiz & Medina (2012).

Stelis posadarum Luer & R.Escobar, Harvard Pap. Bot. 22(2): 100. 2017.

Illustration: Luer 2017b, f. 32.

Distribution: *COL.

Type: Colombia, Luer 6544 (holotype, SEL).

Stelis possoae (Luer) Karremans, Phytotaxa 203(3): 293. 2015. Pleurothallis possoae Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 79: 129-130, f. 6. 2000. Crocodeilanthe possoae (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Illustration: Luer 2000a, f. 6 (p. 139).

Distribution: *COL.

Type: Colombia, Pasmiño & Posso 22 (holotype, PSO; isotype, MO).

Stelis powellii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 17: 16-17. 1922.

Distribution: COL COS HON NIC PAN.

Type: Panama, Powell 247 [holotype, B, destroyed; lectotype, AMES-23778 (Christenson 1991); isolectotype, MO].

Voucher: Colombia, Croat 49910 (MO).

Stelis praecipua Luer, Harvard Pap. Bot. 22(2): 101. 2017.

Illustration: Luer 2017b, f. 33.

Distribution: *COL.

Type: Colombia, Garay 37 (holotype, AMES).

Stelis preclara Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 21, f. 27. 2002.

Illustration: Luer 2002a, f. 27.

Distribution: COL ECU.

Type: Ecuador, Hirtz 1475 (holotype, MO).

Voucher: Colombia, Silverstone-Sopkin 7977 (COL).

Stelis prodigiosa Luer & R.Escobar, Harvard Pap. Bot. 22(1): 55. 2017.

Illustration: Luer 2017a, f. 49.

Distribution: *COL.

Type: Colombia, Luer et al. 7497 (holotype, SEL).

Stelis profusa O.Duque ex Flores-Rojas & Karremans, Phytotaxa 496(2): 141. 2021. Stelis profusa O.Duque, Orquideología 27(2): 145-146. 2010, nom. inval.

Illustration: Duque 2010b, f. 22, ph. 60-62; Karremans et al. 2021, f. 5.

Distribution: *COL.

Type: Colombia, Duque & Alzate 2325 (holotype, JAUM).

Stelis propinqua Ames, Sched. Orch. 6: 55-56. 1923.

Distribution: COL COS NIC.

Type: Costa Rica, Lankester 468 (holotype, AMES).

Voucher: Colombia, Duque 1023 (JAUM).

Stelis prorepens Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 92. 1920.

Distribution: *COL.

Type: Colombia, Lehmann 3520 [holotype, B, destroyed; lectotype (selected here), K-000573934; isolectotypes, AMES-00104935, BM-000084118, BR-00000013247744, NY-00009408].

Stelis pseudocheila (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Effusiella pseudocheila (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007. Pleurothallis pseudocheila Luer & R.Escobar, Orquideología 16(2): 173. 1984. Specklinia pseudocheila (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer & Escobar 1984b, p. 174; Luer 2000a, f. 33 (p. 98).

Distribution: *COL.

Type: Colombia, Luer 8186 (holotype, SEL).

Stelis pubescens (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 141. 2021. Crocodeilanthe pubescens Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 127-128. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 27.

Distribution: *COL.

Type: Colombia, Garcia-Barriga & Jaramillo 20692bis (holotype, AMES; drawing, UGDA).

Stelis pulchella Kunth, Nov. Gen. Sp. (quarto ed.) 1(4): 364, t. 90. 1816. Crocodeilanthe pulchella (Kunth) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia pulchella (Kunth) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis pulchella (Kunth) Lindl., Exot. Fl. 2(14): sub t. 123. 1825 [1824].

Illustration: Luer 1998a, f. 52.

Distribution: COL COS ECU PAN PER.

Type: Ecuador, Humboldt & Bonpland s.n. (holotype, P-00669694; isotype, W).

Voucher: Colombia, Schlim 1031 (K).

Note: The specimen at P was selected as lectotype by Luer (2023), but this is the holotype.

Stelis pulchra Luer & R.Escobar, Harvard Pap. Bot. 21(1): 84. 2016.

Illustration: Luer 2016a, f. 41.

Distribution: *COL.

Type: Colombia, Luer et al. 7505 (holotype, SEL).

Stelis pugiunculi Lindl., Fol. Orchid. ~Stelis~ 17, no. 125. 1852-1855 [1859].

Illustration: Luer 2009a, f. 334; Romero-González & Carnevalli 2000, p. 1042.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Purdie s.n. (K, GH-00104938).

Stelis punicea Luer & R.Escobar, Harvard Pap. Bot. 21(1): 84. 2016.

Illustration: Luer 2016a, f. 42.

Distribution: *COL.

Type: Colombia, Luer et al. 7500 (holotype, SEL).

Stelis purpurea (Ruiz & Pav.) Willd., Sp. Pl. 4: 140. 1805. Humboldtia purpurea Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 235. 1798.

Illustration: Luer 2002a, f. 29; Luer 2004, f. 105 (p. 134); Luer 2007, f. 217; Luer 2009a, f. 105a; Luer 2018b, f. 99, f. 100; Luer 2018c, f. 40, f. 41; Pupulin 2013, f. 50.

Distribution: BOL COL ECU PER.

Type: Peru, Ruiz & Pavón s.n. (holotype, MA).

Voucher: Colombia, Humboldt & Bonpland s.n. (P).

Stelis pusilla Kunth, Nov. Gen. Sp. (quarto ed.) 1: 361. 1816.

Illustration: Luer 2009a, f. 405-405c; Luer 2018b, f. 102-104; Luer 2018c, f. 42-46; RomeroGonzález & Carnevalli 2000, p. 1043.

Distribution: BOL COL DOM ECU HAI PER VEN.

Type: Ecuador, Humboldt & Bonpland s.n. (holotype, P).

Voucher: Colombia, Holton s.n. (K).

Stelis putumayoensis Luer & R.Escobar, Harvard Pap. Bot. 22(1): 56. 2017.

Illustration: Luer 2017a, f. 50.

Distribution: *COL.

Type: Colombia, Luer et al. 3081 (holotype, SEL).

Stelis pyramidalis O.Duque, Orquideología 20(3): 364, 365. 1997.

Illustration: Duque 1997, p. 367, ph. 7-9 (pl. V).

Distribution: *COL.

Type: Colombia, Duque 666 (holotype, JAUM).

Stelis quintella Luer & Hirtz, Harvard Pap. Bot. 22(2): 101. 2017.

Illustration: Luer 2017b, f. 34.

Distribution: *COL.

Type: Colombia, Luer et al. 4589 (holotype, SEL).

Stelis radicans Luer & R.Escobar, Harvard Pap. Bot. 21(2): 216. 2016.

Illustration: Luer 2016b, f. 40.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8469 (holotype, SEL).

Stelis rabeiFoldats, Acta Bot. Venez. 3: 411. 1968.

Illustration: Foldats 1968, f. 35.

Distribution: COL VEN.

Type: Venezuela, Steyermark & Rabe 96760 (holotype, VEN).

Voucher: Colombia, Figueroa-Cardozo 16 (COL).

Stelis ramiromedinae (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 141. 2021. Crocodeilanthe ramiromedinae Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 130-131. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 30.

Distribution: *COL.

Type: Colombia, Schultes & Villarreal 7805 (holotype, AMES; drawing, UGDA).

Stelis rectangularis Luer & R.Escobar, Harvard Pap. Bot. 21(2): 216. 2016.

Illustration: Luer 2016b, f. 41.

Distribution: *COL.

Type: Colombia, Luer et al. 9052 (holotype, SEL).

Stelis recurvilabia (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 141. 2021. Crocodeilanthe recurvilabia Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 132-133. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 32.

Distribution: *COL.

Type: Colombia, Schultes & Cabrera 20106 (holotype, AMES-111705; drawing, UGDA).

Stelis regalis (Luer) Karremans, Lankesteriana 13(3): 329. 2014. Anathallis regalis (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Pleurothallis regalis Luer, Selbyana 5(2): 178. 1979. Specklinia regalis (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer 1999a, f. 22 (p. 131).

Distribution: COL ECU.

Type: Ecuador, Luer et al. 3998 (holotype, SEL).

Voucher: Colombia, Ordóñez 389 (JBB).

Stelis remulifera Luer & R.Escobar, Harvard Pap. Bot. 22(2): 101. 2017.

Illustration: Luer 2017b, f. 35. Fig. 15I.

Distribution: *COL.

Type: Colombia, Luer et al. 7481 (holotype, SEL).

Stelis renilabia (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 141. 2021. Crocodeilanthe renilabia Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 133-134. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 33.

Distribution: *COL.

Type: Colombia, Garay et al. 418 (holotype, AMES; drawing, UGDA).

Stelis repens Cogn., Symb. Antill. 6: 692. 1910.

Illustration: Luer 2009a, f. 407; Luer 2018b, f. 107; Luer & Toscano de Brito 2019, f. 21.

Distribution: BOL COL DOM ECU HAI.

Type: Dominican Republic, Tuerckheim 3329 (holotype, BR; isotypes, AMES, K, US).

Voucher: Colombia, Dueñas 2927 (JBB).

Stelis reptans Pridgeon & M.W.Chase, Lindleyana 17(2): 100. 2002. Pleurothallis scansor Luer, Phytologia 49(3): 216. 1981. Crocodeilanthe scansor (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Stelis scansor (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001, nom. illeg. Non Stelis scansor Rchb.f. [1855].

Illustration: Luer 1998a, f. 57.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 6259 (holotype, SEL).

Voucher: Colombia, Cuatrecasas 19264 (AMES).

Stelis reptata Luer & R.Escobar, Harvard Pap. Bot. 21(1): 85. 2016.

Illustration: Luer 2016a, f. 43.

Distribution: *COL.

Type: Colombia, Luer et al. 3706 (holotype, SEL).

Stelis retroversa O.Duque, Orquideología 20(3): 365, 366. 1997.

Illustration: Duque 1997, p. 368; ph. 1-3 (pl. VII).

Distribution: *COL.

Type: Colombia, Duque 1308 (holotype, JAUM).

Stelis rhamphosa O.Duque, Orquideología 27(2): 149-150. 2010.

Illustration: Duque 2010b, f. 23, ph. 63-64.

Distribution: *COL.

Type: Colombia, Duque 2046 (holotype, JAUM).

Stelis rhodotantha (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Crocodeilanthe rhodotantha (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Humboldtia rhodotantha (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis rhodotantha Rchb.f., Linnaea 22: 825. 1849.

Illustration: Luer 1998a, f. 54.

Distribution: BOL COL ECU.

Type: Colombia, Schlim 1442 (holotype, W; isotype, G).

Stelis rhomboidea Garay, Canad. J. Bot. 34: 353. 1956.

Illustration: Luer 2018a, f. 38. Fig. 15J.

Distribution: *COL.

Type: Colombia, Cuatrecasas 9465 (holotype, US1850742; isotype, AMES).

Stelis rhynchanthera F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 447. 1898.

Illustration: Luer 2018a, f. 39.

Distribution: *COL.

Type: Colombia, Lehmann 6817 (holotype, K).

Stelis ricaurtensis Luer & Hirtz, Harvard Pap. Bot. 22(2): 103. 2017.

Illustration: Luer 2017b, f. 36.

Distribution: *COL.

Type: Colombia, Luer et al. 4595 (holotype, SEL).

Stelis richardiiKarremans, Phytotaxa 496(2): 141. 2021. Crocodeilanthe putumayoensis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 129-130. 2020. Non Stelis putumayoensis Luer & R.Escobar [2017].

Illustration: Kolanowska & Szlachetko 2020b, f. 29.

Distribution: *COL.

Type: Colombia, Schultes & Villarreal 7591 (holotype, AMES; drawing, UGDA).

Stelis rictoria (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Crocodeilanthe rictoria (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Pleurothallis rictoria Rchb.f., Linnaea 41: 14. 1877.

Illustration: Luer 1998a, f. 55.

Distribution: *COL.

Type: Colombia, Warscewicz s.n. (holotype, W).

Stelis rictus O.Duque, Orquideología 27(2): 150, 153. 2010.

Illustration: Duque 2010b, f. 24, ph. 65-66.

Distribution: *COL.

Type: Colombia, Duque 2468 (holotype, JAUM).

Stelis risaraldae Luer & R.Escobar, Harvard Pap. Bot. 22(2): 103. 2017.

Illustration: Luer 2017b, f. 37.

Distribution: *COL.

Type: Colombia, Luer et al. 16775 (holotype, SEL).

Stelis rodrigoi (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Condylago rodrigoiLuer, Orquideología 15(2-3): 118-122. 1982.

Illustration: Luer 1982, p. 119, p. 120; Luer 1987, pl. 1 (p. 23).

Distribution: *COL.

Type: Colombia, Luer 1952 (holotype, SEL; isotype, JAUM).

Stelis romerocastanedoi (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe romerocastanedoi Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 134. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 34.

Distribution: *COL.

Type: Colombia, Catrecasas & Romero Castañeda 25343 (holotype, AMES; drawing, UGDA).

Stelis roseopunctata (Lindl.) R. Bernal, Phytoneuron 22: 5. 2015. Pleurothallis roseopunctata Lindl., Orchid. Linden. 2. 1846. Humboldtia roseopunctata (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891.

Illustration: Luer 1998a, f. 18.

Distribution: COL VEN.

Type: Venezuela, Linden 629 (holotype, K-000674230; isotypes, AMES-00074702, BR0000006584979, G-00168955, P-00485832, P-00485831, W-R-0053234).

Voucher: Colombia, Humboldt & Bonpland s.n. (P, W).

Stelis rubens Schltr., Repert. Spec. Nov. Regni Veg. 8(191-195): 564. 1910. Apatostelis rubens (Schltr.) Garay, Bot. Mus. Leafl. 27(7-9): 190. 1979.

Distribution: BEL COL GUA HON MEX NIC.

Type: Guatemala, von Türckheim II 1061 (holotype, B, destroyed). Guatemala, von Türckheim II 1061 [neotype, AMES-00104966 (24933), illustration of type (Luer 2023)].

Voucher: Colombia, Duque 1842 (JAUM).

Stelis ruris O.Duque, Orquideología 27(2): 153-154. 2010.

Illustration: Duque 2010b, f. 25, ph. 67-68.

Distribution: *COL.

Type: Colombia, Duque 2455 (holotype, JAUM).

Stelis sagittata Zambrano & Solano, Phytotaxa 376(4): 183. 2018. Crocodeilanthe jamiesonii (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004. Humboldtia jamiesonii (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis jamiesonii Lindl., Edwards’s Bot. Reg. 21: sub t. 1797. 1835. Stelis jamiesonii (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 264. 2001, nom. illeg. Non Stelis jamesonii Lindl. [1834].

Illustration: Luer 1998a, f. 28.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K; isotype, AMES).

Voucher: Colombia, Cuatrecasas 18794 (AMES, NY, US).

Stelis salomonica O.Duque, Orquideología 27(2): 15. 2010.

Illustration: Duque 2010b, f. 27, ph. 71-73.

Distribution: *COL.

Type: Colombia, Murillo s.n. (holotype, JAUM).

Stelis salpingantha (Luer & Hirtz) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Crocodeilanthe salpingantha (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Pleurothallis salpingantha Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 65: 50-51, f. 56. 1998.

Illustration: Luer 1998a, f. 56.

Distribution: COL ECU PER VEN.

Type: Ecuador, Luer et al. 10865 (holotype, MO).

Voucher: Colombia, Luer et al. 10335 (MO).

Stelis samson Luer & R.Escobar, Harvard Pap. Bot. 21(1): 85. 2016.

Illustration: Luer 2016a, f. 45.

Distribution: *COL.

Type: Colombia, Luer et al. 10098 (holotype, MO).

Stelis sanguinea Garay, Canad. J. Bot. 34(3): 354. 1956.

Illustration: Garay 1956b, f. 2A.

Distribution: *COL.

Type: Colombia, John 20878 (holotype, AMES00271912; isotype, US-00093590).

Stelis santiagoensis Mansf., Notizbl. Bot. Gart. Berlin-Dahlem 10: 239. 1928.

Illustration: Romero-González & Carnevalli 2000, p. 1046.

Distribution: BOL COL FRE GUY PER VEN.

Type: Peru, Tessmann 4363 (holotype, B, destroyed). Peru, Tessmann 4068 [lectotype (selected here), AMES-00271935; isolectotype, S-R-5896; Illustration, AMES-00104974].

Voucher: Colombia, Duque 1148 (JAUM).

Stelis saurota Luer & R.Escobar, Harvard Pap. Bot. 22(2): 103. 2017.

Illustration: Luer 2017b, f. 38.

Distribution: *COL.

Type: Colombia, Luer et al. 9006 (holotype, SEL).

Stelis scalaris Luer & R.Escobar, Harvard Pap. Bot. 22(2): 104. 2017.

Illustration: Luer 2017b, f. 39, 40.

Distribution: *COL.

Type: Colombia, Luer & Escobar 6576 (holotype, SEL).

Stelis scansor Rchb.f., Bonplandia (Hannover) 3: 241. 1855.

Illustration: Luer 2009a, f. 410.

Distribution: COL ECU.

Type: Colombia, Jameson 448 (holotype, W; isotypes; AMES, BM, BR, G, K, LE).

Stelis scaphoides O.Duque, Orquideologia 20: 369. 1997.

Illustration: Duque 1997, p. 367, ph. 4-6 (pl. VII).

Distribution: *COL.

Type: Colombia, Duque 1814 (holotype, JAUM).

Stelis sceptrumrubrum Luer & R.Escobar, Harvard Pap. Bot. 22(1): 56. 2017.

Illustration: Luer 2017a, f. 51.

Distribution: *COL.

Type: Colombia, Luer et al. 7670 (holotype, SEL).

Stelis schlechterianaGaray, Bot. Mus. Leafl. 27(7-9): 184. 1979. Stelis mirabilis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 32. 1924, nom. illeg. Non Stelis mirabilis Schltr. [1923].

Distribution: *COL.

Type: Colombia, Hopp 140 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis schlimii (Luer) Karremans, Lankesteriana 13(3): 330. 2014. Anathallis schlimii (Luer) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Pleurothallis schlimii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 76: 120, f. 25. 1999. Specklinia schlimii (Luer) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer 1999a, f. 25 (p. 132).

Distribution: *COL.

Type: Colombia, Schlim 1441 (holotype, W; isotype, K-L).

Stelis schmidtchenii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 157. 1924.

Illustration: Luer 2018a, f. 40-42; RomeroGonzález & Carnevalli 2000, p. 1047.

Distribution: COL VEN.

Type: Colombia, Schmidtchen s.n. (holotype, B, lost). Colombia, Luer 17593 [neotype, SEL (Luer 2018a)].

Stelis schneideri (Kolan. & Szlach.) Karremans, comb. nov. Bas.: Crocodeilanthe schneideri Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 136. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 36.

Distribution: *COL.

Type: Colombia, Schneider 276 (holotype, AMES116223; drawing, UGDA).

Stelis schnitteri Schltr., Notizbl. Bot. Gart. BerlinDahlem 7: 528. 1921.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Stelis schomburgkii Fawc. & Rendle, J. Bot. 48: 108. 1910.

Illustration: Luer 2007, f. 242; Luer 2009a, f. 242a, f. 242b; Luer 2018c, f. 47; Romero-González & Carnevalli 2000, p. 1048.

Distribution: COL ECU GUY VEN.

Type: Venezuela, Schomburgk 1025 (holotype, K).

Voucher: Colombia, Viveros 147 (HUQ).

Stelis schultesii (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe schultesii Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 136-137. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 37.

Distribution: *COL.

Type: Colombia, Schultes s.n. (holotype, AMES; drawing, UGDA).

Stelis sclerophylla (Lindl.) Karremans, Lankesteriana 13(3): 330. 2014. Anathallis sclerophylla (Lindl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 250. 2001. Humboldtia sclerophylla (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis sclerophylla Lindl., Edwards’s Bot. Reg. 21, sub. t. 1797 no. 31. 1835. Specklinia sclerophylla (Lindl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 263. 2004.

Illustration: Luer 1999a, f. 26 (p. 132); Santos et al. 2019, f. 1p, f. 11; Luer 1975f, f. 77.

Distribution: BOL BRA COL ECU GUY PER VEN.

Type: Brazil, Martius s.n. (holotype, M; isotype, W).

Voucher: Colombia, Lehmann 6065 (K, AMES, HB, NY, US, W).

Stelis scutella O.Duque, Orquideología 20(3): 370-372. 1997.

Illustration: Duque 1997, p. 367; ph. 7-9 (pl. VII).

Distribution: *COL.

Type: Colombia, Duque 1631 (holotype, JAUM).

Stelis scutellifera Luer & R.Escobar, Harvard Pap. Bot. 22(1): 56. 2017.

Illustration: Luer 2017a, f. 52.

Distribution: *COL.

Type: Colombia, Luer et al. 7516 (holotype, SEL).

Stelis secundosa Luer, Harvard Pap. Bot. 22(2): 104. 2017.

Illustration: Luer 2017b, f. 41.

Distribution: *COL.

Type: Colombia, Escobar 6543 (holotype, SEL).

Stelis sellaformis O.Duque ex Karremans, Phytotaxa 203(3): 293. 2015. Stelis sellaformis O.Duque, Orquideología 27(2): 161-162. 2010, nom. inval.

Illustration: Duque 2010b, f. 28, ph. 74-77.

Distribution: *COL.

Type: Colombia, Duque & Gonzaga 2463 (JAUM).

Stelis semperflorens Luer, Phytologia 49(3): 238-239. 1981.

Distribution: COL PAN.

Type: Panama, Luer et al. 966 (holotype, SEL).

Voucher: Colombia, Gentry 30460 (JAUM, MO).

Stelis septicola Luer & Endara, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 48-49. 2007.

Illustration: Luer 2007, f. 287; Luer 2009a, f. 408.

Distribution: COL ECU.

Type: Ecuador, Lojtnant & Molau 15605 (holotype, AAU).

Voucher: Colombia, Duque 2135 (JAUM).

Stelis sesquipedalis Lindl., Orchid. Linden. 3. 1859.

Illustration: Luer 2017a, f. 21, 22.

Distribution: COL VEN.

Type: Venezuela, Linden 632 (holotype, K).

Voucher: Colombia, Luer et al. 10323 (MO).

Stelis sibundoyensis (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe sibundoyensis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 137-138. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 38.

Distribution: *COL.

Type: Colombia, Medina et al. S18a/07 (holotype, JAUM).

Stelis silverstonei Luer, Orquideología 22(1): 62-64. 2001.

Illustration: Luer 2001b, p. 63; Luer 2016a, f. 5.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 4147 (holotype, CUVC).

Stelis silvestris Luer & R.Escobar, Harvard Pap. Bot. 21(2): 218. 2016.

Illustration: Luer 2016b, f. 42.

Distribution: *COL.

Type: Colombia, Luer et al. 3706 (holotype, SEL).

Stelis simula Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 93. 1920.

Distribution: *COL.

Type: Colombia, Lehmann 6813 (holotype, B, destroyed; lectotype (selected here), AMES00104986; isolectotypes, K-000573875, LE00001406, NY-00009413).

Note: An obscure taxon.

Stelis sobrina Luer & R.Escobar, Harvard Pap. Bot. 22(2): 107. 2017.

Illustration: Luer 2017b, f. 42, 43.

Distribution: *COL.

Type: Colombia, Luer et al. 3016 (holotype, SEL).

Stelis spathilabia (Schltr.) Karremans, Lankesteriana 13(3): 330. 2014. Anathallis spathilabia (Schltr.) Pridgeon & M.W.Chase, Lindleyana 16(4): 251. 2001. Pleurothallis spathilabia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 56. 1924. Specklinia spathilabia (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Luer 1999a, f. 28 (p. 132).

Distribution: COL ECU.

Type: Colombia, Hopp 117 (holotype, B, destroyed).

Stelis spathosa (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 266. 2001. Crocodeilanthe spathosa (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Pleurothallis spathosa Luer & R.Escobar, Orquideología 20: 86. 1996.

Illustration: Luer & Escobar 1996, p. 85; Luer 1998a, f. 60.

Distribution: *COL.

Type: Colombia, Luer et al. 10175 (holotype, MO).

Stelis spathulata Poepp. & Endl., Nov. Gen. Sp. Pl. 1: 46-47, t. 80. 1836.

Illustration: Luer 2009a, f. 412, f. 412a; Luer 2018c, f. 48; Romero-González & Carnevalli 2000, p. 1049.

Distribution: COL COS ECU GUA PAN PER VEN.

Type: Peru, Poeppig s.n. [lectotype, W (Kolanowska 2014)].

Voucher: Colombia, Viveros 785 (HUQ).

Stelis splendens Luer, Harvard Pap. Bot. 22(2): 107. 2017.

Illustration: Luer 2017b, f. 44.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 3784 (holotype, MO; isotype, CUVC).

Stelis stapedia O.Duque, Orquideología 20(3): 372, 373. 1997.

Illustration: Duque 1997, p. 368; ph. 1-3 (pl. VIII).

Distribution: *COL.

Type: Colombia, Duque 1376 (holotype, JAUM).

Stelis stenophylla Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Illustration: Luer 2018a, f. 43; Romero-González & Carnevalli 2000, p. 1050.

Distribution: COL VEN.

Type: Colombia, Wagener s.n. (holotype, W).

Stelis stergiosii (Carnevali & I.Ramírez) Karremans, Phytotaxa 203(3): 293. 2015. Crocodeilanthe stergiosii (Carnevali & I.Ramírez) Carnevali & I.Ramírez, Nuevo Cat. Fl. Vasc. Venezuela 578. 2008. Pleurothallis stergiosii Carnevali & I.Ramírez, Harvard Pap. Bot. 3: 247. 1998.

Illustration: Kolanowska & Szlachetko 2020b, f. 2.

Distribution: COL VEN.

Type: Venezuela, Stergios et al. 2246 (holotype, PORT; isotypes, MO, VEN).

Voucher: Colombia, Foster 1768 (AMES-114016).

Stelis stigmatosa Luer & R.Escobar, Harvard Pap. Bot. 22(2): 107. 2017.

Illustration: Luer 2017b, f. 45.

Distribution: *COL.

Type: Colombia, Luer 6745 (holotype, SEL).

Stelis stipitata Luer & R.Escobar, Harvard Pap. Bot. 21(2): 218. 2016.

Illustration: Luer 2016b, f. 43.

Distribution: *COL.

Type: Colombia, Luer et al. 8856 (holotype, SEL).

Stelis stolonifera Luer & Hirtz, Harvard Pap. Bot. 22(2): 108. 2017.

Illustration: Luer 2017b, f. 46.

Distribution: *COL.

Type: Colombia, Luer et al. 4642 (holotype, SEL).

Stelis straminea Luer & R.Escobar, Harvard Pap. Bot. 21(2): 218. 2016.

Illustration: Luer 2016b, f. 44.

Distribution: *COL.

Type: Colombia, Luer & Escobar 8477 (holotype, SEL).

Stelis striolata Lindl., Fol. Orchid. ~Stelis~ 4, no. 26. 1852-1855 [1859].

Illustration: Luer 2009a, f. 413; Romero-González & Carnevalli 2000, p. 1051.

Distribution: COL ECU PER VEN.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Rivas 26 (HUQ).

Stelis subanaKarremans, Phytotaxa 496(2): 140. 2021. Crocodeilanthe bogotensis Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 111-112. 2020. Non Stelis bogotensis Schltr. [1924].

Illustration: Kolanowska & Szlachetko 2020b, f. 7.

Distribution: *COL.

Type: Colombia, Schiefer 702 (holotype, AMES113802; drawing, UGDA).

Stelis sublesta Luer & R.Escobar, Harvard Pap. Bot. 21(2): 220. 2016.

Illustration: Luer 2016b, f. 45.

Distribution: *COL.

Type: Colombia, Luer et al. 3085 (holotype, SEL).

Stelis suborbiculata (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe suborbiculata Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 138-139. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 39.

Distribution: *COL.

Type: Colombia, Schneider 29 (holotype, AMES115985; drawing, UGDA).

Stelis sulcata O.Duque, Orquideología 27(2): 154, 157. 2010.

Illustration: Duque 2010b, f. 26, ph. 69-70.

Distribution: *COL.

Type: Colombia, Duque 2448 (holotype, JAUM).

Stelis superbiens Lindl., Fol. Orchid. ~Stelis~ (8): 8. 1852-1855 [1859].

Illustration: Luer 2009a, f. 395, f. 414; Luer 2018b, f. 131; Luer 2018c, f. 49-51; Misas Urreta 2005, p. 714; Ortiz & Uribe 2007 (CD Edition). Fig. 14H.

Distribution: BOL COL COS ECU GUA HON MEX PAN PER VEN.

Type: Colombia, Purdie s.n. (holotype, K; isotype, GH).

Stelis tarantula (Luer & Hirtz) Pridgeon & M.W.Chase, Lindleyana 16(4): 267. 2001. Effusiella tarantula (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007. Pleurothallis tarantula Luer & Hirtz, Lindleyana 11(3): 186-187. 1996. Specklinia tarantula (Luer & Hirtz) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Luer 2000a, f. 38 (p. 99); Luer 1996c, f. 57. Fig. 14D.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 13553 (holotype, MO; isotypes, K, QCNE).

Voucher: Colombia, Luer 14212 (MO).

Stelis taurina O.Duque, Orquideología 20(3): 374, 376. 1997.

Illustration: Duque 1997, p. 368; ph. 7-9 (pl. VIII).

Distribution: *COL.

Type: Colombia, Duque 161 (holotype, JAUM).

Stelis tauroculus Luer & R.Escobar, Harvard Pap. Bot. 21(2): 220. 2016.

Illustration: Luer 2016b, f. 46.

Distribution: *COL.

Type: Colombia, Luer et al. 12541 (holotype, MO).

Stelis tenuilabris Lindl., Fol. Orchid. ~Stelis~ 4, no. 20. 1852-1855 [1859].

Illustration: Luer 2007, f. 222; Luer 2018c, f. 52; Romero-González & Carnevalli 2000, p. 1003.

Distribution: COL ECU VEN.

Type: Venezuela, Fendler 1471 (holotype, K; isotypes, AMES, BR, G, GOET).

Voucher: Colombia, Lozano 4141 (COL).

Stelis tenuipetala Garay, Canad. J. Bot. 34(3): 354. 1956.

Illustration: Garay 1956b, f. 2C.

Distribution: *COL.

Type: Colombia, von Sneidern 1097 (holotype, S).

Stelis titanica Luer & R.Escobar, Harvard Pap. Bot. 21(2): 220. 2016.

Illustration: Luer 2016b, f. 47.

Distribution: *COL.

Type: Colombia, Luer et al. 14172 (holotype, MO).

Stelis tolimensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 157. 1924.

Distribution: COL VEN.

Type: Colombia, Schmidtchen s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis tonduziana Schltr., Beih. Bot. Centralbl., Abt. 2 36(2): 393. 1918. Apatostelis tonduziana (Schltr.) Garay, Bot. Mus. Leafl. 27(7-9): 191. 1979.

Distribution: COL COS.

Type: Costa Rica, Jiménez 618 [holotype, B, destroyed; lectotype, AMES-24938, photo of type (Pupulin et al. 2022)].

Voucher: Colombia, Duque 1664 (JAUM).

Stelis tortilis (Luer & R.Escobar) Pridgeon & M.W.Chase, Lindleyana 16(4): 267. 2001. Effusiella tortilis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 107. 2007. Pleurothallis tortilis Luer & R.Escobar, Orquideología 14(2): 180. 1981. Specklinia tortilis (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Illustration: Luer & Escobar 1981a, p. 181; Luer 2000a, f. 40 (p. 99).

Distribution: COL ECU.

Type: Colombia, Luer et al. 3021 (holotype, SEL; isotype JAUM).

Stelis torulosa Luer & R.Escobar, Harvard Pap. Bot. 21(2): 221. 2016.

Illustration: Luer 2016b, f. 48.

Distribution: *COL.

Type: Colombia, Luer et al. 11390 (holotype, MO).

Stelis triangulisepala C.Schweinf., Bot. Mus. Leafl. 15: 22. 1951.

Distribution: COL PER.

Type: Peru, Stork & Horton 10425 (holotype, F-1051159).

Voucher: Colombia, Ortiz 403 (HPUJ).

Stelis trichorrhachis Rchb.f., Bonplandia (Hannover) 3: 71. 1855.

Illustration: Luer 2009a, f. 416.

Distribution: COL ECU VEN.

Type: Colombia, Schlim 168 (holotype, W; isotypes, BR, G, K).

Stelis tridactylon Luer, Selbyana 5(2): 193. 1979.

Illustration: Luer 2009a, f. 417.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 622 (holotype, SEL; isotypes: AMES, K).

Voucher: Colombia, Duque 1143 (JAUM).

Stelis tridentata Lindl., Fol. Orchid. ~Stelis~ (8): 6. 1852-1855 [1859].

Illustration: Barringer 1986, f. 17; Luer 2002a, f. 56; Luer 2009a, f. 56a, f. 56b; Romero-González & Carnevalli 2000, p. 1052.

Distribution: COL COS ECU PER VEN.

Type: Ecuador, Jameson s.n. (holotype, W).

Voucher: Colombia, Duque 1489 (JAUM).

Stelis trifoliacea Luer & R.Escobar, Harvard Pap. Bot. 21(2): 221. 2016.

Illustration: Luer 2016b, f. 49.

Distribution: *COL.

Type: Colombia, de Wilde 3877 (holotype, MO).

Stelis trilobata O.Duque, Orquideología 27(2): 165-166. 2010.

Illustration: Duque 2010b, f. 29, ph. 78-79.

Distribution: *COL.

Type: Colombia, Duque 2452 (holotype, JAUM).

Stelis trimera Luer, Selbyana 30(1): 24. 2009.

Illustration: Luer 2009b, f. 46.

Distribution: *COL.

Type: Colombia, García-Barriga & Jaramillo s.n. (holotype, US; isotype, COL).

Stelis triplaris Luer & R.Escobar, Harvard Pap. Bot. 22(1): 58. 2017.

Illustration: Luer 2017a, f. 53.

Distribution: *COL.

Type: Colombia, Luer et al. 7665 (holotype, SEL).

Stelis triplicata Lindl., Fol. Orchid. ~Stelis~ 2, no. 5. 1852-1855 [1859].

Illustration: Luer 2009a, f. 418, f. 418a; Luer 2018c, f. 53. Fig. 15H.

Distribution: COL ECU.

Type: Ecuador, Jameson 1357 (holotype, K).

Voucher: Colombia, Luer 2079 (SEL).

Stelis triseta Lindl., Fol. Orchid. ~Stelis~ 17, no. 126. 1852-1855 [1859].

Illustration: Luer 2009a, f. 312; Luer 2018b, f. 139, f. 140; Romero-González & Carnevalli 2000, p. 1053; Ortiz & Uribe 2007 (CD Edition).

Distribution: BOL COL ECU PER VEN.

Type: Bolivia, Bridges s.n. (holotype, K).

Voucher: Colombia, Ortiz 540 (HPUJ).

Stelis tritriangulata Luer & R.Escobar, Harvard Pap. Bot. 22(1): 58. 2017.

Illustration: Luer 2017a, f. 54.

Distribution: *COL.

Type: Colombia, Pennell 7001 (holotype, AMES).

Stelis trochophora Luer & R.Escobar, Harvard Pap. Bot. 21(2): 223. 2016.

Illustration: Luer 2016b, f. 50.

Distribution: *COL.

Type: Colombia, Luer et al. 7583 (holotype, SEL).

Stelis trullilabia Luer & R.Escobar, Harvard Pap. Bot. 21(1): 88. 2016.

Illustration: Luer 2016a, f. 46.

Distribution: *COL.

Type: Colombia, Luer et al. 3032 (holotype, SEL).

Stelis truncata Lindl., Companion Bot. Mag. 2: 353. 1836.

Illustration: Luer 2009a, f. 339, f. 339a; Luer 2018b, f. 142, f. 143; Luer 2018c, f. 54-56.

Distribution: BOL COL ECU PER.

Type: Peru, Mathews s.n. (holotype, K).

Voucher: Colombia, Lehmann 8214 (K, AMES, LE, NY).

Stelis tryssa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 88. 2016.

Illustration: Luer 2016a, f. 47.

Distribution: *COL.

Type: Colombia, Luer et al. 8947 (holotype, SEL).

Stelis tsubotae Luer & R.Escobar, Harvard Pap. Bot. 21(2): 223. 2016.

Illustration: Luer 2016b, f. 51.

Distribution: *COL.

Type: Colombia, Luer 16859 (holotype, MO).

Stelis tunguraguae (F.Lehm. & Kraenzl.) Pridgeon & M.W.Chase, Lindleyana 16(4): 267. 2001. Crocodeilanthe tunguraguae (F.Lehm. & Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Pleurothallis tunguraguae F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 439. 1899.

Illustration: Luer 1998a, f. 63; Luer 1975f, f. 84.

Distribution: BOL COL ECU PER.

Type: Ecuador, Lehmann 4504 [holotype, B, destroyed, lectotype, K; isolectotypes, NY, US (Luer 1998a)].

Voucher: Colombia, Escobar 8299 (MO).

Stelis uberis Luer & R.Escobar, Harvard Pap. Bot. 21(2): 223. 2016.

Illustration: Luer 2016b, f. 52.

Distribution: *COL.

Type: Colombia, Luer & Escobar 7582 (holotype, SEL).

Stelis umbelliformis Hespenh. & Dressler, Orquideología 6(1): 21-23. 1971. Apatostelis umbelliformis (Hespenh. & Dressler) Garay, Bot. Mus. Leafl. 27(7-9): 199. 1979.

Distribution: COL PAN.

Type: Panama, Hespenheide & Dressler 2206 (holotype, US; isotypes, K, MO, PH).

Voucher: Colombia, Benavides 4284 (PSO, MO).

Stelis umbriae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 40. 1924.

Distribution: *COL.

Type: Colombia, Hopp 110 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available. Explicitly excluded by Luer (2018a) from the synonymy of S. chamaestelis.

Stelis uncinula Luer & Hirtz, Harvard Pap. Bot. 22(2): 108. 2017.

Illustration: Luer 2017b, f. 47.

Distribution: *COL.

Type: Colombia, Luer et al. 2706 (holotype, SEL).

Stelis uribeorum Luer & R.Escobar, Harvard Pap. Bot. 22(1): 60. 2017.

Illustration: Luer 2017a, f. 55.

Distribution: *COL.

Type: Colombia, Luer 18423 (holotype, MO).

Stelis validipes Luer & R.Escobar, Harvard Pap. Bot. 22(1): 60. 2017.

Illustration: Luer 2017a, f. 56.

Distribution: *COL.

Type: Colombia, Luer et al. 7591 (holotype, SEL).

Stelis vallecaucana (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe vallecaucana Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 140. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 41.

Distribution: *COL.

Type: Colombia, Cuatrecasas 18180 (holotype, AMES-72611; drawing, UGDA).

Stelis valvulosa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 88. 2016.

Illustration: Luer 2016a, f. 48.

Distribution: *COL.

Type: Colombia, Luer et al. 16822 (holotype, MO).

Stelis vanescens Luer, Harvard Pap. Bot. 22(2): 108. 2017.

Illustration: Luer 2017b, f. 48.

Distribution: *COL.

Type: Colombia, Silverstone-Sopkin 4150 (holotype, MO; isotype, CUVC).

Stelis variola Luer & R.Escobar, Harvard Pap. Bot. 22(2): 110. 2017.

Illustration: Luer 2017b, f. 49.

Distribution: *COL.

Type: Colombia, Luer et al. 6825 (holotype, SEL).

Stelis velaticaulis (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 267. 2001. Crocodeilanthe velaticaulis (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004. Humboldtia velaticaulis (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis velaticaulis Rchb.f., Linnaea 22: 824. 1849.

Illustration: Luer 1998a, f. 66, f. 67. Fig. 14C.

Distribution: COL COS ECU PAN PER VEN.

Type: Venezuela, Otto s.n. (holotype, W).

Voucher: Colombia, Lozano 3495 (NY).

Stelis velutina Lindl., Fol. Orchid. ~Stelis~ 17, no. 130. 1852-1855 [1859].

Illustration: Luer 2009a, f. 341.

Distribution: COL ECU PER.

Type: Ecuador, Jameson s.n. (holotype, K).

Voucher: Colombia, Duque 1277 (JAUM).

Stelis venezuelensisFoldats, Acta Bot. Venez. 3: 421. 1968.

Illustration: Foldats 1968, f. 39; Romero-González & Carnevalli 2000, p. 1054; Ortiz & Uribe 2007 (CD Edition).

Distribution: COL VEN.

Type: Venezuela, Steyermark & Dunsterville 98497 (holotype, VEN).

Voucher: An illustration voucher of a Colombian specimen reproduced in Ortiz & Uribe 2007 (CD Edition).

Stelis verecunda Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 158. 1924.

Distribution: *COL.

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis verruculosa Luer & R.Escobar, Harvard Pap. Bot. 22(2): 110. 2017.

Illustration: Luer 2017b, f. 50.

Distribution: *COL.

Type: Colombia, Luer & Escobar 6602 (holotype, SEL).

Stelis vicaria Luer & R.Escobar, Harvard Pap. Bot. 23(1): 45. 2018.

Illustration: Luer 2018a, f. 44.

Distribution: *COL.

Type: Colombia, St. John 20782 (holotype, AMES; isotype, US).

Stelis vigax Luer & R.Escobar, Harvard Pap. Bot. 21(1): 89. 2016.

Illustration: Luer 2016a, f. 49.

Distribution: *COL.

Type: Colombia, Luer et al. 8884 (holotype, SEL).

Stelis vigoris Luer & R.Escobar, Harvard Pap. Bot. 22(2): 112. 2017.

Illustration: Luer 2017b, f. 51.

Distribution: *COL.

Type: Colombia, Luer & Escobar 6591 (holotype, SEL).

Stelis violacea Garay, Canad. J. Bot. 34(3): 355. 1956.

Illustration: Garay 1956b, f. 2E; Romero-González & Carnevalli 2000, p. 1055.

Distribution: COL VEN.

Type: Colombia, von Sneidern 1096 (holotype, S; isotype, AMES-00271910).

Stelis voluptuosa Luer & R.Escobar, Harvard Pap. Bot. 21(1): 89. 2016.

Illustration: Luer 2016a, f. 50.

Distribution: *COL.

Type: Colombia, Luer et al. 8769 (holotype, SEL).

Stelis vulcani Rchb.f., Otia Bot. Hamburg. 1: 19, no. 73. 1878.

Illustration: Luer 2009a, f. 342, f. 342a; Luer 2017a, f. 38; Romero-González & Carnevalli 2000, p. 1056.

Distribution: COL ECU PER VEN.

Type: Ecuador, Lehmann 84 (holotype: W-R0020260).

Voucher: Colombia, Luer et al. 7655 (SEL).

Stelis vulpecula Luer & R.Escobar, Harvard Pap. Bot. 21(1): 91. 2016.

Illustration: Luer 2016a, f. 51.

Distribution: *COL.

Type: Colombia, Luer et al. 17635 (holotype, MO).

Stelis walteri Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 41. 1924.

Distribution: *COL.

Type: Colombia, Hopp 170 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis warszewiczii (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021. Crocodeilanthe warszewiczii Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 141-142. 2020.

Illustration: Kolanowska & Szlachetko 2020b, f. 43.

Distribution: *COL.

Type: Colombia, Schneider 293 (holotype, AMES116221; drawing, UGDA).

Stelis weddelliana (Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 267. 2001. Crocodeilanthe weddelliana (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 103: 309. 2005. Crocodeilanthe weddelliana (Rchb.f.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 257. 2004, nom. inval. Humboldtia weddelliana (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis weddelliana Rchb.f., Xenia Orchid. 3: 24. 1878.

Illustration: Luer 1998a, f. 70.

Distribution: BOL COL.

Type: Bolivia, Mandon 1128 (holotype, W-R-0053642; isotypes, AMES-00074861, AMES-00243103, BM-000082450, BR-0000006571894, BR0000006571818, G-00168969, GH-00243102, GOET-008643, GOET-008655, K-000584261, MPU-018059, NY-00009251, NY00009250, P-00489157, P-00489156, P-00489154, P-00489158, P-00489155, S-R-10422).

Voucher: Colombia, Ewan 16515 (US).

Stelis wercklei Schltr., Repert. Spec. Nov. Regni Veg. Beih. 19: 20. 1923.

Distribution: COL COS NIC.

Type: Costa Rica, Wercklé 15 [holotype, B; destroyed; lectotype, AMES-30439, photo of type (Pupulin 2010)].

Voucher: Colombia, Duque 1791 (JAUM).

Stelis werneri Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 42. 1924.

Distribution: *COL.

Type: Colombia, Hopp 158 (holotype, B, destroyed).

Note: An obscure taxon for which no original material seems to be available.

Stelis xenica Luer & R.Escobar, Harvard Pap. Bot. 21(1): 91. 2016.

Illustration: Luer 2016a, f. 52.

Distribution: *COL.

Type: Colombia, Luer 12485 (holotype, MO).

Stelis zelleri Luer, Harvard Pap. Bot. 22(2): 112. 2017.

Illustration: Luer 2017b, f. 52.

Distribution: *COL.

Type: Colombia, Luer 6504 (holotype, SEL).

Stellamaris Mel.Fernández & Bogarín

Stellamaris pergrata (Ames) Mel.Fernández & Bogarín, Phytotaxa 340(2): 131. 2018. Pleurothallis pergrata Ames, Sched. Orch. 4: 24-25. 1923. Trichosalpinx pergrata (Ames) Luer, Phytologia 54(5): 396. 1983. Tubella pergrata (Ames) Archila, Revista Guatemal. 3(1): 62. 2000 [2009]. Tubella pergrata (Ames) Archila, Revista Guatemal. 17(1): 55. 2014 [2015], nom. superfl.

Illustration: Luer 1997, f. 69, f. 70; Luer 1975e, f. 92. Fig. 16G.

Distribution: COL COS PAN.

Type: Costa Rica, Lankester & Sancho 436 (holotype, AMES).

Voucher: Colombia, Lehmann 8245 (K).

Teagueia (Luer) Luer

Teagueia lehmannii Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 142. 1991.

Illustration: Luer 1991a, pl. 1 (p. 143).

Distribution: *COL.

Type: Colombia, Lehmann s.n. (holotype, K, H.K, 136).

Teagueia phasmida (Luer & R.Escobar) O. Gruss & M. Wolff, Orchid Atlas 417. 2007. Teagueia phasmida (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 121. 2007. nom. superfl. Platystele phasmida Luer & R.Escobar, Orquideología 16(3): 42-45. 1986. Teagueia phasmida Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 139. 1991. nom. inval.

Illustration: Luer & Escobar 1986, p. 43; Luer 1990, pl. 49; Luer 1991d, f. 484.

Distribution: *COL.

Type: Colombia, Escobar et al. 3404 (holotype, MO).

Note: According to the protologue the holotype is at MO, but Luer (1990) states that the holotype is at JAUM and the MO specimen is an isotype.

Teagueia rex (Luer & R.Escobar) O. Gruss & M. Wolff, Orchid Atlas 417. 2007. Platystele rex Luer & R.Escobar, Orquideología 15(2-3): 140. 1982. Teagueia rex (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 112: 121. 2007. nom. superfl. Teagueia rex Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 39: 139. 1991. nom. inval.

Illustration: Luer & Escobar 1982, p. 141; Luer 1990, pl. 59; Luer 1991d, f. 481; Karremans et al. 2016, f. 8f.

Distribution: *COL.

Type: Colombia, Luer et al. 3133 (holotype, SEL; isotype, JAUM).

Trichosalpinx Luer

Trichosalpinx acestrochila Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 88: 112, f. 27. 2002.

Illustration: Luer 2002a, f. 27 (p. 121).

Distribution: *COL.

Type: Colombia, Luer 16908 (holotype, MO).

Trichosalpinx blaisdellii (S. Watson) Luer, Phytologia 54(5): 394. 1983. Pleurothallis blaisdellii S. Watson, Proc. Amer. Acad. Arts 23(2): 284-285. 1888.

Illustration: Luer 1997, f. 2, f. 3, f.4.

Distribution: BEL COL COS ELS GUA HON MEX NIC PAN.

Type: Mexico, Liebmann 7307 (holotype, C).

Voucher: Colombia, Gentry 9335 (COL).

Trichosalpinx ciliaris (Lindl.) Luer, Phytologia 54(5): 395. 1983. Pleurothallis ciliaris (Lindl.) L.O.Williams, Caldasia 1(5): 14. 1942. Specklinia ciliaris Lindl., Edwards’s Bot. Reg. 24: Misc. 31. 1838.

Illustration: Luer 1997, f. 6.

Distribution: BEL COL COS GUA HON MEX NIC.

Type: Mexico, Loddiges s.n. (holotype, K).

Voucher: Colombia, Kolanowska 141 (UGDA).

Trichosalpinx caudata Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 64: 20, f. 5. 1997.

Illustration: Luer 1997, f. 5; Fernández et al. 2014, f. 13, f. 14l.

Distribution: COL COS PAN.

Type: Colombia, Luer 16907 (holotype, MO).

Trichosalpinx memor (Rchb.f.) Luer, Phytologia 54(5): 396. 1983. Humboldtia memor (Rchb.f.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis memor Rchb.f., Bonplandia (Hannover) 4: 330. 1856.

Illustration: Barringer 1986, f. 11; Luer 1997, f. 12, f. 13, f. 14; Stenzel 2007, f. 13; Fernández 2011, f. 5.

Distribution: BOL COL COS CUB ECU GUA MEX NIC PAN PER VEN.

Type: Colombia, Forkel s.n. (holotype, W-R0020793/W-0053202).

Trichosalpinx orbicularis (Lindl.) Luer, Phytologia 54(5): 396. 1983. Lepanthes orbiculata Lindl. ex Rchb.f., Xenia Orchid. 1: 152. 1856. Humboldtia orbicularis (Lindl.) Kuntze, Revis. Gen. Pl. 2: 668. 1891. Pleurothallis orbicularis (Lindl.) Lindl., Edwards’s Bot. Reg. 28: Misc. 79. 1842. Specklinia orbicularis Lindl., Edwards’s Bot. Reg. 24: Misc. 31. 1838.

Illustration: Luer 1997, f. 16; Luer 1975b, f. 33. Fig. 16H.

Distribution: BRA COL COS ECU FRE GUY L.A. PAN PER SUR VEN.

Type: Guyana, Loddiges s.n. (holotype, K).

Voucher: Colombia, Arévalo 341 (COL).

Trichosalpinx semilunata (Luer) Luer, Phytologia 54(5): 397. 1983. Pleurothallis semilunata Luer, Selbyana 3: 184. 1976. Tubella semilunata (Luer) Archila, Revista Guatemal. 17(1): 56. 2014 [2015], nom. superfl. Tubella semilunata (Luer) Archila, Revista Guatemal. 3(1): 64. 2000 [2009].

Illustration: Luer & Escobar 1984b, p. 181, p. 182; Luer 1997, f. 77.

Distribution: COL ECU.

Type: Ecuador, Luer et al. 523 (holotype, SEL).

Voucher: Colombia, Luer et al. 9044 (holotype, SEL).

Note: This taxon is clearly not related to other Trichosalpinx species in the strict sense, and is more likely a relative of Stellamaris. However, without DNA data to confirm its phylogenetic placement, a transfer to any other genus is premature.

Trisetella Luer

Trisetella abbreviata Luer, Phytologia 47: 82. 1980.

Illustration: Luer 1989, f. 1 (p. 77).

Distribution: COL ECU.

Type: Ecuador, Luer 2209 (holotype, SEL).

Voucher: Colombia, Arévalo 923 (COL).

Trisetella escobarii Luer, Lindleyana 1(3): 188. 1986.

Illustration: Luer 1989, f. 6 (p. 87).

Distribution: *COL.

Type: Colombia, Luer et al. 10324 (holotype, MO).

Trisetella gemmata (Rchb.f.) Luer, Phytologia 47(2): 57. 1980. Masdevallia gemmata Rchb.f., Gard. Chron. 19: 294. 1883. Triaristella gemmata (Rchb.f.) Luer, Selbyana 2(2-3): 205. 1978. nom. illeg. Triaristellina gemmata (Rchb.f.) Rauschert, Feddes Repert. 94(7-8): 469. 1983. nom. superfl.

Illustration: Luer 1989, f. 8 (p. 91).

Distribution: *COL.

Type: Colombia, O’Brien s.n. (holotype, W).

Trisetella tenuissima (C.Schweinf.) Luer, Phytologia 47(2): 58. 1980. Masdevallia tenuissima C.Schweinf., Bot. Mus. Leafl. 4(7): 111. 1937. Triaristella tenuissima (C.Schweinf.) Luer, Selbyana 2(2-3): 206. 1978. nom. illeg. Triaristellina tenuissima (C.Schweinf.) Rauschert, Feddes Repert. 94(7-8): 469. 1983. nom. superfl.

Illustration: Luer 1989, f. 17 (p. 109).

Distribution: COL PAN.

Type: Panama, Allen 587 (holotype, MO).

Voucher: Colombia, Luer 13087 (MO).

Trisetella triaristella (Rchb.f.) Luer, Phytologia 47(2): 58. 1980. Masdevallia triaristella Rchb.f., Gard. Chron., n.s. 6(138): 226. 1876. Triaristellina triaristella (Rchb.f.) Rauschert, Feddes Repert. 94(7- 8): 469. 1983. nom. superfl.

Illustration: Luer 1989, f. 18 (p. 111). f. 19 (p. 112); Williams & Williams 1894, p. 505.

Distribution: COL COS ECU PAN.

Type: Costa Rica, Endrés 566 (holotype, W).

Voucher: Colombia, Fernández-Alonso 12369 (COL).

Trisetella triglochin (Rchb.f.) Luer, Phytologia 47(2): 58. 1980. Masdevallia triglochin Rchb.f., Gard. Chron., n.s. 8(204): 648. 1877. Triaristella triglochin (Rchb.f.) Luer, Selbyana 2(2-3): 206. 1978. nom. illeg. Triaristellina triglochin (Rchb.f.) Rauschert, Feddes Repert. 94(7-8): 469. 1983. nom. superfl.

Illustration: Luer 1989, f. 20-24 (p. 115-119); Baquero & Mogrovejo 2021, f. 4d.

Distribution: BOL BRA COL COS ECU PAN PER VEN.

Type: Ecuador, Lehmann 68 (holotype, W).

Voucher: Colombia, Cogollo 3241 (FMB).

Trisetella vittata (Luer) Luer, Phytologia 47(2): 58. 1980. Masdevallia vittata Luer, Phytologia 39: 233-234. 1978. Triaristella vittata (Luer) Luer, Selbyana 2(2-3): 206. 1978. nom. illeg. Triaristellina vittata (Luer) Rauschert, Feddes Repert. 94(7-8): 469. 1983. nom. superfl.

Illustration: Luer 1989, f. 25 (p. 121); Baquero & Mogrovejo 2021, f. 4c.

Distribution: COL ECU.

Type: Ecuador, Luer 1756 (holotype, SEL).

Voucher: Colombia, Vieira 112 (JAUM).

Zootrophion Luer

Zootrophion aguirrei P.Ortiz, Orquideología 28(1): 11. 2011.

Illustration: Ortiz 2011, p. 8, p. 14.

Distribution: *COL.

Type: Colombia, Ortiz 1374 (holotype, HPUJ).

Note: Ortiz 1358 indicated as isotype, but is a paratype.

Zootrophion alvaroi (Garay) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 215. 2004. Cryptophoranthus alvaroi Garay, Orquideología 5: 15. 1970.

Illustration: Luer 2004, f. 1 (p. 228).

Distribution: COL ECU.

Type: Colombia, Escobar 372 (holotype, AMES).

Zootrophion antioquianum Uribe Vélez & Sauleda, New World Orchid. Nomencl. Notes 64: 1. 2020.

Illustration: Uribe-Velez & Sauleda 2020, p. 2, p. 3, p. 4(top).

Distribution: *COL.

Type: Colombia, Arrodeau s.n. (holotype, HPUJ).

Zootrophion argus (Kraenzl.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 215. 2004. Cryptophoranthus argus Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 437-438. 1921.

Illustration: Luer 2004, f. 2 (p. 228).

Distribution: *COL.

Type: Colombia, Schlim 762 [Lectotype, K; isotypes, W, NY (Luer 2004)].

Zootrophion atropurpureum (Lindl.) Luer, Selbyana 7(1): 80. 1982. Cryptophoranthus atropurpureus (Lindl.) Rolfe, Gard. Chron. 2: 698. 1887. Humboldtia atropurpurea (Lindl.) Kuntze, Revis. Gen. Pl. 2: 667. 1891. Pleurothallis atropurpurea (Lindl.) Lindl., Edwards’s Bot. Reg. 81. 1842. Specklinia atropurpurea Lindl., Edwards’s Bot. Reg. 21: 1797. 1836.

Illustration: Cogniaux 1896, f. 1 (p. 114); Luer 2004, f. 3 (p. 229); Rysy 2007, f. 1; Stenzel 2007, f. 16; Loefgren 1918, pl. XIXB; Misas Urreta 2005, p. 766, p. 767.

Distribution: BOL BRA COL CUB DOM ECU HAI JAM PER.

Type: Jamaica, unknown collector [lectotype (selected here), K-000463225].

Voucher: Colombia, Misas Urreta 67a (HPUJ).

Zootrophion dayanum (Rchb.f.) Luer, Selbyana 7(1): 82. 1982. Cryptophoranthus dayanus (Rchb.f.) Rolfe, Gard. Chron. 3(2): 693, f. 134. 1887. Masdevallia dayana Rchb.f., Gard. Chron. 14: 295. 1880.

Illustration: Luer 2004, f. 5, f. 6 (p. 229); Rysy 2007, f. 2; Williams & Williams 1894, p. 487. Fig. 16K.

Distribution: BOL COL ECU PER VEN.

Type: Colombia, Day s.n. (holotype, W; isotype, K).

Zootrophion dodsonii (Luer) Luer, Selbyana 7(1): 82. 1982. Cryptophoranthus dodsonii Luer, Selbyana 5(2): 145. 1979.

Illustration: Luer 2004, f. 7 (p. 230).

Distribution: COL ECU.

Type: Ecuador, Gentry & Schupp 9202 (holotype, SEL).

Voucher: Colombia, Luer 2965 (SEL).

Zootrophion endresianum (Kraenzl.) Luer, Selbyana 7(1): 82. 1982. Cryptophoranthus endresianus Kraenzl., Repert. Spec. Nov. Regni Veg. 17(492/503): 437. 1921.

Illustration: Luer 2004, f. 8 (p. 230); Rysy 2007, f. 3.

Distribution: COL COS ECU NIC.

Type: Costa Rica, Endrés s.n. (holotype, K).

Voucher: Colombia, Luer 2350 (SEL).

Zootrophion hypodiscus (Rchb.f.) Luer, Selbyana 7(1): 84. 1982. Cryptophoranthus hypodiscus (Rchb.f.) Rolfe, Gard. Chron. 2: 693. 1887. Masdevallia hypodiscus Rchb.f., Gard. Chron. 2: 234. 1878.

Illustration: Luer 2004, f. 11 (p. 231).

Distribution: COL ECU.

Type: Colombia, Lehmann s.n. (holotype, W).

Zootrophion ildephonsi P.Ortiz, Orquideología Supl. 1: 37. 2012.

Illustration: Ortiz 2012, f. 32-34.

Distribution: *COL.

Type: Colombia, Ortiz 1344 (holotype, HPUJ, lost).

Zootrophion lappaceum Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 223. 2004.

Illustration: Luer 2004, f. 12 (p. 231).

Distribution: *COL.

Type: Colombia, Luer 14289 (holotype, MO).

Zootrophion machaqway A.Doucette & J. Portilla, Phytotaxa 257(3): 244. 2016.

Illustration: Doucette et al. 2016b, f. 10, f. 11h; Uribe-Velez & Sauleda 2023, p. 2-6.

Distribution: COL ECU.

Type: Ecuador, Doucette 315 (holotype, WIS).

Voucher: Colombia, Aguirre s.n. (HPUJ).

Note: This species is unusual among Zootrophion species for its prolific habit. It was described from cultivation in Ecuador and the Colombian specimen was also prepared from cultivated material (under the name Z. griffin Luer in Uribe-Vélez & Sauleda 2023).

Zootrophion oblongifolium (Rolfe) Luer, Selbyana 7(1): 84. 1982. Cryptophoranthus oblongifolius Rolfe, Bull. Misc. Inform. Kew 1895: 5. 1895.

Illustration: Luer 2004, f. 14 (p. 231); Rysy 2007, f. 6. Fig. 16L.

Distribution: COL ECU PER.

Type: Colombia, Charlesworth s.n. (holotype, K; isotype, BR).

Zootrophion trivalve (Luer & R.Escobar) Luer, Selbyana 7: 84. 1982. Cryptophoranthus trivalvis Luer & R.Escobar, Orquideología 14: 118. 1981.

Illustration: Luer & Escobar 1981a, p. 119; Luer 2004, f. 15 (p. 232).

Distribution: *COL.

Type: Colombia, Escobar 1658 (holotype, JAUM; isotype, SEL).

Zootrophion ximenae (Luer & Hirtz) Pfahl, Internet Orchid Sp. Photo Encycl. Nomencl. Notes 3(2): 1. 2014. Epibator ximenae Luer & Hirtz, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 204. 2004.

Illustration: Luer 2004, f. 3 (p. 204); Sauleda et al. 2023, p. 4-7.

Distribution: COL? ECU.

Type: Ecuador, Hirtz & Hirtz 6294 (holotype, MO).

Voucher: Colombia, Mejia s.n. (HPUJ)

Note: The Colombian voucher for this taxon comes from cultivated material described under the name Epibator mejia Sauleda, Uribe Vélez & Szlach. of uncertain provenance (Sauleda et al. 2023). Additionally, no such specimen is physically present at HPUJ as indicated by the authors.

Additional taxa based on Colombian types

Acianthera adeodata P.Ortiz, O.Pérez & E.Parra, Orquideología 27(1): 56-59. 2010.

Accepted Name: Acianthera madisonii .

Type: Colombia, Pérez-Escobar et al. 512 (holotype, VALLE).

Note: The large leaf decurrent on the ramicaul, short raceme emerging above the base of the leaf, the gapping, fleshy flowers, with a yellow dorsal sepal, and purple lateral sepals are consistent with A. madisonii, a rare species described from Ecuador.

Acianthera caucanensis Uribe Vélez, Sauleda & Szlach., New World Orchid. Nomencl. Notes 111: 1. 2022.

Accepted Name: Pleurothallis macrophylla .

Type: Colombia, Aguirre s.n. (holotype, HPUJ).

Note: Inexplicably described in the distantly related genus Acianthera, this taxon is inseparable from P. macrophylla. The type specimen is said to be conserved at “HJPU”, which we assume refers to HPUJ. However, like other taxa described in this journal, the holotype does not seem to have been deposited.

Acianthera ortizii Uribe Vélez, Sauleda & Szlach., New World Orchid. Nomencl. Notes 110: 1-2. 2022.

Accepted Name: Pleurothallis macrophylla .

Type: Colombia, Aguirre s.n. (holotype, HPUJ).

Note: Inexplicably described in the distantly related genus Acianthera, this taxon is inseparable from P. macrophylla. The type specimen is said to be conserved at “HJPU”, which we assume refers to HPUJ. However, like other taxa described in this journal, the holotype does not seem to have been deposited.

Acostaea glandulata P.Ortiz, Orquideología 15(2,3): 166-168. 1982.

Accepted Name: Specklinia colombiana .

Type: Colombia, Ortiz 892 (holotype, COL).

Barbosella caespitifica (F.Lehm. & Kraenzl.) Garay, Orquideología 9(2): 114. 1974.

Accepted Name: Barbosella prorepens .

Type: Colombia, Lehmann 3305 (holotype, lost?).

Barbosella longipes Schltr., Repert. Spec. Nov. Regni Veg. 7: 118. 1920.

Accepted Name: Barbosella cucullata .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 41, No. 158). Colombia, Lehmann H.K.313 [neotype, K (Luer 2000a), here rejected and selected as epitype].

Brachionidium floribundum Garay, Canad. J. Bot. 34: 736. 1956.

Accepted Name: Brachionidium tuberculatum .

Type: Colombia, Renz 5253 (holotype, AMES; isotype, herb. Renz).

Crocodeilanthe asymmetrica Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 108-109. 2020.

Accepted Name: Pleurothallis xanthochlora .

Type: Colombia, Garay et al. 223 (holotype, AMES; drawing, UGDA).

Note: Described as a unique species of Crocodeilanthe, this taxon clearly belongs in Pleurothallis instead (Karremans et al. 2021) and, despite the claims of having asymmetric flower, is referable to P. xanthochlora.

Crocodeilanthe elegans (Kunth) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 256. 2004.

Accepted Name: Stelis roseopunctata .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; fragment, W).

Crocodeilanthe velaticauloides Kolan. & Szlach., Phyton. Annales Rei Botanicae 60: 140-141. 2020.

Accepted Name: Stelis pennelliana .

Type: Colombia, Pennell 2043 (holotype, AMES; drawing, UGDA).

Note: The holotype of this taxon is an isotype of S. pennelliana, a species that was inexplicably not even mentioned despite being conspecific.

Cryptophoranthus auriculatus Garay, Orquideología 5(1): 16. 1970.

Accepted Name: Ophidion pleurothallopsis .

Type: Colombia, Escobar 399 (holotype, AMES).

Cryptophoranthus lehmannii Rolfe, Bull. Misc. Inform. Kew 26. 1922, nom. superfl.

Accepted Name: Zootrophion dayanum .

Type: Colombia, Lehmann s.n. (holotype, K).

Note: This is a superfluous name because all requirements for valid publication were met before.

Cryptophoranthus lehmannii Rolfe, Orchid Rev. 11: 303. 1903.

Accepted Name: Zootrophion dayanum .

Type: Colombia, Lehmann s.n. [lectotype (selected here), K-000573922; isolectotypes, K-000573923, K-000573924. K-000573925, K-000573926, K-000573927].

Note: Even though Rolfe probably had no intent to formally publish this taxon here and tried to do so later, the fact is that all requirements for valid publication were met here. Several specimens were made overtime of one or several cultivated plants and the type is a mixed collection.

Cryptophoranthus moorei Rolfe, Bull. Misc. Inform. Kew 30. 1906, nom. superfl.

Accepted Name: Zootrophion hypodiscus .

Illustration: Luer 2004, f. 11 (p. 231).

Type: Colombia, unknown (holotype, K).

Note: This is a superfluous name because all requirements for valid publication were met before.

Cryptophoranthus moorei Rolfe, Orchid Rev. 11: 304. 1903.

Accepted Name: Zootrophion hypodiscus .

Type: Colombia, unknown [lectotype (selected here), K-000573984; isolectotypes, K-000573985, K-000573986].

Note: Even though Rolfe probably had no intent to formally publish this taxon here and attempt to do so later, the fact is that all requirements for valid publication were met here. Several specimens were made overtime of one or several cultivated plants and the type is a mixed collection.

Dendrobium elegans Kunth, Nov. Gen. Sp. 1: 358. 1816.

Accepted Name: Stelis roseopunctata .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; fragment, W).

Draconanthes trachysepala (Schltr.) Kolan., S.Nowak & Szlach., Turkish J. Bot. 44: 199. 2020.

Accepted Name: Draconanthes aberrans .

Type: Colombia, Lehmann 6910 [holotype, B, destroyed; lectotype, Schlechter’s floral analysis published by Mansfeld (1929: Pl. 34, No. 130)]. Colombia, Luer 12520 [epitype, MO (Luer 1996b)].

Note: In a rather unconvincing treatment of the D. aberrans complex (Kolanowska et al. 2020b), the authors separate several taxa on the basis of slight differences in illustrations and photographs by several different authors, some very precise, others quite rudimentary, with no regards to properly documenting extant natural populations and assessing variation. Some of the illustrations published by the authors are mislabeled, compounding the confusion.

Dracula callifera (Schltr.) Luer, Selbyana 2: 194. 1978.

Accepted Name: Dracula houtteana .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 25, No. 95)].

Dracula carderi (Rchb.f.) Luer, Selbyana 2: 194. 1978.

Accepted Name: Dracula inaequalis .

Type: Colombia, Carder s.n. (holotype, W).

Dracula carderiopsis (F.Lehm. & Kraenzl.) Luer, Selbyana 2: 194. 1978.

Accepted Name: Dracula houtteana .

Type: Colombia, Lehmann 2335 [lectotype, G-00168644; isolectotypes, BR-0000009973237, W].

Dracula lactea (Kraenzl.) Luer, Selbyana 2: 195. 1978.

Accepted Name: Dracula velutina .

Type: Colombia, Kalbreyer 1545 (holotype, B, destroyed).

Dracula lowii (Rolfe) Luer, Selbyana 2: 196. 1978.

Accepted Name: Dracula spectrum .

Type: Colombia, Courtauld s.n. (holotype, K).

Dracula medellinensis (Kraenzl.) Luer, Selbyana 2: 196. 1978.

Accepted Name: Dracula radiosa .

Type: Colombia, Boxal s.n. (holotype, W).

Dracula microglochin (Rchb.f.) Luer, Selbyana 2: 196. 1978.

Accepted Name: Dracula velutina .

Type: Colombia, Lehmann 3217 (holotype, W).

Dracula mosquerae (F.Lehm. & Kraenzl.) Luer, Selbyana 2: 196. 1978.

Accepted Name: Dracula houtteana .

Type: Colombia, Lehmann 7001 [lectotype, K-001457838]. Colombia, Lehmann 4302 (syntype).

Dracula platycrater (Rchb.f.) Luer, Selbyana 2: 196. 1978.

Accepted Name: Dracula spectrum .

Type: Colombia, Lehmann 3363 (holotype, W; isotypes, BR, BM, G).

Dracula quilichaoënsis (F.Lehm. & Kraenzl.) Luer, Selbyana 2(2/3): 197. 1978.

Accepted Name: Dracula trichroma .

Type: Colombia, Lehmann 2667 (holotype, K; isotype, BM, BR, G, MO, HBG-501778).

Dracula senilis (Rchb.f.) Luer, Selbyana 2: 197. 1978.

Accepted Name: Dracula chimaera .

Type: Colombia, Low & Co s.n. (holotype, K, W).

Dracula iricolor (Rchb.f.) Luer & R.Escobar, Selbyana 7: 65. 1982, sphalm.

Accepted Name: Dracula trichroma .

Type: Colombia, Roezl s.n. (holotype, W).

Note: This is a misspelling of the illegitimate name Dracula tricolor (see Karremans et al. 2019).

Dracula trinema (Rchb.f.) Luer, Selbyana 2: 198. 1978.

Accepted Name: Dracula velutina .

Type: Colombia, Lehmann s.n. (holotype, W).

Dracula vagabunda Luer & R.Escobar, Orquideología 15(1): 35-39

Accepted Name: Dracula pusilla .

Type: Colombia? [Uncertain origin], Escobar 1755 (holotype, JAUM; isotype, SEL).

Note: The type was initially assigned with doubt to Colombia, this taxon was later expressly excluded from the country by Luer (1993).

Dryadella misasii Luer & R.Escobar, Orquideología 13(2): 141-145. 1979.

Accepted Name: Dryadella gnoma .

Type: Colombia, Escobar 1451 (holotype, JAUM).

Dryadella popayanensis (F.Lehm. & Kraenzl.) Luer, Selbyana 2: 209. 1978.

Accepted Name: Dryadella simula .

Type: Colombia, Lehmann 184 (holotype, AMES; isotypes, G, K, NY, RJ, W).

Elongatia jimii (Luer) Luer, Harvard Pap. Bot. 17(2): 368. 2012, nom. inval.

Accepted Name: Pleurothallis lueriana .

Type: Colombia, Luer 20126 (holotype, MO).

Epibator mejiaSauleda, Uribe Vélez & Szlach., New World Orchid. Nomencl. Notes 117: 2. 2023.

Accepted Name: Zootrophion ximenae .

Type: Colombia? [uncertain origin], Mejia s.n. (holotype, HPUJ).

Lepanthes arethusaLuer & R.Escobar, Orquideología 21(2): 135-137. 1999.

Accepted Name: Lepanthes venusta .

Type: Colombia, Luer et al. 17633 (holotype, MO).

Lepanthes auspicataLuer & R.Escobar, Orquideología 21(2): 138-140. 1999.

Accepted Name: Lepanthes satyrica .

Type: Colombia, Silverstone-Sopkin et al. 2909 (holotype, CUVC; isotype, MO).

Lepanthes bifurcataLuer & R.Escobar, Orquideología 21(2): 140-142. 1999.

Accepted Name: Lepanthes georgii .

Type: Colombia, Luer 16639 (holotype, MO).

Note: Based on the same type number as L. georgii but strictly speaking not illegitimate because the holotype of the first is an isotype of the latter.

Lepanthes caetanoae O.Pérez, E.Parra & Kolan., Orquideología 27: 47. 2010.

Accepted Name: Lepanthes declivis .

Type: Colombia, Pérez et al. 512 (holotype, VALLE).

Lepanthes caucana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 159. 1924, nom. inval.

Accepted Name: Lepanthes tracheia .

Type: Colombia, Lehmann 3562 [holotype, B. destroyed; lectotype (selected here), BM-000082181; isolectotype, W-R-0054518].

Lepanthes dapaënsis P.Ortiz, O.Pérez & E.Parra, Orquideología 28(1): 23. 2011.

Accepted Name: Lepanthes tomentosa .

Type: Colombia, Pérez & Parra 751 (holotype, CUVC).

Lepanthes darienensis Sauleda & Uribe Vélez, New World Orchid. Nomencl. Notes 92: 1. 2020, nom. inval.

Accepted Name: Lepanthes sanjuanensis .

Note: This taxon is most probably a synonym of L. sanjuanensis, and was described without designating a type specimen. Mere citation of a locality and herbarium does not fulfill the requirement for valid publication.

Lepanthes dasyphylla Rchb.f., Flora 69: 557. 1886.

Accepted Name: Andinia pilosella .

Type: Colombia, Lehmann 3368 [lectotype (selected here), W-R-0014040; type illustr., W-R0054559].

Lepanthes elizabethiae O.Pérez, Kolan. & E.Parra, Phytotaxa 79(2): 58. 2013.

Accepted Name: Lepanthes lycocephala .

Type: Colombia, Pérez et al. 999 (holotype, CUVC).

Note: The description and accompanying illustration of this taxon correspond very well with Luer’s L. lycocephala (Luer 1996b, Luer & Thoerle 2012a).

Lepanthes guaduasensisLuer & R.Escobar, Orquideología 21(2): 145-147. 1999.

Accepted Name: Lepanthes macrostylis .

Type: Colombia, Escobar 5242 (holotype, MO).

Lepanthes hispida Luer & R.Escobar, Orquideología 16(2): 146. 1984.

Accepted Name: Lepanthes dunstervilleorum .

Type: Colomba, Luer et al. 7503 (holotype, SEL).

Lepanthes janus Luer & R.Escobar, Amer. Orchid Soc. Bull. 54(6): 730-732. 1985.

Accepted Name: Lepanthes norae .

Type: Colombia, Luer et al. 10283 (holotype, MO).

Lepanthes ospinaeGaray, Orquideología 4: 15, 84. 1969.

Accepted Name: Andinia nummularia .

Type: Colombia, Ospina 102a (holotype, AMES).

Lepanthes palmae Kolan., Szlach. & Medina Tr., Ann. Bot. Fenn. 59: 42. 2022.

Accepted Name: Lepanthes caudata .

Type: Colombia, Suarez et al. S19/27 (holotype, JAUM; isotype JAUM).

Note: The description and accompanying photographs of this taxon correspond very well with L. caudata (Luer 1996b), a species found in the Departments of Cauca, Nariño and Putumayo in southern Colombia that has been recorded to be variable both in shape and color of the flowers.

Lepanthes palmaritoensis Garay, Canad. J. Bot. 34: 251. 1956.

Accepted Name: Andinia pilosella .

Type: Colombia, Grant 9628 (holotype, US; isotypes, AMES, WIS).

Lepanthes queremalensis O.Pérez, Kolan. & Rinc.Useche, Syst. Bot. 38(2): 316. 2013.

Accepted Name: Lepanthes ankistra .

Type: Colombia, Pérez et al. 1100 (holotype, CUVC).

Note: The crispate leaf margin and subfalcate, tailed, and acute petals, and cymbiform appendix of the lip with an opening at the apex which are diagnostic of this taxon correspond very well with L. ankistra.

Lepanthes oteroi Luer, Orquideología 22(1): 42-45. 2001.

Accepted Name: Lepanthes elata .

Type: Colombia, Otero 274 (holotype, CUVC; isotype, MO).

Lepanthes peperomioides Schltr., Repert. Spec. Nov. Regni Veg. 7: 97. 1920.

Accepted Name: Andinia pilosella .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 33, No. 126)].

Lepanthes porphyrea Luer & R.Escobar, Amer. Orchid Soc. Bull. 53(9): 922. 1984.

Accepted Name: Lepanthes agglutinata .

Type: Colombia, Luer et al. 10071 (holotype, MO; isotype, SEL).

Lepanthes scopulifera Luer, Revista Soc. Boliv. Bot. 4(1): 9. 2003, nom. illeg.

Accepted Name: Lepanthes penicillifera .

Type: Colombia, Niessen & Uribe 036A (holotype, MO).

Lepanthes torquipetala Szlach. & Archila, Ann. Bot. Fenn. 56: 232. 2019.

Accepted Name: Lepanthes cyrtostele .

Type: Colombia, Cuatrecasas 20051B (holotype, COL; isotypes, COL, UGDA).

Note: This taxon is based solely on an herbarium specimen rehydrated and drawn by one person and described by others based only on that illustration. The extremely unusual petal morphology is therefore more likely a misinterpretation by the illustrator than something actually occurring in nature.

Lepanthes trachysepala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 100. 1920.

Accepted Name: Draconanthes aberrans .

Type: Colombia, Lehmann 6910 [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 34, No. 130)]. Colombia, Luer 12520 [neotype, MO (Luer 1996b), here rejected and selected as epitype].

Note: In a rather unconvincing treatment of the D. aberrans complex (Kolanowska et al. 2020b), the authors separate several taxa on the basis of slight differences in illustrations and photographs by several different authors, some very precise, others quite rudimentary, with no regards to properly documenting extant natural populations and assessing variation. Some of the illustrations published by the authors are mislabeled, compounding the confusion. We follow Pupulin et al. (2022) in recognizing Schlechter’s floral analyses reproduced by Mansfeld as original material and therefore reject Luer’s neotype in favor of a lectotype.

Lepanthes tricuspis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 101. 1920.

Accepted Name: Lepanthes mucronata .

Type: Colombia, Lehmann 4551 [holotype, B, destroyed; lectotype (selected here), K-000584318; isolectotypes, K-000584319, AMES-00100715, GH-00100716, MO-1106490].

Lothiania bilabiata Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 203. 1925.

Accepted Name: Masdevallia platyglossa .

Type: Colombia, Lehmann 3761 (holotype, G).

Masdevallia affinis Lindl., Orchid. Linden. 5. 1846.

Accepted Name: Masdevallia laevis .

Type: Colombia, Linden 1273 (holotype, K-L; isotypes, BR-0000006570439, G-00168825, W-R39210).

Masdevallia affinis subsp. maculigera (Schltr.) Luer, Lindleyana 3: 17. 1988.

Accepted Name: Masdevallia laevis .

Type: Colombia, Hopp 168 (holotype, B, destroyed). Colombia, Luer et al. 12551 (neotype, MO).

Masdevallia affinis subsp. petiolaris (Schltr.) Luer, Lindleyana 3: 17. 1988.

Accepted Name: Masdevallia laevis .

Type: Colombia, Madero s.n. (holotype, B, destroyed).

Masdevallia antioquiensis F.Lehm. & Kraenzl. ex Kraenzl., Bot. Jahrb. Syst. 26: 456. 1898.

Accepted Name: Masdevallia molossus .

Type: Colombia, Lehmann 7227 (holotype, K-000077750; isotypes, HBG-501770, LE00006496).

Masdevallia asperrima Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 133. 1922.

Accepted Name: Masdevallia melanoxantha .

Type: Colombia, Kalbreyer 757 (holotype, B, destroyed).

Masdevallia atroviolacea Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 130. 1922.

Accepted Name: Masdevallia mooreana .

Type: Colombia, Kalbreyer 1083 (holotype, B, destroyed). Colombia, Luer et al. 10198 [neotype, MO (Luer 2000c)].

Masdevallia auropurpurea Rchb.f. & Warsz., Bonplandia (Hannover) 2: 115. 1854.

Accepted Name: Masdevallia bicolor .

Type: Colombia, Warszewicz s.n. (holotype, W-R47222).

Masdevallia bathyschista Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 76. 1920.

Accepted Name: Masdevallia fasciata .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 25, No. 94)].

Masdevallia burfordiensis O’Brien, Gard. Chron. ser. 3, 33: 99-100. 1903.

Accepted Name: Masdevallia angulata .

Type: Ecuador, Lehmann s.n. (holotype, K-000077818).

Masdevallia backhousiana Rchb.f., Gard. Chron. 1: 716. 1879.

Accepted Name: Dracula chimaera .

Type: Colombia, Backhouse s.n. (holotypes, K, W).

Masdevallia bilabiata (Kraenzl.) Garay, Orquideología 5: 80. 1970.

Accepted Name: Masdevallia platyglossa .

Type: Colombia, Lehmann 3761 (holotype, G).

Masdevallia callifera Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 77. 1920.

Accepted Name: Dracula houtteana .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 25, No. 95)].

Masdevallia calopterocarpa Rchb.f., Flora 69: 560. 1886.

Accepted Name: Masdevallia amanda .

Type: Colombia (holotype, W).

Masdevallia calyptrata Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 1: 83. 1895.

Accepted Name: Masdevallia corniculata .

Type: Colombia (holotype, B, destroyed).

Masdevallia campyloglossa subsp. ortegesiana Luer, Lindleyana 3: 21. 1988.

Accepted Name: Masdevallia campyloglossa .

Type: Colombia, Ortgies s.n. (holotype, W-R39109).

Masdevallia carderi Rchb.f., Gard. Chron. 1: 784. 1883.

Accepted Name: Dracula inaequalis .

Type: Colombia, Carder s.n. (holotype, W).

Masdevallia carderi var. mosquerae F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 138. 1925.

Accepted Name: Dracula houtteana .

Type: Colombia, Lehmann 7001 [lectotype (selected here), K-001457838]. Colombia, Lehmann 4302 (syntype).

Masdevallia carderiopsis Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 422. 1921.

Accepted Name: Dracula houtteana .

Type: Colombia, Lehmann 2335 [lectotype (selected here), G-00168644; isolectotypes, BR0000009973237, W (not seen)].

Masdevallia casta Kraenzl., Notizbl. Bot. Gart. BerlinDahlem 8: 128. 1922.

Accepted Name: Masdevallia tubulosa .

Type: Colombia, Kalbreyer 1176 (holotype, B, destroyed).

Masdevallia chlorotica Kraenzl., Bull. Misc. Inform. Kew 107. 1925.

Accepted Name: Masdevallia laevis .

Type: Colombia, Lehmann 8506 (holotype, K-000077770; isotypes, AMES-00101223, G-00168819, HBG-501772, NY-00009050, W-R7962).

Masdevallia confusa Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 161. 1925.

Accepted Name: Masdevallia laevis .

Type: Colombia, Lehmann 522 (holotype, W; isotype, G).

Masdevallia corniculata var. inflata (Rchb.f.) H.J.Veitch, Man. Orchid. Pl. 5: 37. 1889.

Accepted Name: Masdevallia corniculata .

Type: Colombia, Bull s.n. (holotype, W).

Masdevallia chrysoneura F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 453. 1898.

Accepted Name: Masdevallia uncifera .

Type: Colombia, Lehmann 6753 [lectotype (selected here), K-000077774, excluding the illustration and far left plant element; isolectotypes, AMES00101227, LE-00006498, NY-00009051].

Note: Luer (2001a) selected a specimen at K as lectotype wihtout further specification. Two sheets bear tickets with the name M. chrysoneura in Kraenzlin’s writting, but neither is determined by him as type. K-000077774 has been determined as type by Garay. It is a mixed collection with some elements conforming with the protologue, selected here as lectotype. The second ticket is found on a type sheet of M. pantherina.

Masdevallia chrysochaete F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 159. 1925. Masdevallia chrysochaete F.Lehm., Bot. Jahrb. Syst. 26: 452. 1898, nom. inval.

Accepted Name: Masdevallia strumifera .

Type: Colombia, Lehmann 8868 [lectotype (selected here), K-000077803; isolectotype, AMES00082880 (not seen)]. Colombia, Lehmann 7034 (syntypes, AMES-00101226, LE-00001377).

Masdevallia cryptocopis Rchb.f. ex Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 182. 1925.

Accepted Name: Masdevallia picturata .

Type: Colombia, Kalbreyer 602 (holotype, W-R50757/W-0029150).

Masdevallia cucutillensis Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 127. 1922.

Accepted Name: Masdevallia caudata .

Type: Colombia, Kalbreyer 1156 (holotype, B, destroyed).

Masdevallia deorsum Rolfe, Orchid Rev. 8: 255. 1900.

Accepted Name: Masdevallia caesia .

Type: Colombia, Lehmann s.n. (holotype, K).

Note: This appears not to have a Latin description anywhere.

Masdevallia dermatantha Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 426. 1921.

Accepted Name: Masdevallia campyloglossa .

Type: Colombia (holotype, Woolward 1896, tab.12).

Masdevallia echinocarpa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 78-79. 1920.

Accepted Name: Diodonopsis erinacea .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral. analysis published by Mansfeld (1929: Pl. 26, No. 97)].

Masdevallia ellipes Rchb.f., Linnaea 41: 11. 1877.

Accepted Name: Masdevallia peristeria .

Type: Colombia, Roezl s.n. (holotype, W-R38721).

Masdevallia ephippium Rchb.f., Bot. Zeitung (Berlin) 390. 1873.

Accepted name: Masdevallia trochilus .

Type: Colombia, Wallis s.n. [lectotype (selected here), W-R-47258; tracing, W-R-21218].

Note: Reichenbach did not cite a collector in the 1873 protologue, but listed materials of Wallis, Roezl and Patin a year later in Xenia Orch. The plate reproduced there is clearly based on the Roezl material, but the original protologue does not describe a leaf, which indeed is abscent in the Wallis materials. Of the latter the author also prepared an unpublished sketch and therefore we believe it is more likely what he had originally at hand.

Masdevallia estradae var. delicata Rchb.f., Gard. Chron. n.s., 17: 525. 1882.

Accepted Name: Masdevallia estradae .

Type: Colombia, Bull s.n. [lectotype (selected here), W-R-50937, top illustr.].

Masdevallia fertilis Kraenzl., Kew Bull. 108. 1925.

Accepted Name: Masdevallia campyloglossa .

Type: Colombia, Lehmann 7037 (holotype, K; isotypes, BR, HBG-501773).

Masdevallia fissa Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 429. 1921.

Accepted Name: Masdevallia heteroptera .

Type: Colombia, Schmidtchen s.n. (holotype, W-R-11189).

Masdevallia flaccida Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 425. 1921.

Accepted Name: Masdevallia uncifera .

Type: Colombia, Lehmann s.n. (holotype, W-R83488).

Masdevallia frontinoensis Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 421. 1921.

Accepted Name: Masdevallia herradurae .

Type: Colombia, Sander s.n. (holotype, W-R15806).

Masdevallia gomeziana F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 434. 1921.

Accepted Name: Masdevallia laevis .

Type: Colombia, Lehmann 2700 [lectotype (selected here), W-R-80918; isolectotypes, AMES00101254, BM-00084406, BM-000084408, G-00168800, K-001458733, LE-00011167, LE00011165, NY-00009055].

Masdevallia gustavi Rchb.f., Gard. Chron. n.s., 3: 461. 1875.

Accepted Name: Masdevallia amanda .

Type: Colombia, Shuttleworth sub Bull s.n. [lectotype (selected here), W-R-47347, bottom specimens]. Colombia, Wallis s.n. (syntype, W-R47347, flowers top right).

Note: Reichenbach clearly cites syntypes collected by Wallis and Shuttleworth, both of which are found on a single sheet at W-R.

Masdevallia haematosticta Rchb.f., Flora 69: 559. 1886.

Accepted Name: Masdevallia peristeria .

Type: Colombia, Lehmann s.n. [lectotype (selected here), W-R-38735, type illustr.].

Masdevallia harryana Rchb.f., Gard. Chron. 1421. 1871.

Accepted Name: Masdevallia coccinea .

Type: Colombia, Veitch s.n. (holotype, W).

Masdevallia harryana var. carnea Linden, Lindenia 8: 95, t. 382. 1893.

Accepted Name: Masdevallia coccinea .

Masdevallia harryana var. illustris Linden, Lindenia 8: 95, t. 382. 1893.

Accepted Name: Masdevallia coccinea .

Masdevallia harryana var. kegeljani Linden, Lindenia 8: 95, t. 382. 1893.

Accepted Name: Masdevallia coccinea .

Masdevallia harryana var. laeta Rchb.f., Gard. Chron. n.s., 11: 716. 1879.

Accepted Name: Masdevallia coccinea .

Type: Colombia, Lawrence s.n. (holotype, W-R34073).

Masdevallia harryana var. lilacina Linden, Lindenia 8: 95, t. 382. 1893.

Accepted Name: Masdevallia coccinea .

Masdevallia harryana var. reginae Linden, Lindenia 8: 95, t. 382. 1893.

Accepted Name: Masdevallia coccinea .

Masdevallia heterotepala Rchb.f., Flora 69: 561. 1886.

Accepted Name: Masdevallia campyloglossa .

Type: Colombia, Lehmann 3655 (holotype, W-R14495).

Masdevallia hoppii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 24. 1924.

Accepted Name: Masdevallia pachyantha .

Type: Colombia, Hopp 141 (holotype, B, destroyed).

Masdevallia ignea var. marschalliana Rchb.f., Gard. Chron. 1872: 351. 1872.

Accepted Name: Masdevallia ignea .

Type: Colombia, Marshall s.n. [lectotype (selected here), W-R-47374, top right flower ‘72’].

Masdevallia ignea var. stobartiana Rchb.f., Gard. Chron. n.s., 15: 136. 1881.

Accepted Name: Masdevallia ignea .

Type: [Colombia] unknown collector [lectotype (selected here), W-R-31507].

Masdevallia ignea var. thomasonii Gentil, Pl. Cult. Serres Jard. Bot. Brux. 199. 1907.

Accepted Name: Masdevallia ignea .

Masdevallia inflata Rchb.f., Gard. Chron. 2: 716. 1881.

Accepted Name: Masdevallia corniculata .

Type: Colombia, Bull 674 (holotype, W-R-50957).

Masdevallia kalbreyeri Rchb.f. ex Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 169. 1925.

Accepted Name: Masdevallia urceolaris .

Type: Colombia, Kalbreyer 545 (holotype, W-R52945).

Masdevallia lactea Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 420. 1921.

Accepted Name: Dracula velutina .

Type: Colombia, Kalbreyer 1545 (holotype, B, destroyed).

Masdevallia lepida Rchb.f., Bonplandia (Hannover) 3: 69. 1855.

Accepted Name: Masdevallia laevis .

Type: Colombia, Wagener 575 (holotype, W-R38793; isotype, K-L).

Masdevallia lindenii André, Ill. Hort. 17: 226, t. 42. 1870.

Accepted Name: Masdevallia coccinea .

Type: Colombia, Wallis s.n.

Masdevallia lindenii var. grandiflora Linden, Lindenia 1: 73, t. 34. 1885.

Accepted Name: Masdevallia coccinea .

Masdevallia lindenii var. harryana (Rchb.f.) André, Ill. Hort. t. 142. 1872.

Accepted Name: Masdevallia coccinea .

Type: Colombia, Veitch s.n. (holotype, W).

Masdevallia lowii Rolfe, Gard. Chron. 1: 416. 1890.

Accepted Name: Dracula spectrum .

Type: Colombia, Courtauld s.n. (holotype, K).

Masdevallia macrochila Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 307. 1890.

Accepted Name: Dracula chestertonii .

Type: Colombia (holotype, LE).

Masdevallia maculigera Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 24. 1924.

Accepted Name: Masdevallia laevis .

Type: Colombia, Hopp 168 (holotype, B, destroyed). Colombia, Luer et al. 12551 (neotype, MO).

Masdevallia maxillariiformis F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 433. 1921.

Accepted Name: Masdevallia strumifera .

Type: Colombia, Lehmann 2452 [lectotype (selected here), W-R-0080914; isolectotypes, BM000082157, BR-0000006570170].

Note: Contrary to other taxa described by Kraenzlin, none of these materials had been annotated as type.

Masdevallia medellinensis Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 420. 1921.

Accepted Name: Dracula radiosa .

Type: Colombia, Boxal s.n. (holotype, W).

Masdevallia megaloglossa Luer & R.Escobar, Orquideología 15(2-3): 134. 1982.

Accepted Name: Masdevallia vargasii .

Type: Colombia, Escobar 1984 (holotype, JAUM; isotype, SEL).

Masdevallia metallica F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 70. 1925.

Accepted Name: Masdevallia caesia .

Type: Colombia, Lehmann 2689 [lectotype (selected here), BM-000084429; isolectotype, BR0000006570828].

Masdevallia microglochin Rchb.f., Flora 69: 558. 1886.

Accepted Name: Dracula velutina .

Type: Colombia, Lehmann 3217 (holotype, W).

Masdevallia militaris Rchb.f. & Warsz., Bonplandia (Hannover) 2: 115. 1854.

Accepted Name: Masdevallia coccinea .

Type: Colombia, Warszewicz s.n. (holotype, W-R0047333).

Masdevallia muriculata Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 99-100. 1925.

Accepted Name: Diodonopsis pygmaea .

Type: Colombia, Lehmann 2734 [lectotype (selected here), G-00168771].

Masdevallia ocanensis Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 429. 1921.

Accepted Name: Masdevallia picturata .

Type: Colombia, Bruchmüller s.n. (holotype, W-R14996/W-0029146).

Masdevallia ochthodes Rchb.f., Bonplandia (Hannover) 3: 70. 1855.

Accepted Name: Scaphosepalum verrucosum .

Type: Colombia, Wagener 564 (holotype, W).

Masdevallia oligantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 79. 1920.

Accepted Name: Masdevallia amanda .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 26, No. 99)].

Masdevallia olivacea Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 20. 1977.

Accepted Name: Masdevallia angulifera .

Type: Colombia, Lehmann 3207 (holotype, G; isotypes, BM-000082135, BR-0000006570743).

Masdevallia ortgiesiana Rchb.f. ex Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 74. 1925. Masdevallia ortgiesiana Rchb.f. ex Rolfe, Orchid Rev. 3: 48. 1895, nom. nud.

Type: Colombia, Ortgies s.n. [lectotype (selected here), W-R-39109, type illustr.]

Note: Kraenzlin validates Rchb.f.’s unpublished name. A lectotype is selected among the materials examined according to the protologue.

Masdevallia pachyantha var. hoppii (Schltr.) Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 63. 1925.

Accepted Name: Masdevallia pachyantha .

Type: Colombia, Hopp 141 (holotype, B, destroyed).

Masdevallia paisbambae F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 425. 1921.

Accepted Name: Masdevallia klabochiorum .

Type: Colombia, Lehmann 3719 (types BM, BR, G).

Masdevallia pallida (Woolward) Luer, Selbyana 2(4): 376. 1978.

Accepted Name: Masdevallia xanthina .

Type: Unknown, Lehmann s.n. (holotype, K).

Masdevallia palmensis Kraenzl., Bull. Misc. Inform. Kew 100. 1925.

Accepted Name: Masdevallia fasciata .

Type: Colombia, Lehmann 4286 (holotype, K-000077731; isotypes, G-00168773, HBG501776).

Masdevallia pantherina F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 454. 1898.

Accepted Name: Masdevallia laevis .

Type: Colombia, Lehmann 6752 [lectotype (selected here), K-000077772; isolectotypes, AMES00101291, AMES-00271898, G [not seen], K-000077773, LE-00006504, NY-00009061, US00093525, W-R-0007961].

Note: The materials of M. pantherina have been mixed with those of M. chrysoneura. There are tickets with the names of both species on sheets at K and W, sometimes on the same sheet.

Masdevallia petiolaris Schltr., Repert. Spec. Nov. Regni Veg. 7: 80. 1920.

Accepted Name: Masdevallia laevis .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 27, No. 100)].

Masdevallia peristeria subsp. haematosticta (Rchb.f.) Luer, Lindleyana 3(1): 44. 1988.

Accepted Name: Masdevallia peristeria .

Type: Colombia, Lehmann s.n. (holotype, W).

Masdevallia picturata var. minor Cogn., Fl. Bras. 3(4): 329-330. 1896.

Accepted Name: Masdevallia picturata .

Type: Colombia, Lehmann 2732 (holotype, BM000084425; isotypes, BR, G).

Masdevallia platycrater Rchb.f., Flora 69: 559. 1886.

Accepted Name: Dracula spectrum .

Type: Colombia, Lehmann 3363 (holotype, W; isotypes, BR, BM, G).

Masdevallia popayanensis F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 416. 1921.

Accepted Name: Dryadella simula .

Type: Colombia, Lehmann 184 (holotype, AMES; isotypes, G, K, NY, RJ, W).

Masdevallia porcelliceps Rchb.f., Gard. Chron. n.s., 19: 10. 1883.

Accepted Name: Masdevallia macroglossa .

Type: Colombia, O’Brien s.n. (holotype, W-R0039201).

Masdevallia quilichaoënsis F.Lehm. & Kraenzl., Bull. Misc. Inform. Kew 1925: 110-111. 1925.

Accepted Name: Dracula trichroma .

Type: Colombia, Lehmann 2667 (holotype, K; isotypes, BM, BR, G, MO, HBG-501778).

Masdevallia remotiflora Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 415. 1921.

Accepted Name: Masdevallia amanda .

Type: Colombia, Wallis 243 (holotype, W-R0014992).

Masdevallia restrepioidea Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 134. 1922.

Accepted Name: Masdevallia fasciata .

Type: Colombia, Kalbreyer 359 (syntype, B, destroyed); 1714 (syntype, B, destroyed).

Masdevallia roezlii var. rubra hort. ex Williams, Orchid Growers’ Man. ed. 6 6: 398. 1885.

Accepted Name: Dracula roezlii .

Masdevallia sarcophylla Kraenzl., Bull. Misc. Inform. Kew 100. 1925.

Accepted Name: Masdevallia campyloglossa .

Type: Colombia, Lehmann 10041 (holotype, K).

Masdevallia schmidtchenii Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 432. 1921.

Accepted Name: Masdevallia molossus .

Type: Colombia, Schmidtchen s.n. (holotype, W-R-0011186).

Masdevallia senilis Rchb.f., Gard. Chron. 2: 489. 1885.

Accepted Name: Dracula chimaera .

Type: Colombia, Low & Co s.n. (holotype, K, W ).

Masdevallia shuttleworthii Rchb.f., Gard. Chron. n.s., 3: 170. 1875.

Accepted Name: Masdevallia caudata .

Type: Colombia, Shuttleworth s.n. [lectotype (selected here), W-R-0050948, top right illustr.].

Masdevallia shuttleworthii var. xanthocorys Rchb.f., Gard. Chron. n.s., 17: 366. 1882.

Accepted Name: Masdevallia caudata .

Type: Colombia, Lawrence s.n. [lectotype (selected here), W-R-0050948, bottom right illustr.].

Masdevallia subumbellata Kraenzl., Rep. Spec. Nov. Beihefte 34: 107. 1925.

Accepted Name: Masdevallia bicolor .

Type: Colombia, Lehmann 4129 [lectotype (selected here), W-R-0007980; isolectotypes, AMES00106808, G-00168767). Ecuador, Hübsch s.n. (syntype, W-R-0016017).

Masdevallia sororcula Rchb.f., Gard. Chron. 1887(2): 713. 1887.

Accepted Name: Masdevallia mooreana .

Type: [Colombia], Lawrence s.n. (holotype, W-R0038791).

Masdevallia stenantha F.Lehm. & Kraenzl. ex Krendl, Bot. Jahrb. Syst. 26: 455. 1898.

Accepted Name: Masdevallia tubulosa .

Type: Colombia, Lehmann 6740 (holotype, K; isotypes, AMES, NY).

Masdevallia tenuipes Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 81. 1920.

Accepted Name: Masdevallia herradurae .

Type: Colombia, Madero, s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 27, No. 102)].

Masdevallia tricarinata F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26(5): 456. 1899.

Accepted Name: Pabstiella tripterantha .

Type: Colombia, Lehmann 8240 (holotype, B, destroyed).

Masdevallia trinema Rchb.f., Flora 69: 558. 1886.

Accepted Name: Dracula velutina .

Type: Colombia, Lehmann s.n. (holotype, W).

Masdevallia trinemoides Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 418. 1921.

Accepted Name: Masdevallia fasciata .

Type: Colombia, Schmidtchen s.n. (holotype, W).

Masdevallia trionyx Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 131. 1922.

Accepted Name: Masdevallia falcago .

Type: Colombia, Kalbreyer 738 (holotype, B).

Masdevallia ventricularia var. brevicaudata Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 19. 1925.

Accepted Name: Masdevallia ventricularia .

Type: Colombia, Lehmann 23 (holotype, W-R0083470).

Note: Lehmann 2006, 4137, 4310, 4311, 7032, Schmidtchen s.n., and Kalbreyer 1529 have been cited as syntypes. But these don’t fit Kraenzlin’s description and we believe he was merely listing these as materials studied under the name M. ventricularia. At W-R, Kraenzl. explicitly determined a holotype for this taxon.

Masdevallia ventricularia var. longicaudata F.Lehm., Gard. Chron. III, 23: 238. 1898.

Accepted Name: Masdevallia filaria .

Note: This name was never published.

Masdevallia wageneriana var. colombiana Braas, Orquideología 12: 27. 1977.

Accepted Name: Masdevallia pteroglossa .

Type: Colombia, Escobar 1161 (holotype, JAUM).

Masdevallia wallisii var. stupenda Rchb.f., Gard. Chron. 1: 270. 1885.

Accepted Name: Dracula chimaera .

Type: Colombia, Shuttleworth s.n. (holotype, W).

Masdevallia winniana Rchb.f., Gard. Chron. 2: 198. 1881.

Accepted Name: Dracula roezlii .

Type: Colombia, Winn s.n. (holotype, K, W).

Masdevallia xanthina var. pallida Woolward, Monogr. Genus Masdevallia . 1896.

Accepted Name: Masdevallia xanthina .

Type: Unknown, Lehmann s.n. (holotype, K).

Masdevallia xerophila F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 431. 1921.

Accepted Name: Masdevallia pteroglossa .

Type: Colombia, Lehmann 125 (holotype, W-R0083529).

Masdevallia xipheres Rchb.f., Linnaea 41: 12. 1877.

Accepted Name: Porroglossum muscosum .

Type: Colombia, Roezl s.n. (holotype, W-R0050781).

Muscarella antioquiensis Sauleda, Uribe Vélez & Szlach., New World Orchid. Nomencl. Notes 118: 2. 2023.

Accepted Name: Andreettaea zephyrina .

Type: Colombia, Mejia s.n. (holotype, HPUJ).

Muscarella tsubotae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 105: 121-122. 2006.

Accepted Name: Andreettaea barberiana .

Type: Colombia, Luer 16763 (holotype, MO).

Neooreophilus medinae Kolan. & Szlach., Diversity 14(372): 20. 2022.

Accepted Name: Andinia ursula .

Type: Colombia, Medina T. S14/28 (holotype, JAUM; photo of type, MEDEL).

Note: Inspection of specimens from the type locality of N. medinae and A. ursula reveal the two taxa are indistinguishable.

Octomeria longerepens Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 58-59. 1924.

Accepted Name: Octomeria graminifolia .

Type: Colombia, Hopp 80 (holotype, B, destroyed). Colombia, Hirtz 4128 [neotype, MO (Luer 2010)].

Octomeria schultesii Pabst, Arq. Bot. Estado São Paulo 3(5): 268-269, pl. 67, f. B. 1962.

Accepted Name: Octomeria longifolia .

Type: Colombia, Schultes & López 10148 (holotype, IAN; isotype, AMES).

Platystele cuculligera P.Ortiz, Orquideología 14(3): 210. 1981.

Accepted Name: Platystele alucitae .

Type: Colombia, Ortiz 969 (holotype, COL).

Platystele culex Luer & R.Escobar, Phytologia 49(3): 198. 1981.

Accepted Name: Platystele examen-culicum .

Type: Colombia, Luer 2313 (holotype, SEL).

Platystele escobariana Garay, Orquideología 6: 15. 1971.

Accepted Name: Platystele schmidtchenii .

Type: Colombia, Escobar 155 (holotype, AMES).

Platystele gemmula Luer, Phytologia 49(3): 198. 1981.

Accepted Name: Platystele microglossa .

Type: Colombia, Luer et al. 4581 (holotype, SEL).

Platystele hypsitera Luer & R.Escobar, Orquideología 16(1): 14. 1983.

Accepted Name: Platystele pisifera .

Type: Colombia, Luer et al. 7898 (holotype, SEL).

Platystele porphyroglossa P.Ortiz, Orquideología 14(3): 216. 1981.

Accepted Name: Platystele gyroglossa .

Type: Colombia, Ortiz 971 (holotype, COL).

Pleurothallis andrei Luer & R.Escobar, Orquideología 20: 34. 1996.

Accepted Name: Pleurothallis lindenii .

Type: Colombia, Luer et al. 16744 (holotype, MO).

Pleurothallis antioquiensis (Schltr.) P.H. Allen, Revista Fac. Nac. Agron., Medellin 8(32): 442. 1948.

Accepted Name: Restrepia sanguinea .

Type: Colombia, Madero s.n. (holotype, B, destroyed).

Pleurothallis apoxys Luer & R.Escobar, Orquideología 14(2): 132. 1981.

Accepted Name: Pleurothallis supervacanea .

Type: Colombia, Luer 2087 (holotype, SEL; isotype, JAUM).

Pleurothallis arachnion Luer, Phytologia 47(2): 74. 1980.

Accepted Name: Pleurothallis quadricaudata .

Type: Colombia, Luer et al. 4606 (holotype, SEL).

Pleurothallis arevaloi Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 43-44. 1924.

Accepted Name: Specklinia picta .

Type: Colombia, Hopp 121,161 (holotype, B, destroyed).

Pleurothallis asymmetrica (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 135. 2021.

Accepted Name: Pleurothallis xanthochlora .

Type: Colombia, Garay et al. 223 (holotype, AMES; drawing, UGDA).

Pleurothallis aves-seriales Luer & R.Escobar, Orquideología 14(2): 134. 1981.

Accepted Name: Pleurothallis penduliflora .

Type: Colombia, Escobar et al. 956 (holotype, JAUM).

Pleurothallis birchenallii Rolfe, Bull. Misc. Inform. Kew 365. 1909.

Accepted Name: Pleurothallis glossopogon .

Type: Colombia, Birchenall s.n. (holotype, K).

Pleurothallis biserrula Rchb.f., Bonplandia (Hannover) 3: 71. 1855.

Accepted Name: Pleurothallis glossopogon .

Type: Colombia, Wagener s.n. [L551] (holotype, W-R-0053225; isotype; K-L).

Pleurothallis bogotensis Lindl., Orchid. Linden. 1. n. 3. 1846.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Linden 1271 (holotype, K-L; isotype, W-R-0053235!, left).

Pleurothallis bogotensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 162-163. 1924, nom. illeg.

Accepted Name: Stelis pulchella .

Type: Colombia, Hopf s.n. (holotype, B, destroyed). Colombia, Hartweg s.n. [neotype, K (Luer 1998a)].

Pleurothallis brachyantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 44. 1924.

Accepted Name: Anathallis citrina .

Type: Colombia: Hopp 21 (holotype, B; destroyed).

Pleurothallis brevicaulis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 101. 1920.

Accepted Name: Specklinia gracillima .

Type: Colombia, Lehmann 8205 [holotype, B, destroyed; lectotype, AMES-02151693 (Luer 2006); islectotype, K-000584172].

Pleurothallis caliensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 102-103. 1920.

Accepted Name: Pleurothallopsis tubulosa .

Type: Colombia, Lehmann 8226 [holotype, probably destroyed at B; lectotype (selected here), K-000583672; isolectotype, K-000583671; illustration of type, Schlechter’s floral analysis published by Mansfeld (1929: Pl. 35, No. 133)].

Pleurothallis cardiantha Rchb.f., Linnaea 41: 15. 1877 [1876].

Accepted Name: Pleurothallis bivalvis .

Type: Colombia, Roezl s.n. (holotype, W-R0025680).

Pleurothallis chachatoynsis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 45. 1924.

Accepted Name: Pleurothallis microcardia .

Type: Colombia, Hopp 48 (holotype, B, destroyed).

Pleurothallis chloroleuca Lindl., Orchid. Linden. 2. 1846.

Accepted Name: Pleurothallis laurifolia .

Type: Colombia, Linden 1269 (holotype, K-L000079869; isotypes, GENT-10168516, W-R-0024877).

Pleurothallis chlamydopus Schltr., Repert. Spec. Nov. Regni Veg. 7: 103. 1920.

Accepted Name: Stelis ligulata .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 35, No. 134)]. Colombia, Lehmann 4508 [neotype, AMES; isoneotype, NY (Luer 1998a), here rejected and selected as epitypes].

Pleurothallis corralensis Garay, Arch. Jard. Bot. Rio de Janeiro 13: 42. 1955.

Accepted Name: Pleurothallis coriacardia .

Type: Colombia, Lehmann 8207 [holotype, B, destroyed; lectotype, K; isolectotype, AMES (Luer 2005)].

Pleurothallis cundinamarcae Schltr., Notizbl. Bot. Gart. Berlin-Dahlem 7: 528. 1921.

Accepted Name: Pleurothallis microcardia .

Type: Colombia, Schnitter s.n. (holotype, B, destroyed).

Pleurothallis cyclochila Lindl., Fol. Orchid. ~Pleurothallis~ 8. 1859.

Accepted Name: Pleurothallis antennifera .

Type: Colombia, Triana 124 (holotype, K-L; isotype, W-R-0053853; tracing of type, W-R0007810).

Pleurothallis cymbicalli Pabst, Arq. Bot. Estado São Paulo 3(5): 268, t. 67, f. A. 1962.

Accepted Name: Myoxanthus trachychlamys .

Type: Colombia, Schultes 8584 (holotype, IAN; isotype, AMES).

Pleurothallis daguensis F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26(3-4): 444. 1899.

Accepted Name: Acianthera ciliata .

Type: Colombia, Lehmann 8167 (holotype, B; destroyed).

Pleurothallis decora Luer & R.Escobar, Orquideología 16(1): 20. 1983.

Accepted Name: Pleurothallis nutans .

Type: Colombia, Luer et al. 7922 (holotype, SEL; isotype, JAUM).

Pleurothallis decurva Lindl., Fol. Orchid. ~Pleurothallis~ 8. 1859.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Jameson 19 (holotype, K-L).

Pleurothallis elegans (Kunth) Lindl., Edwards’s Bot. Reg. 28(Misc): 70. 1842.

Accepted Name: Stelis roseopunctata .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; fragment, W).

Pleurothallis espinalii Luer, Orquideología 21(2): 122-125. 1999.

Accepted Name: Stelis convoluta .

Type: Colombia, Espinal & Ramos 3041 (holotype, CUVC).

Pleurothallis falcipetala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 163. 1924.

Accepted Name: Gravendeelia chamaelepanthes .

Type: Colombia, Lehmann 2045 (holotype, B, destroyed; lectotype (selected here), BM-000082184. Colombia, Lehmann 8245 [neotype, K (Luer 1997), here rejected].

Note: Among the indet materials at BM we found an isotype of this obscure taxon and determined it as G. chamaelepanthes. Based on the description, Luer mistakenly believed this name applied to Trichosalpinx (Stellamaris) pergrata. The neotype is rejected here.

Pleurothallis florulenta Linden & Rchb.f., Bonplandia (Hannover) 3: 223. 1855.

Accepted Name: Specklinia picta .

Type: Colombia, Linden 55 (holotype,W; isotype, AMES).

Pleurothallis fornix Luer & R.Escobar, Orquideología 16(2): 160. 1984.

Accepted Name: Pleurothallis calceolaris .

Type: Colombia, Escobar et al. 8758 (holotype, SEL).

Pleurothallis fuscata Braid, Bull. Misc. Inform. Kew 1924: 200-201. 1924.

Accepted Name: Pleurothallis bivalvis .

Type: Colombia (holotype, K).

Pleurothallis granularis Luer & R.Escobar, Orquideología 16(2): 162. 1984.

Accepted Name: Pleurothallis platysepala .

Type: Colombia, Luer 8183 (holotype, SEL).

Pleurothallis hartwegii Lindl., Ann. Mag. Nat. Hist. 15: 106. 1845, nom. illeg.

Accepted Name: Stelis laevigata .

Type: Colombia, Hartweg 1405 (holotype, K-L000674224; isotypes, BM-000082364; LD-1246759, W-R-0053633).

Pleurothallis hawkesii Flickinger, Orchid Rev. 71: 336. 1963.

Accepted Name: Restrepia brachypus .

Type: Colombia, Moore s.n. (lectotype, K).

Pleurothallis hemirhoda Lindl. & Paxton, Paxt. Fl. Gard. 3: 133, no. 612, f. 299. 1852.

Accepted Name: Pleurothallis nuda .

Type: Colombia, Linden s.n. (holotype, W; type illustration, K-L).

Pleurothallis hirtzii Luer, Phytologia 47(2): 76. 1980.

Accepted Name: Pleurothallis colossus .

Type: Colombia, Luer et al. 4618 (holotype, SEL).

Pleurothallis hopfiana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 164. 1924.

Accepted Name: Stelis ligulata .

Type: Colombia, Hopf s.n. (holotype, B, destroyed). Colombia, Luer et al. 6658 [neotype, SEL (Luer 1998a)].

Pleurothallis hoppii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 46. 1924.

Accepted Name: Pleurothallis glossopogon .

Type: Colombia, Hopp 122 (holotype, B, destroyed).

Pleurothallis humidicolaGaray, Svensk Bot. Tidskr. 47: 212. 1953.

Accepted Name: Specklinia gracillima .

Type: Colombia, Sneidern 109 (holotype, S).

Pleurothallis ineziae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 47. 1924.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Hopp 10 (holotype, B, destroyed).

Pleurothallis jimiiLuer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 237, 242, f. 9. 2004, nom. inval.

Accepted Name: Pleurothallis lueriana .

Type: Colombia, Luer 20126 (holotype, MO).

Pleurothallis lancifera Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 48-49. 1924.

Accepted Name: Trichosalpinx orbicularis .

Type: Colombia, Hopp 67 (holotype, B, destroyed).

Pleurothallis langlassei Schltr., Repert. Spec. Nov. Regni Veg. 7: 105. 1920.

Accepted Name: Stelis cassidis .

Type: Colombia, Langlassé 97 [holotype, B, destroyed; lectotype, AMES; isolectotypes, G, K, US (Luer 1998a)].

Pleurothallis lansbergii Regel, Index Sem. (St. Petersburg) 17. 1856.

Accepted Name: Pleurothallis bivalvis .

Type: Colombia (holotype, LE).

Pleurothallis lauta Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 105. 1920.

Accepted Name: Muscarella zephyrina .

Type: Colombia, Lehmann 8206 [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 36, No. 138)].

Pleurothallis leontoglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 106. 1920, nom. illeg.

Accepted Name: Pleurothallis coriacardia .

Type: Colombia, Lehmann 8207 [holotype, B, destroyed; lectotype, K; isolectotype, AMES (Luer 2005)].

Pleurothallis lepanthoides Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 106-107. 1920.

Accepted Name: Karma arbuscula .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 37, No. 140)].

Pleurothallis lindenii var. pumila Rchb.f., Bonplandia (Hannover) 3: 72. 1855.

Accepted Name: Pleurothallis inflata .

Type: Colombia, Wagener s.n. (holotype, K).

Pleurothallis listrostachys Rchb.f., Bonplandia (Hannover) 3: 71. 1855.

Accepted Name: Stelis sclerophylla .

Type: Colombia, Wagener 561 (holotype, W; isotypes, BR, K-L).

Pleurothallis mariae I. Bock, Orchidee (Hamburg) 36(3): 128. 1981.

Accepted Name: Pleurothallis discoidea .

Type: Colombia, Bock s.n. (holotype, BOCK).

Pleurothallis medellinensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 236. 1920.

Accepted Name: Pabstiella tripterantha .

Type: Colombia, Lehmann 8240 (holotype, B, destroyed).

Pleurothallis monocardia Rchb.f., Bonplandia (Hannover) 3: 72. 1855.

Accepted Name: Pleurothallis cordata .

Type: Colombia, Wagener s.n. (holotype, W).

Pleurothallis nasuta Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 164. 1924.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Schnnitter s.n. (holotype, B?, destroyed).

Pleurothallis navicularis Lindl., Fol. Orchid. ~Pleurothallis~ 6-7. 1859.

Accepted Name: Pleurothallis loranthophylla .

Type: Colombia, Purdie s.n. (holotype, K; illustration of type, K-L).

Pleurothallis northenae Luer, Amer. Orchid Soc. Bull. 45(3): 201. 1976.

Accepted Name: Pleurothallis latipetala .

Type: Colombia, Luer 031 (holotype, SEL).

Pleurothallis nubigena Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 50. 1924, nom. illeg.

Accepted Name: Stelis ligulata .

Type: Colombia, Hopp 30 (holotype, B, destroyed). Colombia, Lehmann 6924 [neotype, AMES; isoneotypes, NY, US (Luer 1998a)].

Pleurothallis ochroleuca Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 108. 1920.

Accepted Name: Stelis aurea .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 37, No. 143)].

Pleurothallis ospinae R.E. Schult., Caldasia 7(35): 354. 1957.

Accepted Name: Restrepia antennifera .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Pleurothallis papillisepala Schltr., Repert. Spec. Nov. Regni Veg. 7: 109. 1920.

Accepted Name: Stelis convoluta .

Type: Colombia, Lehmann 4518 [holotype, B, destroyed; lectotype (selected here), K-000584304; isolectotypes, AMES00074589, LE-00001422, US-00093672].

Pleurothallis patula Schltr., Repert. Spec. Nov. Regni Veg. 7: 110. 1920.

Accepted Name: Stelis ligulata .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 38, No. 145)]. Colombia, Lehmann 4508 [neotype, AMES (Luer 1998a), here rejected].

Pleurothallis pendula Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 51. 1924.

Accepted Name: Pleurothallis lindenii .

Type: Colombia, Hopp 172 (holotype, B, destroyed).

Pleurothallis phaeantha Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 52. 1924.

Accepted Name: Pleurothallis bivalvis .

Type: Colombia, Hopp 1 (holotype, B, destroyed).

Pleurothallis platycardium Schltr., Repert. Spec. Nov. Regni Veg. 7: 529. 1921.

Accepted Name: Pleurothallis cordata .

Type: Colombia, Schnitter s.n. (holotype, S).

Pleurothallis pleiostachys Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 53. 1924.

Accepted Name: Stelis cassidis .

Type: Colombia, Hopp 174 (holotype, B, destroyed). Colombia, Jameson 398 [neotype, AMES, isonetoype, K (Luer 1998a)].

Pleurothallis potamophila Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 53. 1924.

Accepted Name: Stelis rhodotantha .

Type: Colombia, Hopp 119 (holotype, B, destroyed). Colombia, Lehmann 4502 [neotype, K; isonetoypes, AMES, LE-00001424 (Luer 1998a)].

Pleurothallis pteroglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 112. 1920.

Accepted Name: Stelis pulchella .

Type: Colombia, Triana 565 [holotype, B, destroyed; lectotype, US (Luer 1998a)].

Pleurothallis pristis F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 443. 1899.

Accepted Name: Stelis dialissa .

Type: Colombia, Lehmann 6032 [holotype, B, destroyed; lectotype, AMES (Luer 2009a); lectotype, AMES-14772 (Luer 2018c)].

Pleurothallis puberula Klotzsch, Allg. Gartenzeitung 22(50): 393-394. 1854.

Accepted Name: Restrepiella ophiocephala .

Type: Colombia, Warscewicz s.n. (holotype?).

Pleurothallis pulvinipes Schltr., Notizbl. Bot. Gart. Berlin-Dahlem 7: 529. 1921.

Accepted Name: Pleurothallis macrocardia .

Type: Colombia, Schnitter s.n. (holotype, S).

Pleurothallis pyxos Luer & R.Escobar, Orquideología 14: 168. 1981.

Accepted Name: Karma amygdalora .

Type: Colombia, Luer et al. 3047 (holotype, SEL; isotype, JAUM).

Pleurothallis reichenbachiana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 8: 131. 1921.

Accepted Name: Pleurothallis laurifolia .

Type: Colombia, Roezl s.n. (holotype, W; isotype, AMES).

Pleurothallis rhaphidopus Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 54-55. 1924.

Accepted Name: Pleurothallis phyllocardioides .

Type: Colombia, Hopp 83 (holotype, B, destroyed).

Pleurothallis rhodocardiaGaray, Svensk Bot. Tidskr. 47(2): 204-206. 1953.

Accepted Name: Pleurothallis rowleei .

Type: Colombia, von Sneidern 1603 (holotype, S; isotype, AMES).

Pleurothallis rhomboglossa Rchb.f., Flora 71(10): 154. 1888, nom. illeg.

Accepted Name: Pleurothallis laurifolia .

Type: Colombia, Roezl s.n. (holotype, W; isotype, AMES).

Pleurothallis roezlii Rchb.f., Linnaea 61: 13. 1877.

Accepted Name: Pleurothallis macrophylla .

Type: Colombia, Röezl s.n. (holotype, P).

Pleurothallis ruscifolia var. caquetana Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 55. 1924. Accepted Name: Pleurothallis ruscifolia.

Type: Colombia, Hopp 72 (holotype, B, destroyed).

Pleurothallis scapha Rchb.f., Gard. Chron. 2: 162. 1874.

Accepted Name: Pleurothallis sirene .

Type: Colombia, Day 358 (holotype, W).

Pleurothallis scaphioglottis Schltr., Repert. Spec. Nov. Regni Veg. 7: 113. 1920.

Accepted Name: Pleurothallis canaligera .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 39, No. 150)].

Pleurothallis schistopetala Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 113. 1920.

Accepted Name: Muscarella zephyrina .

Type: Colombia, Smith 2482 (holotype, US; isotype, BM-000082524; BR, K, NY).

Pleurothallis schnitteri Schltr., Notizbl. Bot. Gart. Berlin-Dahlem 7: 530. 1921.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Schnnitter s.n. (holotype, B?, destroyed).

Pleurothallis smithii Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 114. 1920.

Accepted Name: Pleurothallis coriacardia .

Type: Colombia, Smith 2362 (holotype, K; isotypes, AMES, NY, US ).

Pleurothallis sphenochila Lindl., Fol. Orchid. ~Pleurothallis~ 22. 1859.

Accepted Name: Stelis pulchella .

Type: Colombia, Schlim 1031 (holotype, K).

Pleurothallis stelidioides Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 116. 1920.

Accepted Name: Stelis magdalenae .

Type: Colombia, Lehmann 8198 [holotype, B, destroyed; lectotype, AMES-14836/AMES00074771 (Luer 1999a emend Toscano de Brito 2018); isolectotypes, AMES-00074772, G-00168981, HB [not seen], K-000584276, US00093702! W-R-0007966].

Pleurothallis stenophylla F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 442. 1899.

Accepted Name: Stelis melanopus .

Type: Colombia, Lehmann 4515 [lectotype (selected here), K-000584275; isolectotypes, LE00006522, US-00093703].

Pleurothallis sugdenii Luer, Orquideología 20(2): 229.

Accepted Name: Pleurothallis discoidea .

Type: Colombia, Sugden 34 (holotype, K).

Pleurothallis superposita Schltr., Repert. Spec. Nov. Regni Veg. 14: 386. 1916.

Accepted Name: Stelis catenata .

Type: Ecuador, Sodiro 29 [holotype, B, destroyed; lectotype, QPLS (Luer 1999a)].

Pleurothallis trianae Schltr., Repert. Spec. Nov. Regni Veg. 7: 117. 1920.

Accepted Name: Stelis galeata .

Type: Colombia, Triana 569 [holotype, B, destroyed; lectotype, US (Luer 1998a); isolectotypes, P-00489121, P-00489122, COL-000363332].

Pleurothallis tsubotae Luer & R.Escobar, Orquideología 20: 88. 1996.

Accepted Name: Andreettaea barberiana .

Type: Colombia, Luer 16763 (holotype, MO).

Pleurothallis trachytheca F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26(3-4): 444-445. 1899.

Accepted Name: Trichosalpinx orbicularis .

Type: Colombia, Lehmann 8224 [holotype, B, destroyed; lectotype, K-000584211; isolectotypes, AMES-00074813, AMES-00287046, NY00009245 (Luer 1997)].

Pleurothallis tunguraguae var. minor Garay, Canad. J. Bot. 34(2): 255. 1956.

Accepted Name: Stelis tunguraguae .

Type: Colombia, Aristide-Joseph A194 (holotype, US-888402).

Pleurothallis urosepala F.Lehm. & Kraenzl., Bot. Jahrb. Syst. 26: 442. 1899.

Accepted Name: Stelis sclerophylla .

Type: Colombia, Lehmann 6065 (holotype, K-000584267; isotypes, AMES00074828, AMES-00074829, AMES-00074830, AMES00074832, AMES-00074833, AMES-00074834, BM-000074326, K-000584266, K-000584268, LE-00006524, NY-00009249, US-00093715).

Pleurothallis venosa Rolfe, Kew Bull. 30. 1906.

Accepted Name: Stelis galeata .

Type: Colombia, Sander s.n. (holotype, K; isotype, AMES).

Pleurothallis wendlandiana Rchb.f., Flora 71(9): 153-154. 1888.

Accepted Name: Pleurothallis laurifolia .

Type: Colombia, Wendland s.n. [lectotype (selected here), W-R-0024879, type illustr.].

Pleurothallis werneri Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 57. 1924.

Accepted Name: Pleurothallis phalangifera .

Type: Colombia, Hopp 90 (holotype, B, destroyed).

Porroglossum colombianum Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 83. 1920.

Accepted Name: Porroglossum mordax .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 27, No. 103)].

Porroglossum xipheres (Rchb.f.) Garay, Bot. Mus. Leafl. 21: 252. 1967.

Accepted Name: Porroglossum muscosum .

Type: Colombia, Roezl s.n. (holotype, W).

Restrepia antennifera var. angustifolia Kraenzl., Bot. Jahrb. Syst. 26: 451. 1899.

Accepted Name: Barbosella cucullata .

Type: Colombia, Lehmann 6883 (lectotype, K).

Restrepia antennifera subsp. erythroxantha (Rchb.f.) Mohr, Leafl. Schlechter Inst. 2: 13. 1996.

Accepted Name: Restrepia elegans .

Type: Venezuela, Moritz 854 (holotype, W; isotype, G).

Restrepia antennifera subsp. hemsleyana (Schltr.) H.Mohr, Leafl. Schlechter Inst. 2: 13. 1996.

Accepted Name: Restrepia antennifera .

Type: Colombia, Sander s.n. (holotype, K).

Restrepia antennifera subsp. leontoglossa (Schltr.) H.Mohr, Leafl. Schlechter Inst. 2: 13. 1996.

Accepted Name: Restrepia trichoglossa .

Type: Colombia, Madero s.n. (holotype, B, destroyed). Colombia, Luer 17618 [neotype, MO (Luer 1996a)].

Restrepia antennifera subsp. striata (Rolfe) H.Mohr, Leafl. Schlechter Inst. 2: 13. 1996.

Accepted Name: Restrepia brachypus .

Type: Colombia, Moore s.n. (lectotype, K).

Restrepia antioquiensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 118. 1920.

Accepted Name: Restrepia sanguinea .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 42, No. 159)].

Restrepia aristulifera f. leathersii Luer, Harvard Pap. Bot. 17(2): 360. 2012.

Accepted Name: Restrepia aristulifera .

Type: Colombia, Leathers s.n. (holotype, MO).

Restrepia biflora Regel, Ann. Sci. Nat., Bot., sér. 4, 6: 373. 1856.

Accepted Name: Pleurothallis nuda .

Type: Colombia, Lamberg s.n. (holotype, LE).

Restrepia brachypus subsp. serrilabia (Schltr.) H.Mohr, Leafl. Schlechter Inst. 2: 14. 1996.

Accepted Name: Restrepia trichoglossa .

Type: Colombia, Madero s.n. (holotype, B, destroyed). Colombia, Luer 17548 [neotype, MO (Luer 1996a)].

Restrepia caespitifica F.Lehm. & Kraenzl., Repert. Spec. Nov. Regni Veg. 24(22-25): 358-359. 1928.

Accepted Name: Barbosella prorepens .

Type: Colombia, Lehmann 3305 (holotype, lost?).

Restrepia caucana Schltr., Repert. Spec. Nov. Regni Veg. 7: 119. 1920.

Accepted Name: Restrepia contorta .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 42, No. 160)]. Colombia, Luer 16681 [neotype, MO (Luer 1996a), here rejected and selected as epitype].

Restrepia dentata Rolfe, Bull. Misc. Inform. Kew 208. 1892.

Accepted Name: Restrepia aspasicensis .

Type: Colombia, Moore s.n. (holotype, K).

Restrepia hemsleyana Schltr., Repert. Spec. Nov. Regni Veg. 7: 199. 1920.

Accepted Name: Restrepia antennifera .

Type: Colombia, Sander s.n. (holotype, K).

Restrepia leontoglossa Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 120. 1920.

Accepted Name: Restrepia trichoglossa .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 42, No. 161)]. Colombia, Luer 17618 [neotype, MO (Luer 1996a), here rejected and selected as epitype].

Restrepia maculata Lindl., Orchid. Linden. 4. 1846.

Accepted Name: Restrepia contorta .

Type: Colombia, Linden 1270 [lectotype, K (Luer 1996a); isolectotypes, BR, W].

Restrepia muscifera subsp. shuttleworthii (Rolfe) H.Mohr, Leafl. Schlechter Inst. 2: 8, t. 2. 1996. Accepted Name: Restrepia muscifera.

Type: Colombia, Shuttleworth s.n. (holotype, K).

Restrepia schlimii Rchb.f., Linnaea 41: 45. 1877.

Accepted Name: Restrepia nittiorhyncha .

Type: Colombia, Schlim 1158 (holotype, K; isotype, G).

Restrepia serrilabia Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 121. 1920.

Accepted Name: Restrepia trichoglossa .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 42, No. 162)]. Colombia, Luer 17548 [neotype, MO (Luer 1996a), here rejected and selected as epitype].

Restrepia shuttleworthii Rolfe, Bull. Misc. Inform. Kew 1892(65-66): 138-139. 1892.

Accepted Name: Restrepia muscifera .

Type: Colombia, Shuttleworth s.n. (holotype, K).

Restrepia striata Rolfe, Gard. Chron., ser. 3 9: 137. 1891.

Accepted Name: Restrepia brachypus .

Type: Colombia, Moore s.n. (lectotype, K).

Restrepia vittata Lindl., J. Hort. Soc. London 3: 314. 1848.

Accepted Name: Pleurothallis nuda .

Type: Colombia, Linden s.n. (holotype, W).

Restrepia vittata var. biflora Regel, Ann. Sci. Nat., Bot., sér. 4, 6: 373. 1856.

Accepted Name: Pleurothallis nuda .

Type: Colombia, Lamberg s.n. (holotype, LE).

Restrepiella bicallosa (Luer & R.Escobar) Braas & H.Mohr, Orchideen 33: 154. 1982.

Accepted Name: Pleurothallopsis norae .

Type: Colombia, Luer 2332 (holotypes, SEL, JAUM ).

Restrepiopsis bicallosa Luer & R.Escobar, Selbyana 2: 201. 1978.

Accepted Name: Pleurothallopsis norae .

Type: Colombia, Luer 2332 (holotypes, SEL, JAUM ).

Scaphosepalum antioquiense Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 135. 1922.

Accepted Name: Porroglossum mordax .

Type: Colombia, Kalbreyer 1715; 2361 (syntypes, B, destroyed).

Scaphosepalum escobarianum Garay, Orquideología 6(1): 14. 1971.

Accepted Name: Scaphosepalum grande .

Type: Colombia, Escobar 712 (holotype, AMES).

Scaphosepalum nutans Kraenzl., Notizbl. Bot. Gart. Berlin-Dahlem 8: 136. 1922.

Accepted Name: Scaphosepalum breve .

Type: Colombia, Kalbreyer 1471 (holotype, B, destroyed).

Scaphosepalum ochthodes (Rchb.f.) Pfitzer, Nat. Pflanzenfam. 2(6): 39. 1888.

Accepted Name: Scaphosepalum verrucosum .

Type: Colombia, Wagener 564 (holotype, W).

Scaphosepalum platypetalum Schltr., Repert. Spec. Nov. Regni Veg. 7: 75. 1920.

Accepted Name: Scaphosepalum swertiifolium .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 25, No. 93)].

Scaphosepalum reversum Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 436. 1921.

Accepted Name: Scaphosepalum antenniferum .

Type: Colombia, unknown (holotype, W).

Scaphosepalum rolfeanum Kraenzl., Repert. Spec. Nov. Regni Veg. Beih. 34: 211. 1925.

Accepted Name: Scaphosepalum pulvinare .

Type: Colombia, Boxall s.n. [lectotype, W (Luer 1988)].

Scaphosepalum sneiderniiGaray, Svensk Bot. Tidskr. 47: 202. 1953.

Accepted Name: Scaphosepalum odontochilum .

Type: Colombia, v. Sneidern 950 (holotype, S).

Scaphosepalum swertiifolium subsp. exiguum Luer & R.Escobar, Monogr. Syst. Bot. Missouri Bot. Gard. 26: 91, pl. 37-38. 1988.

Accepted Name: Scaphosepalum swertiifolium .

Type: Colombia, Escobar 2453 (holotype, SEL).

Scaphosepalum xipheres (Rchb.f.) Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 220. 1920.

Accepted Name: Porroglossum muscosum .

Type: Colombia, Roezl s.n. (holotype, W).

Specklinia florulenta (Linden & Rchb.f.) Pridgeon & M.W.Chase, Lindleyana 16(4): 257. 2001.

Accepted Name: Specklinia picta .

Type: Colombia, Linden 55 (holotype,W; isotype, AMES).

Specklinia lepanthoides (Schltr.) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 261. 2004.

Accepted Name: Karma arbuscula .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype, Schlechter’s floral analysis published by Mansfeld (1929: Pl. 37, No. 140).

Specklinia pereziana Kolan., Biodiv. Res. Conserv. 24: 19-22. 2011.

Accepted Name: Lankesteriana barbulata .

Type: Colombia, Kolanowska & Pérez 263 (holotype, UGDA-spirit).

Specklinia tsubotae (Luer & R.Escobar) Luer, Monogr. Syst. Bot. Missouri Bot. Gard. 95: 264. 2004.

Accepted Name: Muscarella barberiana .

Type: Colombia, Luer 16763 (holotype, MO).

Stelis abrupta Rchb.f., Linnaea 41: 7. 1877.

Accepted Name: Stelis purpurea .

Type: Colombia, Roezl s.n. (holotype, W; drawing of type, AMES).

Stelis apiculata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 84. 1920, nom. illeg.

Accepted Name: Stelis oblonga .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 28, No. 105)].

Stelis arevaloi Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 25. 1924.

Accepted Name: Stelis lamellata .

Type: Colombia, Hopp 145 (holotype, B, destroyed). Colombia, Luer et al. 13844 [neotype, MO (Luer 2009a)].

Stelis bracteata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 26-27. 1924, nom. illeg.

Accepted Name: Stelis braccata .

Type: Colombia, Hopp 58 (holotype, B, destroyed).

Stelis bractescensGaray, Bot. Mus. Leafl. 27(7-9): 185. 1979.

Accepted Name: Stelis braccata .

Type: Colombia, Hopp 58 (holotype, B, destroyed).

Stelis bruchmuelleri Rchb.f. ex Hook.f., Bot. Mag. 106: t. 6521. 1880.

Accepted Name: Stelis eublepharis .

Type: Colombia, Veitch & Sons s.n. (holotype, K).

Stelis caespitifica Rchb.f., Bonplandia (Hannover) 3: 71. 1855.

Accepted Name: Stelis nitens .

Type: Colombia, Wagener s.n. (holotype, W; isotype, AMES-00287033).

Stelis caespitula Luer & R.Escobar, Harvard Pap. Bot. 21(1): 62. 2016.

Accepted Name: Stelis silverstonei .

Type: Colombia, Luer 13221 (holotype, MO).

Stelis cairoënsis Luer, Selbyana 30: 20-21. 2009.

Accepted Name: Stelis lindenii .

Type: Colombia, Silverstone-Sopkin 8000 (holotype, CUVC; isotype, MO).

Stelis canaliculata Rchb.f., Gard. Chron. 1718. 1872.

Accepted Name: Stelis tenuilabris .

Type: Colombia, Saunders 2707 (holotype, W).

Stelis caucae Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 85. 1920.

Accepted Name: Stelis purpurea .

Type: Colombia, Lehmann 6923 [holotype, B, destroyed; lectotype (selected here), K-000573941; isolectotypes, LE-00001388, NY-00009391].

Stelis citrina Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 86. 1920.

Accepted Name: Stelis pusilla .

Type: Colombia, Lehmann 3692 [lectotype K-000573947 (Luer 2009a); isolectotypes, BR, AMES-00104676, BM-000084109, NY-00009392, US-00093546].

Stelis convallarioides Garay, Canad. J. Bot. 34(3): 351-352. 1956.

Accepted Name: Stelis superbiens .

Type: Colombia, Archer 2143 (holotype, US1519162).

Stelis dazae Garay, Bot. Mus. Leafl. 27(7-9): 202 (index). 1980, nom. inval.

Accepted Name: Stelis polyclada .

Type: Colombia, Hopp 151 (holotype, B, destroyed).

Stelis dendrophila Luer & R.Escobar, Harvard Pap. Bot. 22(1): 34. 2017.

Accepted Name: Stelis atra .

Type: Colombia, Luer & Escobar 7619 (holotype, SEL).

Stelis dynamica Luer & R.Escobar, Harvard Pap. Bot. 22(1): 36. 2017.

Accepted Name: Stelis attenuata .

Type: Colombia, Luer et al. 7466 (holotype, SEL).

Stelis elegans (Kunth) Pridgeon & M.W.Chase, Lindleyana 16(4): 262. 2001, nom. illeg.

Accepted Name: Stelis roseopunctata .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; fragment, W).

Stelis elegantula Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 28. 1924.

Accepted Name: Stelis pusilla .

Type: Colombia, Hopp 9 (holotype, B, destroyed). Ecuador, Hirtz & Hirtz 7542 [neotype, MO (Luer 2009a)].

Stelis endocharis Rchb.f., Linnaea 41: 8. 1877.

Accepted Name: Stelis purpurea .

Type: Colombia, Roezl s.n. (holotype, W).

Stelis ephippium Luer & R.Escobar, Harvard Pap. Bot. 22(1): 39. 2017, nom. inval.

Accepted Name: Stelis sesquipedalis .

Distribution: COL VEN.

Type: Colombia, Luer et al. 10323 (holotype, MO).

Note: Invalid for citation of two holotypes.

Stelis exilipes Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 28. 1924.

Accepted Name: Stelis striolata .

Type: Colombia, Hopp 41 (holotype, B, destroyed).

Stelis floribunda Kunth, Nov. Gen. Sp. (folio ed.) 1: 362. 1815 [1816].

Accepted Name: Stelis oblonga .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Stelis fruticulus Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 88. 1920.

Accepted Name: Stelis truncata .

Type: Colombia, Lehmann 8214 [holotype, B, destroyed; lectotype (selected here), K-000573931; isolectotypes, AMES-00084207, LE-00001393, NY-00009395].

Stelis gustavii O.Duque, Orquideología 20(3): 348-350. 1997.

Accepted Name: Stelis dressleri .

Type: Colombia, Duque 1835 (holotype, JAUM).

Stelis hemicardia Rchb.f., Linnaea 41: 8. 1877.

Accepted Name: Stelis purpurea .

Type: Colombia, Roezl s.n. (holotype, W).

Stelis herbiola Lindl., Fol. Orchid. ~Stelis~ 10. 1852- 1855 [1859].

Accepted Name: Stelis pusilla .

Type: Colombia, Holton s.n. (holotype, K).

Stelis inedita Luer & R.Escobar, Harvard Pap. Bot. 22(1): 48. 2017.

Accepted Name: Stelis vulcani .

Type: Colombia, Luer et al. 7655 (holotype, SEL).

Stelis insignis Ames, Sched. Orch. 1: 5. 1922.

Accepted Name: Stelis oblonga .

Type: Colombia, Pittier 1299 (holotype, US531497; AMES, 25099).

Stelis lobata Rchb.f., Bonplandia (Hannover) 3: 241. 1855.

Accepted Name: Stelis elongata .

Type: Colombia, Jameson 481 (holotype, W; isotypes, BM, BR, G, LE).

Stelis macrocarpa Kunth, Nov. Gen. Sp. (quarto ed.) 1: 363. 1816.

Accepted Name: Stelis purpurea .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P).

Stelis melanoxantha Rchb.f., Bonplandia (Hannover) 3: 71. 1855.

Accepted Name: Stelis purpurea .

Type: Colombia, Wagener s.n. (holotype, W).

Stelis minimiflora Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 31. 1924.

Accepted Name: Stelis braccata .

Type: Colombia, Hopp 171 (holotype, B, destroyed).

Stelis mononeura Lindl., Fol. Orchid. ~Stelis~ 4, no. 25. 1852-1855 [1859].

Accepted Name: Stelis euspatha .

Type: Colombia, Jameson s.n. (holotype, K).

Stelis myriantha Schltr., Repert. Spec. Nov. Regni Veg. 7: 90. 1920, nom. illeg.

Accepted Name: Stelis tenuilabris .

Type: Colombia, Lehmann 6920 [holotype, B, destroyed; lectotype (selected here), K-000573957; isolectotype, LE-00001401]. Colombia, Luer et al. 12035 [neotype, MO (Luer 2009a), here rejected].

Stelis navicularis Garay, Canad. J. Bot. 34: 353. 1956.

Accepted Name: Stelis colombiana .

Type: Colombia, John 20875 (holotype, AMES; isotype, US).

Stelis pachyphylla Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 91. 1920.

Accepted Name: Stelis purpurea .

Type: Colombia, Madero s.n. [holotype, B, destroyed; lectotype (selected here), Schlechter’s floral analysis published by Mansfeld (1929: Pl. 31, No. 116)].

Stelis patella O.Duque, Orquideología 27(2): 134, 137, Fotos 48-50, fig. 18. 2010.

Accepted Name: Stelis asseris .

Type: Colombia, Duque 196 (holotype, JAUM).

Stelis pastoensis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 35. 1924.

Accepted Name: Stelis oblongifolia .

Type: Colombia, Hopp 150 (holotype, B, destroyed).

Stelis picta Pridgeon & M.W.Chase, Lindleyana 17(2): 100. 2002.

Accepted Name: Stelis roseopunctata .

Type: Colombia, Humboldt & Bonpland s.n. (holotype, P; isotype, W).

Stelis pinguis Luer & R.Escobar, Harvard Pap. Bot. 21(1): 82. 2016.

Accepted Name: Stelis cochlearis .

Type: Colombia, Luer et al. 8747 (holotype, SEL).

Stelis pleurothalloides Ames, Orchidaceae 7: 132. 1922.

Accepted Name: Stelis furfuracea .

Type: Colombia, Pittier 1156 (holotype, US; isotype, AMES).

Stelis prionota Luer & R.Escobar, Harvard Pap. Bot. 21(2): 216. 2016.

Accepted Name: Stelis cochlearis .

Type: Colombia, Luer 16696 (holotype, MO).

Stelis purdiaei Lindl., Fol. Orchid. ~Stelis~ 5, no. 31. 1852-1855 [1859].

Accepted Name: Stelis pugiunculi .

Type: Colombia, Purdie s.n. (holotype, K; isotype, GH-00104938).

Stelis ringens Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 37. 1924.

Accepted Name: Stelis velutina .

Type: Colombia, Hopp 49 (holotype, B, destroyed).

Stelis robusta Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 38. 1924, nom. illeg.

Accepted Name: Stelis purpurea .

Type: Colombia, Hopp 56 (holotype, B, destroyed).

Stelis robustiorGaray, Bot. Mus. Leafl. 27: 185. 1979.

Accepted Name: Stelis purpurea .

Type: Colombia, Hopp 56 (holotype, B, destroyed).

Stelis scandens Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 39. 1924, nom. illeg.

Accepted Name: Stelis polyclada .

Type: Colombia, Hopp 151 (holotype, B, destroyed).

Stelis stella O.Duque, Orquideología 20(3): 373, 374. 1997.

Accepted Name: Stelis tridactylon .

Type: Colombia, Duque 1143 (holotype, JAUM).

Stelis tenuis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 94. 1920.

Accepted Name: Stelis pusilla .

Type: Colombia, Lehmann 6039 [holotype, B, destroyed; lectotype (selected here), K-000573948; isolectotypes, AMES-00105004, NY-00009416, LE-00001407, US-00093597].

Stelis tonsoria Luer, Phytologia 49(3): 239. 1981.

Accepted Name: Stelis langlassei .

Type: Colombia, Luer et al. 4579 (holotype, SEL).

Stelis trianaei Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 94. 1920.

Accepted Name: Stelis scansor .

Type: Colombia, Triana 566 [holotype, B, destroyed; lectotype (selected here), P-00489435; isolectotypes, AMES-00084457, BM-000082571, US-00093598].

Stelis umbelliformis subsp. brevipedunculata Carnevali & I.Ramírez, Novon 3(2): 121. 1993.

Accepted Name: Stelis umbelliformis .

Type: Colombia, Benavides 4284 (holotype, PSO; isotype, MO).

Stelis vagans Ames, Sched. Orch. 1: 8. 1922.

Accepted Name: Stelis prorepens .

Type: Colombia, Pittier 1061 (holotype, US531263).

Stelis velaticauloides (Kolan. & Szlach.) Karremans, Phytotaxa 496(2): 142. 2021.

Accepted Name: Stelis pennelliana .

Type: Colombia, Pennell 2043 (holotype, AMES; drawing, UGDA).

Stelis virgulata Schltr., Repert. Spec. Nov. Regni Veg. Beih. 27: 40. 1924.

Accepted Name: Stelis truncata .

Type: Colombia, Hopp 44 (holotype, B).

Trichosalpinx carinatus Rykacz., Szlach. & Kolan., Ann. Bot. Fenn. 55(1-3): 146. 2018.

Accepted Name: Pendusalpinx patula .

Type: Colombia, Schneider 353 (holotype, AMES; isotype, UGDA).

Note: This appears to be based on an imprecise illustration of a dried specimen of P. patula.

Trichosalpinx dithele Luer & R.Escobar, Orquideología 16: 181. 1984.

Accepted Name: Trichosalpinx semilunata .

Type: Colombia, Luer et al. 9044 (holotype, SEL).

Trichosalpinx falcipetala (Schltr.) Luer, Phytologia 54(5): 395. 1983.

Accepted Name: Stellamaris pergrata .

Type: Colombia, Lehmann 2045 (holotype, B, destroyed). Colombia, Lehmann 8245 [neotype, K (Luer 1997)].

Trichosalpinx lancifera (Schltr.) Luer, Phytologia 54(5): 396. 1983.

Accepted Name: Trichosalpinx orbicularis .

Type: Colombia, Hopp 67 (holotype, B, destroyed).

Trichosalpinx pyxos (Luer & R.Escobar) Luer, Phytologia 54(5): 397. 1983.

Accepted Name: Karma amygdalora .

Type: Colombia, Luer et al. 3047 (holotype, SEL).

Trichosalpinx vertex Luer & R.Escobar, Orquideología 16(2): 186. 1984.

Accepted Name: Karma tenuis .

Type: Colombia, Luer & Escobar 8488 (holotype, SEL; isotypes, COL, JAUM).

Excluded Taxa

Acianthera capillaris (Lindl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015) and Bernal et al. (2016), here treated as Acianthera floribunda.

Acianthera cogniauxiana (Schltr.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher.

Acianthera ellipsophylla (L.O. Williams) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Acianthera glumacea (Lindl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Acianthera rubroviridis (Lindl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher.

Acianthera strupifolia (Lindl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Anathallis acuminata (Kunth) Pridgeon & M.W.Chase.

Treated here as a synonym of Stelis aurea.

Anathallis ciliolata (Schltr.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher.

Anathallis dolichopus (Schltr.) Pridgeon & M.W.Chase.

This is a Central American species. Colombian materials are referable to Stelis sclerophylla.

Anathallis funerea (Barb.Rodr.) Luer.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher. It is highly likely to occur in the country.

Anathallis marginata (Barb.Rodr.) F.Barros & Barberena.

Cited by Betancur et al. (2015), no available voucher.

Anathallis microphyta (Barb.Rodr.) C.O.Azevedo & Van den Berg.

Cited by Betancur et al. 2015, no available voucher.

Anathallis stenophylla (F.Lehm. & Kraenzl.) Pridgeon & M.W.Chase.

Treated here as a synonym of Stelis melanopus.

Anathallis taracuana (Schltr.) F.Barros & Barberena.

Cited by Betancur et al. (2015), no available voucher.

Anathallis ypirangae (Kraenzl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Ancipitia jimii (Luer) Luer.

An invalid name, treated here as a synonym of Pleurothallis lueriana.

Andinia persimilis (Luer & Sijm) Karremans & S.Vieira-Uribe.

Colombian specimens labeled A. aff. persimilis in Karremans & S.Vieira-Uribe (2020) are not that species and belong to un undescribed taxon.

Dracula dodsonii (Luer) Luer.

Cited by Bernal et al. (2016) and Govaerts (2022), no available voucher.

Dracula pusilla (Rolfe) Luer.

Cited by Bernal et al. (2016), no available voucher.

Dryadella perpusilla (Kraenzl.) Luer.

Cited by Betancur et al. (2015), no available voucher.

Dryadella summersii (L.O. Williams) Luer.

Cited by Betancur et al. (2015), no available voucher.

Echinosepala uncinata (Fawc.) Pridgeon & M.W.Chase.

Colombian specimens are treated as E. isthmica following Pupulin et al. (2020).

Frondaria caulescens (Lindl.) Luer.

Colombian specimens are treated as F. colombiana following Szlachetko et al. (2018).

Lepanthes actias-luna Luer & Hirtz.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes capitana Rchb.f.

Cited by Betancur et al. (2015), most likely a misidentified voucher.

Lepanthes crista-pulli Luer & R.Escobar.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes erinacea Rchb.f.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes esmeralda Luer & Hirtz.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes hagsateri Salazar & Soto Arenas.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes helicocephala Rchb.f.

Cited by Bernal et al. (2016), no available voucher.

Lepanthes juninensis Schltr.

Cited by Betancur et al. (2015), most likely based on a misidentified voucher.

Lepanthes lindleyana Oerst. & Rchb.f.

Cited by Betancur et al. 2015, most likely based on a misidentified voucher.

Lepanthes lloensis Luer.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes megalostele Luer.

Cited by Betancur et al. (2015) based on a misidentified specimen in Misas Urreta (2005).

Lepanthes nanegalensis Rchb.f.

Cited by Betancur et al. (2015) and Bernal et al. (2016), no available voucher.

Lepanthes pan Luer & Dalström.

Cited by Betancur et al. (2015) based on a misidentified specimen in Misas Urreta (2005).

Lepanthes plectilis Luer & Hirtz.

Cited by Betancur et al. 2015, no available voucher.

Lepanthes pumila C.Schweinf.

Cited by Betancur et al. 2015, most likely based on misidentified vouchers.

Lepanthes rabeiFoldats, Acta Bot. Venez. 3: 41. 1968.

Cited by Betancur et al. (2015), here treated as a synonym of L. aquila-borussiae.

Lepanthes rotundifolia L.O. Williams.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes rubripetala Stimson.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes rupestris Stimson.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes rupicola Schltr.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes ruscifolia Rchb.f.

Cited by Betancur et al. (2015), no available voucher.

Lepanthes saltatrix Luer & Hirtz.

Cited by Bernal et al. (2016) and Govaerts (2022), no available voucher.

Lepanthes samacensis Ames.

Cited by Betancur et al. (2015), most likely a misidentified voucher.

Lepanthes unitrinervis Carnevali & I. Ramírez.

Cited by Betancur et al. (2015), no available voucher.

Masdevallia alphonsiana A.Doucette ex L.E.Matthews.

Cited by Govaerts (2022), no available voucher.

Masdevallia biflora Regel.

Cited by Govaerts (2022), no available voucher.

Masdevallia bulbophyllopsis Kraenzl.

Cited by Bernal et al. (2016), no available voucher.

Masdevallia ensata Rchb.f.

Cited by Bernal et al. (2016) and Govaerts (2022), no available voucher.

Masdevallia fulvescens Rolfe.

Cited by Govaerts (2022), no available voucher.

Masdevallia kuhniorum Luer.

Cited by Govaerts (2022), no available voucher.

Masdevallia lehmannii Rchb.f.

Cited by Bernal et al. (2016), no available voucher.

Masdevallia maculata Klotzsch & H.Karst.

Cited by Govaerts (2022), no available voucher.

Masdevallia navicularis Garay & Dunst.

Cited by Bernal et al. (2016), no available voucher.

Masdevallia pachygyne Kraenzl.

Cited by Govaerts (2022), no available voucher.

Masdevallia virgo-cuencae Luer & Andreetta.

Cited by Bernal et al. (2016) and Govaerts (2022), no available voucher.

Masdevallia xiphium Rchb.f. ex Kraenzl.

Cited by Govaerts (2022), no available voucher.

Myoxanthus speciosus (Luer) Luer.

Here treated as a synonym of M. exasperatus following Rojas-Alvarado et al. (2021).

Octomeria amazonica Pabst.

Cited by Betancur et al. (2015) and Bernal et al. (2016), a synonym of O. colombiana sensu Luer (2010).

Octomeria chloidophylla (Rchb.f.) Garay.

Cited by Betancur et al. (2015) and Govaerts (2022), no available voucher.

Octomeria erosilabia C.Schweinf.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), a synonym of O. longifolia sensu Luer (2010).

Octomeria minor C.Schweinf.

Cited by Betancur et al. (2015), no available voucher.

Octomeria schultesii Pabst.

Cited by Bernal et al. (2016) and Govaerts (2022), a synonym of O. longifolia sensu Luer (2010).

Octomeria surinamensis H.Focke.

Here treated as a synonym of O. tridentata Lindl. following Luer (2010).

Pabstiella parvifolia (Lindl.) Luer.

Cited by Bernal et al. (2016) and Govaerts (2022), but this is a Brazilian endemic. Colombian materials are referable to Andreettaea cryptophyta (Barb.Rodr.) A.Doucette (see Karremans & Vieira-Uribe 2020).

Platystele brenneri Luer.

Cited by Betancur et al. (2015) based on a misidentified specimen in Misas Urreta (2005).

Platystele compacta (Ames) Ames.

Cited by Betancur et al. (2015) based on a misidentified specimen in Misas Urreta (2005).

Platystele cornejoi Luer.

Cited by Cornejo (2014) as “seen” in Colombia, but no voucher is currently available.

Platystele dressleri Luer.

Cited by Betancur et al. (2015) based on a specimen in Misas Urreta (2005), and cited by Ortiz (1977) based a specimen therein. Those photographs and drawings suggest Platystele argentosa Luer & R.Escobar rather than P. dressleri.

Platystele ornata Garay.

Cited by Betancur et al. (2015), no available voucher.

Platystele taylorii Luer.

Cited by Betancur et al. (2015), no available voucher.

Platystele ximenae Luer & Hirtz.

Cited by Govaerts (2022), no available voucher.

Pleurothallis apopsis Luer.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis archicolonae Luer.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis canaliculata Rchb.f.

Cited by Betancur et al. (2015), this name seems to be unpublished.

Pleurothallis capillaris Lindl., Edwards’s Bot. Reg. 21: sub t. 1797. 1836.

Here treated as Acianthera floribunda.

Pleurothallis cornualis Luer.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher.

Pleurothallis cuscifolia R.Br.

Cited by Betancur et al. (2015), but this is simply a misspelled name.

Pleurothallis foliosa Hook.

Cited by Betancur et al. (2015), this name applies to a Gomesa species.

Pleurothallis gargantua Luer.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis longipedicellata Ames & C.Schweinf.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis luctuosa Rchb.f.

Colombian materials are referable to Pleurothallis tenuisepala Mark Wilson (see Wilson et al. 2019).

Pleurothallis miserrima Lindl.

Cited by Betancur et al. (2015), this name seems to be unpublished.

Pleurothallis ophiocephala Lindl., Edwards’s Bot. Reg. 24: Misc. 34. 1838.

This widespread species could occur in Colombia. However, at this time we are not able to cite any voucher to support its inclusion. A widely cited specimen at JAUM with the number Ospina 81 (Betancur et al. 2015, Bernal et al. 2016) is actually a specimen of Pleurothallis peduncularis Lindl. (= Myoxanthus parahybunensis) not Pleurothallis peduncularis Hook. (= Pleurothallis ophiocephala).

Pleurothallis parviflora Luer.

Cited by Betancur et al. (2015), but the voucher is of Andreettaea cryptophyta (Barb.Rodr.) A.Doucette (see Karremans & Vieira-Uribe 2020).

Pleurothallis perryi Luer.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis raymondi Rchb.f.

Cited by Betancur et al. (2015), but this is simply a misspelled name.

Pleurothallis subsinuata Lindl.

Cited by Betancur et al. (2015) and Govaerts (2022), no available voucher. This taxon is of unknown country of origin and definitively belongs in genus Specklinia.

Pleurothallis suspensa Luer.

Cited by Betancur et al. (2015) and Govaerts (2022), no available voucher.

Pleurothallis tentaculata (Poepp. & Endl.) Lindl.

Cited by Betancur et al. (2015), Bernal et al. (2016) and Govaerts (2022), no available voucher.

Pleurothallis tetrachaeta Luer & Hirtz.

Cited by Betancur et al. (2015), no available voucher.

Pleurothallis trachysepala Kraenzl.

Here treated as a synonym of P. cordata.

Pleurothallis rubella Luer.

Here treated as a synonym of Platystele aurea Garay.

Pleurothallis virens Lindl.

Cited by Betancur et al. (2015), this name seems to be unpublished.

Restrepia roseola Luer & R.Escobar.

Cited by Bernal et al. (2016), no available voucher.

Restrepiella ophiocephala (Lindl.) Garay & Dunst., Venez. Orchid. Ill. Vol. 1. 4: 266. 1966.

This widespread species could occur in Colombia. However, at this time we are not able to cite any voucher to support its inclusion. A widely cited specimen at JAUM with the number Ospina 81 (Betancur et al. 2015, Bernal et al. 2016) is actually a specimen of Pleurothallis peduncularis Lindl. (= Myoxanthus parahybunensis) not Pleurothallis peduncularis Hook. (= Restrepiella ophiocephala).

Specklinia alata (A. Rich. & Galeotti) Solano & Soto Arenas.

Cited by Betancur et al. (2015), this name is a synonym of Andreettaea marginata (Rich.) A.Doucette a taxon known only from northern Middle America. The authors probably mean another Andreettaea or Specklinia species.

Specklinia aristata (Hook.) Pridgeon & M.W.Chase.

Cited by Dodson (1992), Misas Urreta (2005), Betancur et al. (2015), Bernal et al. (2016), and Govaerts (2022). It is likely that this species occurs in Colombia, however none of the materials that we have inspected bearing this name are correctly identified. The specimen Misas-Urreta 114 (HPUJ) corresponds to Andreettaea clavigera, while the photograph in Ortiz & Uribe 2007 (CD Edition) depicts Andreettaea barberiana.

Specklinia bicornis (Luer) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Specklinia endotrachys (Rchb.f.) Pridgeon & M.W.Chase.

Colombian specimens cited by Misas Urreta (2006) and Luer (2007) are referable to S. dunstervillei, the true S. endotrachys is endemic to Costa Rica.

Specklinia lanceola (Sw.) Lindl.

Cited by Bernal et al. (2016) and Govaerts (2022), no available voucher. The voucher cited by Betancur et al. (2015) is an unrelated species in Specklinia subgen. Sarcinula Karremans.

Specklinia remotiflora Pupulin & Karremans.

Colombian specimens cited by Pupulin et al. (2012) are referable to S. dunstervillei.

Specklinia uniflora (Lindl.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Stelis argentea Lindl.

Cited by Betancur et al. (2015), but this is simply a misspelled name.

Stelis deregularis Barb.Rodr.

Cited by Betancur et al. (2015), no available voucher. It is likely to occur in Colombia given widespread this species is.

Stelis flaccida (Klinge) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015) and Bernal et al. (2016). This is an obscure taxon of unknown origin and it belongs in genus Specklinia.

Stelis humilis Lindl.

Cited by Dorr (2014) as likely to occur as it has been collected close to the border in Venezuela, no available voucher.

Stelis grossilabris Rchb.f.

Treated here as a synonym of Stelis grandiflora.

Stelis guatemalensis Schltr.

Treated here as a synonym of Stelis pardipes.

Stelis muscifera Lindl.

Treated here as a synonym of Stelis grandiflora.

Stelis nanegalensis Lindl.

Treated here as a synonym of Stelis superbiens.

Stelis neorubens Chiron.

Treated here as a synonym of Stelis montserrati.

Stelis ophioglossoides (Jacq.) Sw.

Cited by Betancur et al. (2015), no available voucher.

Stelis polybotrya Lindl.

Cited by Bernal et al. (2016), no available voucher.

Stelis punctulata (Rchb.f.) Soto Arenas.

Type erroneously labeled Neu Granada, but known only from Central America (see Luer 2006)

Stelis punoensis C.Schweinf.

Treated here as a synonym of Stelis dupliciformis.

Stelis quinquenervia C.Schweinf.

Cited by Bernal et al. (2016), no available voucher.

Stelis rostrata Luer.

Cited by Betancur et al. (2015), this is a nomenclatural synonym of S. septicola.

Stelis scabrida Lindl.

Cited by Betancur et al. (2015), this is a synonym of S. ophioglossoides a species from the Lesser Antilles.

Stelis segoviensis (Rchb.f.) Pridgeon & M.W.Chase.

Cited by Betancur et al. (2015), no available voucher.

Stelis transversalis Ames.

Cited by Bernal et al. (2016), no available voucher.

Stelis tricardium Lindl.

Cited by Dodson (1992), no available voucher.

Stelis villosa (Knowles & Westc.) Pridgeon & M.W.Chase.

Cited by Bernal et al. (2016), no available voucher.

Stelis vulcanica Schltr.

Cited by Betancur et al. (2015), this is a synonym of S. superbiens.

Stelis wagneri (Schltr.) Pridgeon & M.W.Chase.

Cited by Luer (1998b), Bernal et al. (2016), no available voucher.

Stelis yangaensis Luer & Hirtz.

Cited by Betancur et al. (2015), no available voucher.

Teagueia teaguei (Luer) Luer.

Cited by Betancur et al. (2015), no available voucher.

Trichosalpinx cedralensis (Ames) Luer.

Cited by Betancur et al. (2015), no available voucher.

Trichosalpinx montana (Barb.Rodr.) Luer.

Betancur et al. (2015) cite Ospina 86 (JAUM) as voucher, but this a misidentified specimen.

Trichosalpinx quitensis (Rchb.f.) Luer.

Bernal et al. (2016) cite Ospina 86 (JAUM) as voucher, but this a misidentified specimen.

Trichosalpinx xiphochila (Rchb.f.) Luer.

Cited by Betancur et al. (2015), this is an obscure taxon from Venezuela which probably belongs in genus Platystele instead of Trichosalpinx.

Yolanda floribunda (Garay) Brieger.

Treated here as a synonym of Brachionidium tuberculatum.

Zootrophion gracilentum (Rchb.f.) Luer.

Cited by Betancur et al. (2015), no available voucher.

Zootrophion griffin Luer.

Cited by Uribe-Vélez & Sauleda (2023), the illustrated species is actually Zootrophion machaqway A.Doucette & J. Portilla.

Zootrophion moorei (Rolfe) Luer.

Treated here as a synonym of Zootrophion hypodiscus.

Zosterophyllanthos fornix (Luer & R.Escobar) Szlach. & Marg.

Treated here as a synonym of Pleurothallis calceolaris.

Zosterophyllanthos granularis (Luer & R.Escobar) Szlach. & Marg.

Treated here as a synonym of Pleurothallis platysepala.

Zosterophyllanthos monocardius (Rchb.f.) Szlach. & Marg.

Treated here as a synonym of Pleurothallis cordata.

Zosterophyllanthos northenae (Luer) Szlach. & Marg.

Treated here as a synonym of Pleurothallis latipetala.

Zosterophyllanthos pulvinipes (Schltr.) Szlach. & Marg.

Treated here as a synonym of Pleurothallis macrocardia.